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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.70.762" ID-GBIF-Dataset="b8fceb5a-9f78-4dc4-8fa2-92fe20bb2c42" ID-PMC="PMC3088446" ID-Pensoft-Pub="1313-2970-70-1" ID-PubMed="21594041" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-70-1" ModsDocOrigin="ZooKeys 70" ModsDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an “oceanic” island" checkinTime="1451250880545" checkinUser="pensoft" docAuthor="Rowson, B, Warren, B. H. &amp; Ngereza, C. F." docDate="2010" docId="8FCA78D1BA57CEB3608647DBBF3E8407" docLanguage="en" docName="ZooKeys 70: 1-39" docOrigin="ZooKeys 70" docSource="http://dx.doi.org/10.3897/zookeys.70.762" docTitle="Allopeas gracile Hutton 1834" docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FFB2FFB6591BFFBAFFFF0472FFE6A56D" masterDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island" masterLastPageNumber="39" masterPageNumber="1" pageNumber="16" updateTime="1668164835731" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rowson, B</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Warren, B. H.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ngereza, C. F.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>70</mods:number>
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<mods:start>1</mods:start>
<mods:end>39</mods:end>
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<treatment ID-GBIF-Taxon="159360813" LSID="urn:lsid:plazi:treatment:8FCA78D1BA57CEB3608647DBBF3E8407" httpUri="http://treatment.plazi.org/id/8FCA78D1BA57CEB3608647DBBF3E8407" lastPageId="16" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection pageId="15" pageNumber="16" type="nomenclature">
<paragraph pageId="15" pageNumber="16">
17.
<taxonomicName authority="Hutton, 1834" authorityName="Hutton" authorityYear="1834" class="Gastropoda" family="Achatinidae" genus="Allopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allopeas gracile" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="gracile">Allopeas gracile (Hutton, 1834)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="reference_group">
<paragraph pageId="15" pageNumber="16">
<taxonomicName class="Gastropoda" family="Acteonidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus gracile" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="gracile">Bulimus gracile</taxonomicName>
<bibRefCitation author="Hutton, T" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="23" pageNumber="24" pagination="81 - 93" title="On the land shells of India." volume="3" year="1834">Hutton 1834</bibRefCitation>
: 84, 93
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="notes">
<paragraph pageId="15" pageNumber="16">Notes.</paragraph>
<paragraph lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Pemba specimens reach a large size (13.5
<normalizedToken originalValue="×">x</normalizedToken>
3.4 mm, occasionally 15.0
<normalizedToken originalValue="×">x</normalizedToken>
3.5 mm, up to 8 whorls), always being relatively narrower than
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Opeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opeas lamoense" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="lamoense">Opeas lamoense</taxonomicName>
. The maximum size is substantially greater than given by most authors (e.g.
<bibRefCitation author="Gerlach, J" journalOrPublisher="Backhuys, Leiden, The Netherlands" pageId="22" pageNumber="23" title="Terrestrial and freshwater Mollusca of the Seychelles islands" year="2006 b">Gerlach 2006b</bibRefCitation>
: 9.4 mm;
<bibRefCitation author="Griffiths, OL" journalOrPublisher="Bioculture Press, Mauritius" pageId="23" pageNumber="24" title="A field guide to the non-marine molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion) and the Northern Dependencies of Mauritius" year="2006">Griffiths and Florens 2006</bibRefCitation>
: 13.5 mm). Said to be restricted to &quot;waste places&quot; by
<bibRefCitation author="Verdcourt, B" editor="Burgess, ND" journalOrPublisher="IUCN, Gland, Switzerland" pageId="24" pageNumber="25" title="Molluscs" volumeTitle="Coastal Forests of Eastern Africa" year="2000">Verdcourt (2000)</bibRefCitation>
this species nonetheless occurs in some forest habitats (Table 2).
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Allopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allopeas gracile" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="gracile">Allopeas gracile</taxonomicName>
is widespread in the tropics but its native range is uncertain.
<bibRefCitation author="Voeltzkow, A" journalOrPublisher="Witu-Inseln und Zanzibar-Archipel. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart" pageId="24" pageNumber="25" title="Reise in Ostafrika in den Jahren 1903 - 1905. Wissenschaftliche Ergebnisse" year="1923">
<pageBreakToken pageId="16" pageNumber="17" start="start">Voeltzkow</pageBreakToken>
(1923)
</bibRefCitation>
noted that &quot;
<taxonomicName genus="Opea" lsidName="Opea gracile subsp. johanninum" pageId="16" pageNumber="17" rank="subSpecies" species="gracile" subSpecies="johanninum">Opea gracile johanninum</taxonomicName>
(Mor.)&quot; (sic) was numerous in rotting logs on Pemba;
<bibRefCitation author="Haas, F" journalOrPublisher="Zoologische Jahrbuecher" pageId="23" pageNumber="24" pagination="387 - 430" title="Die Binnenmollusken der Voeltzkow'schen Reisen in Ostafrika und den ostafrikanischen Inseln" volume="57" year="1929">Haas (1929)</bibRefCitation>
gives the record from Chake Chake.
<bibRefCitation author="Fischer-Piette, E" journalOrPublisher="Memoires du Museum National D'Histoire Naturelle (Nouvelle Serie: Serie A, Zoologie)" pageId="22" pageNumber="23" pagination="1 - 77" title="Les mollusques terrestres des Iles Comores." volume="84" year="1974">Fischer-Piette and Vukadinovic (1974)</bibRefCitation>
consider
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus johanninus" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="johanninus">Bulimus johanninus</taxonomicName>
Morelet, 1877, described from the Comoros, a synonym of
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Allopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allopeas gracile" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="gracile">Allopeas gracile</taxonomicName>
. Recent authors (e.g.
<bibRefCitation author="Neubert, E" journalOrPublisher="Fauna of Arabia" pageId="23" pageNumber="24" pagination="333 - 461" title="Annotated checklist of the terrestrial and freshwater molluscs of the Arabian Peninsula with descriptions of new species." volume="17" year="1998">Neubert 1998</bibRefCitation>
,
<bibRefCitation author="Gerlach, J" journalOrPublisher="Backhuys, Leiden, The Netherlands" pageId="22" pageNumber="23" title="Terrestrial and freshwater Mollusca of the Seychelles islands" year="2006 b">Gerlach 2006b</bibRefCitation>
,
<bibRefCitation author="Griffiths, OL" journalOrPublisher="Bioculture Press, Mauritius" pageId="23" pageNumber="24" title="A field guide to the non-marine molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion) and the Northern Dependencies of Mauritius" year="2006">Griffiths and Florens 2006</bibRefCitation>
) rather than
<bibRefCitation author="Verdcourt, B" journalOrPublisher="B Verdcourt, Maidenhead, UK" pageId="24" pageNumber="25" title="A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi)" year="2006">Verdcourt (2006)</bibRefCitation>
and
<bibRefCitation author="Rowson, B" journalOrPublisher="Journal of Conchology" pageId="24" pageNumber="25" pagination="425 - 466" title="Land molluscs of Zanzibar Island (Unguja), Tanzania with the description of a new species of Gulella (Pulmonata: Streptaxidae)." volume="39" year="2007">Rowson (2007)</bibRefCitation>
are followed here in treating this species in
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Allopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allopeas" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Allopeas</taxonomicName>
as distinct from
<taxonomicName class="Gastropoda" family="Achatinidae" genus="Lamellaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lamellaxis" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Lamellaxis</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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