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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1023.61553" ID-GBIF-Dataset="b33816ea-cc6c-4ebe-b7cb-b70f2bb0775e" ID-PMC="PMC7973067" ID-Pensoft-Pub="1313-2970-1023-151" ID-Pensoft-UUID="5FE215C4917B5466ADD35FF62CFE5AD8" ID-PubMed="33776518" ID-ZooBank="14A62011A6484C6F91FA89EF25132191" ModsDocID="1313-2970-1023-151" checkinTime="1615560747517" checkinUser="pensoft" docAuthor="Yan, Weifeng, Shi, Hongliang, Liang, Hongbin &amp; Shi, Juan" docDate="2021" docId="886987A3A2E95F5DA34AF26F13793CE9" docLanguage="en" docName="ZooKeys 1023: 151-169" docOrigin="ZooKeys 1023" docPubDate="2021-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1023.61553" docTitle="Deltomerodes conaensis sp. nov." docType="treatment" docVersion="5" id="5FE215C4917B5466ADD35FF62CFE5AD8" lastPageNumber="151" masterDocId="5FE215C4917B5466ADD35FF62CFE5AD8" masterDocTitle="Two new species of the genus Deltomerodes Deuve, 1992 (Coleoptera, Carabidae, Patrobini) from Xizang, China" masterLastPageNumber="169" masterPageNumber="151" pageNumber="151" updateTime="1668149884416" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the genus Deltomerodes Deuve, 1992 (Coleoptera, Carabidae, Patrobini) from Xizang, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yan, Weifeng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7338-344X</mods:nameIdentifier>
<mods:affiliation>College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Juan</mods:namePart>
<mods:affiliation>College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier type="email">shi_juan@263.net</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1023</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>151</mods:start>
<mods:end>169</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1023.61553</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1023.61553</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1023-151</mods:identifier>
<mods:identifier type="ZooBank">14A62011A6484C6F91FA89EF25132191</mods:identifier>
<mods:identifier type="Pensoft-UUID">5FE215C4917B5466ADD35FF62CFE5AD8</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="151" type="taxonomy">
<treatment ID-GBIF-Taxon="180331276" LSID="urn:lsid:zoobank.org:act:52BCBE25-A05D-4FC4-A151-C81D2380673A" httpUri="http://treatment.plazi.org/id/886987A3A2E95F5DA34AF26F13793CE9" lastPageNumber="151" pageId="0" pageNumber="151">
<subSubSection pageId="0" pageNumber="151" type="nomenclature">
<paragraph pageId="0" pageNumber="151">
<taxonomicName LSID="http://zoobank.org/52BCBE25-A05D-4FC4-A151-C81D2380673A" authority="sp. nov." authorityName="sp. nov." class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">Deltomerodes conaensis sp. nov.</taxonomicName>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Deltomerodes conaensis sp. nov. (holotype). Habitus and labels (the white dots represent the setiferous pores on the left side of the specimen). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1023.61553.figure15" httpUri="https://binary.pensoft.net/fig/519161" pageId="0" pageNumber="151">Figs 15</figureCitation>
<figureCitation captionStart="Figures 16–19" captionStartId="F8" captionText="Figures 16 - 19. Deltomerodes conaensis sp. nov. (holotype) 16 first two segments of antenna 17 lateral side of fifth metatarsomere 18 labium 19 pronotum. Scale bars: 0.5 mm (a for 16, 17, b for 18, 19)." figureDoi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151">, 16-19</figureCitation>
<figureCitation captionStart="Figures 20–23" captionStartId="F9" captionText="Figures 20 - 23. Deltomerodes conaensis sp. nov. (holotype) 20 left paramere of aedeagus 21 right paramere of aedeagus 22 median lobe of aedeagus, left lateral view (22 A), dorsal view (22 B) 23 endophallus of aedeagus, left lateral view (23 A), dorsal view (23 B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151">, 20-23</figureCitation>
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">, 24-28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="type locality">
<paragraph pageId="0" pageNumber="151">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<materialsCitation ID-GBIF-Occurrence="3063627306" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="China" county="Cona County" elevation="5019" latitude="28.3037" location="Rama La" longLatPrecision="6" longitude="91.8834" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
:
<collectingCounty>Cona County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:886987A3A2E95F5DA34AF26F13793CE9:BEC21E7F8A50CEEC91FF6662DD6DE686" country="China" county="Cona County" latitude="28.3037" longLatPrecision="6" longitude="91.8834" name="Rama La" stateProvince="Xizang">Rama La</location>
pass (
<geoCoordinate degrees="28.3037" direction="north" orientation="latitude" precision="5" value="28.3037">28.3037°N</geoCoordinate>
,
<geoCoordinate degrees="91.8834" direction="east" orientation="longitude" precision="5" value="91.8834">91.8834°E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.019" unit="m" value="5019.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="5.019" unit="m" value="5019.0">5019 m</elevation>
</quantity>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="type material">
<paragraph pageId="0" pageNumber="151">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3063627301" collectingDate="2019-07-24" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="IZAS" collectorName="Paratypes" country="China" county="Lhoka" elevation="5019" latitude="28.3037" location="Quchomo Xiang" longLatPrecision="6" longitude="91.8834" municipality="Cona county" specimenCode="BJFU 2019" specimenCount="17" specimenCount-female="7" specimenCount-male="9" stateProvince="Xizang" typeStatus="holotype">
male (
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
), &quot;
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
,
<collectingCounty>Lhoka</collectingCounty>
pref.,
<collectingMunicipality>Cona county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:886987A3A2E95F5DA34AF26F13793CE9:35CCF23A7BB5859DAA55D3200A44F217" county="Lhoka" latitude="28.3037" longLatPrecision="6" longitude="91.8834" municipality="Cona county" name="Quchomo Xiang" stateProvince="Xizang">Quchomo Xiang</location>
,
<location LSID="urn:lsid:plazi:treatment:886987A3A2E95F5DA34AF26F13793CE9:2C0B0763AC18527AFF79544AB11FC7E0" county="Lhoka" latitude="28.3037" longLatPrecision="6" longitude="91.8834" municipality="Cona county" name="Rama La" stateProvince="Xizang">Rama La</location>
pass, alpine meadow,
<geoCoordinate degrees="28.3037" direction="north" orientation="latitude" precision="5" value="28.3037">28.3037°N</geoCoordinate>
,
<geoCoordinate degrees="91.8834" direction="east" orientation="longitude" precision="5" value="91.8834">91.8834°E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.019" unit="m" value="5019.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="5.019" unit="m" value="5019.0">5019m</elevation>
</quantity>
&quot;; &quot; under stone,
<date value="2019-07-24">
<collectingDate value="2019-07-24">2019. VII. 24</collectingDate>
</date>
, Shi HL, Yan WF &amp; Zhu PZ lgt., Exp.
<specimenCode collectionCode="BJFU">BJFU 2019</specimenCode>
&quot;; &quot;
<typeStatus>HOLOTYPE</typeStatus>
<specimenCount>♂</specimenCount>
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov., des. YAN &amp; SHI, 2021&quot; [red label].
<emphasis bold="true" italics="true" pageId="0" pageNumber="151">
<typeStatus>
<collectorName>Paratypes</collectorName>
</typeStatus>
</emphasis>
:
<specimenCount type="male">9 males</specimenCount>
and
<specimenCount type="female">7 females</specimenCount>
(
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
), same data as holotype but labeled as
<typeStatus>paratypes</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="diagnosis">
<paragraph pageId="0" pageNumber="151">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="151">The new species can be distinguished within the genus by the combination of: head generally with two pairs of setae between eyes and neck constriction; pronotum subcordate, lateral margins slightly sinuate before posterior angles, which are slightly pointed outwards; humeri distinct, rounded; scutellar pore present; elytral discal pores only present on third and seventh intervals; fifth meso- and metatarsomeres with three to four pairs of setae ventrally; endophallus consisting of a straight and weakly sclerotized proximal copulatory piece and two distal spiniform pieces.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="description">
<paragraph pageId="0" pageNumber="151">Description.</paragraph>
<paragraph pageId="0" pageNumber="151">
Habitus as in Figure
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Deltomerodes conaensis sp. nov. (holotype). Habitus and labels (the white dots represent the setiferous pores on the left side of the specimen). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1023.61553.figure15" httpUri="https://binary.pensoft.net/fig/519161" pageId="0" pageNumber="151">15</figureCitation>
. Medium-sized for a
<taxonomicName authorityName="Kirkby" authorityYear="1837" lsidName="" pageId="0" pageNumber="151" rank="tribe" tribe="Patrobini">Patrobini</taxonomicName>
species (L =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.06" metricValueMax="1.15" metricValueMin="0.97" unit="mm" value="10.6" valueMax="11.5" valueMin="9.7">9.7-11.5 mm</quantity>
; EW =
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" unit="mm" value="3.45" valueMax="3.7" valueMin="3.2">3.2-3.7 mm</quantity>
).
</paragraph>
<caption doi="10.3897/zookeys.1023.61553.figure15" httpUri="https://binary.pensoft.net/fig/519161" pageId="0" pageNumber="151" start="Figure 15" startId="F7">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 15.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>holotype</typeStatus>
). Habitus and labels (the white dots represent the setiferous pores on the left side of the specimen). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">General appearance</emphasis>
:
</emphasis>
Dorsal side dark brown to black, shiny, without metallic luster; head, mandibles, pronotum and elytra brown; antennae and legs a little paler, palpi reddish-brown; ventral side largely dark brown, abdominal sternum brown. Dorsal side smooth except for basal foveae of pronotum slightly punctate. Microsculpture invisible on head and pronotum, isodiametric on elytra, distinct in apical third, hardly visible near base.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Head</emphasis>
:
</emphasis>
Broad and ovate; surface smooth, without coarse punctures. Mandibles not elongated, right mandible with a weakly protruding tooth beneath basal inner margin; terminal maxillary palpomere broadest in middle; ligula with two adjacent apical setae. Antennae pubescent beginning with third antennomere; first antennomere with only one long seta; second antennomere glabrous except for apical ring of setae (Fig.
<figureCitation captionStart="Figures 16–19" captionStartId="F8" captionText="Figures 16 - 19. Deltomerodes conaensis sp. nov. (holotype) 16 first two segments of antenna 17 lateral side of fifth metatarsomere 18 labium 19 pronotum. Scale bars: 0.5 mm (a for 16, 17, b for 18, 19)." figureDoi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151">16</figureCitation>
). Eyes small, very slightly convex; temporae tumid, much longer than eye diameter; neck constriction shallower than in
<taxonomicName lsidName="D. ovicollis" pageId="0" pageNumber="151" rank="species" species="ovicollis">
<emphasis italics="true" pageId="0" pageNumber="151">D. ovicollis</emphasis>
</taxonomicName>
sp. nov. Frontal furrows short and rather shallow, extended to level of posterior edge of eyes, slightly divergent posteriorly; with two setae between eyes and neck constriction on each side (two specimens with three or four setae on one side), including one supraorbital seta near middle level of eyes, and one seta between posterior margins of eyes and neck constriction, not adjoining to neck constriction. Mentum with two deep longitudinal foveae basally, tooth narrow and bifid, apical notch shallow; submentum with two setae on each side in both sexes (Fig.
<figureCitation captionStart="Figures 16–19" captionStartId="F8" captionText="Figures 16 - 19. Deltomerodes conaensis sp. nov. (holotype) 16 first two segments of antenna 17 lateral side of fifth metatarsomere 18 labium 19 pronotum. Scale bars: 0.5 mm (a for 16, 17, b for 18, 19)." figureDoi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151">18</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151" start="Figures 16–19" startId="F8">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figures 16-19.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>holotype</typeStatus>
)
<emphasis bold="true" pageId="0" pageNumber="151">16</emphasis>
first two segments of antenna
<emphasis bold="true" pageId="0" pageNumber="151">17</emphasis>
lateral side of fifth metatarsomere
<emphasis bold="true" pageId="0" pageNumber="151">18</emphasis>
labium
<emphasis bold="true" pageId="0" pageNumber="151">19</emphasis>
pronotum. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(a for
<emphasis bold="true" pageId="0" pageNumber="151">16, 17</emphasis>
, b for
<emphasis bold="true" pageId="0" pageNumber="151">18, 19</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Pronotum</emphasis>
:
</emphasis>
Longitudinal-subcordate, rather narrow,
<collectionCode country="Thailand" name="Paleontological Collections">PW</collectionCode>
/
<collectionCode country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
= 1.00-1.15, slightly wider than head,
<collectionCode country="Thailand" name="Paleontological Collections">PW</collectionCode>
/HW = 1.11-1.21, widest near anterior third, fairly convex, moderately constricted posteriorly. Anterior margin near straight; anterior angles rounded, somewhat distinct, not protruding anteriorly; lateral margins broadly rounded in front, with slight sinuation before posterior angles; posterior margin nearly straight to slightly rounded; posterior angles obtuse to rectangular, apex sharp, slightly pointed outwards. Anterior transverse impression shallow and smooth; basal foveae shallow, finely punctate and wrinkled; disc smooth, median line shallow, obliterated at both extremities; lateral grooves narrow, not punctate. Lateral margins each with one seta before middle (two specimens with two setae on one side or both sides), another seta situated a little before posterior angle (Fig.
<figureCitation captionStart="Figures 16–19" captionStartId="F8" captionText="Figures 16 - 19. Deltomerodes conaensis sp. nov. (holotype) 16 first two segments of antenna 17 lateral side of fifth metatarsomere 18 labium 19 pronotum. Scale bars: 0.5 mm (a for 16, 17, b for 18, 19)." figureDoi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151">19</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Elytra</emphasis>
:
</emphasis>
Oblong-ovate, convex; EL/EW = 1.69-1.74, EW/
<collectionCode country="Thailand" name="Paleontological Collections">PW</collectionCode>
= 1.64-1.74, EL/
<collectionCode country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
= 2.80-2.84, widest near posterior third; humeri distinct, rounded, with small humeral tooth. Intervals slightly convex; inner striae well incised, finely punctate basally, lateral striae shallower; scutellar stria inconspicuous, only with one or two punctures and a superficial impression, scutellar pore present at base of second interval; third interval with three to four setiferous pores, all adjoining the third stria, the first one near basal sixth; seventh interval with one or two setiferous pores near basal sixth; umbilicate series composed of nine to twelve pores, basal ones not aggregated.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Ventral side</emphasis>
:
</emphasis>
Prosternum smooth, propleuron densely punctate; mesosternum and mesopleuron wrinkled, mesopleuron with sporadic coarse punctures; mesepimeron narrow, slightly widened laterally, suture separating mesepisternum and mesosternum joins anterior margin of mesepimeron; metepisternum rather long and narrow, not punctate. Lateral areas of abdominal sternites finely rugose, sternites IV to
<collectionCode country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
each with one pair of setae near middle; sternite VII with one seta on each side in males and two in females. Female sternite VIII with anterolateral apophyses long, with both longitudinal and transverse depigmentation distinct, posterointernal depigmentation present (Fig.
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">26</figureCitation>
). Female tergite VIII with anterolateral apophyses quite short; longitudinal carinae absent, transverse ones short and laterally placed, median sclerotization absent, basal depigmentation indistinct, apical depigmentation absent (Fig.
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">25</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Legs</emphasis>
:
</emphasis>
Males with basal two protarsomeres slightly expanded, second protarsomere distinctly wider than third, which is near triangular; fourth protarsomere slightly bilobed; metatrochanter normal, apex not protruding; all tarsomeres pubescent dorsally; fifth meso- and metatarsomeres with three or four pairs of setae ventrally (Fig.
<figureCitation captionStart="Figures 16–19" captionStartId="F8" captionText="Figures 16 - 19. Deltomerodes conaensis sp. nov. (holotype) 16 first two segments of antenna 17 lateral side of fifth metatarsomere 18 labium 19 pronotum. Scale bars: 0.5 mm (a for 16, 17, b for 18, 19)." figureDoi="10.3897/zookeys.1023.61553.figures16-19" httpUri="https://binary.pensoft.net/fig/519162" pageId="0" pageNumber="151">17</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Male genitalia</emphasis>
:
</emphasis>
Median lobe of aedeagus strongly bent at base, gutter-shaped and opened dorsally; in lateral view apical lamella moderately curved dorsally; in dorsal view apical lamella gradually attenuated towards apex and faintly twisted rightwards, apex rounded, forming a prominent hook to the right (Fig.
<figureCitation captionStart="Figures 20–23" captionStartId="F9" captionText="Figures 20 - 23. Deltomerodes conaensis sp. nov. (holotype) 20 left paramere of aedeagus 21 right paramere of aedeagus 22 median lobe of aedeagus, left lateral view (22 A), dorsal view (22 B) 23 endophallus of aedeagus, left lateral view (23 A), dorsal view (23 B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151">22</figureCitation>
). Armature of endophallus consisting of a straight and weakly sclerotized proximal copulatory piece and two spiniform distal pieces, the dorsal one large and stout, the left one much narrower (Fig.
<figureCitation captionStart="Figures 20–23" captionStartId="F9" captionText="Figures 20 - 23. Deltomerodes conaensis sp. nov. (holotype) 20 left paramere of aedeagus 21 right paramere of aedeagus 22 median lobe of aedeagus, left lateral view (22 A), dorsal view (22 B) 23 endophallus of aedeagus, left lateral view (23 A), dorsal view (23 B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151">23</figureCitation>
). Left paramere larger than right one, both short and gradually contracted towards apex but the left one more truncated at apex, both with two long apical setae and one or two short subapical setae (Figs
<figureCitation captionStart="Figures 20–23" captionStartId="F9" captionText="Figures 20 - 23. Deltomerodes conaensis sp. nov. (holotype) 20 left paramere of aedeagus 21 right paramere of aedeagus 22 median lobe of aedeagus, left lateral view (22 A), dorsal view (22 B) 23 endophallus of aedeagus, left lateral view (23 A), dorsal view (23 B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151">20</figureCitation>
,
<figureCitation captionStart="Figures 20–23" captionStartId="F9" captionText="Figures 20 - 23. Deltomerodes conaensis sp. nov. (holotype) 20 left paramere of aedeagus 21 right paramere of aedeagus 22 median lobe of aedeagus, left lateral view (22 A), dorsal view (22 B) 23 endophallus of aedeagus, left lateral view (23 A), dorsal view (23 B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151">21</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1023.61553.figures20-23" httpUri="https://binary.pensoft.net/fig/519163" pageId="0" pageNumber="151" start="Figures 20–23" startId="F9">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figures 20-23.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>holotype</typeStatus>
)
<emphasis bold="true" pageId="0" pageNumber="151">20</emphasis>
left paramere of aedeagus
<emphasis bold="true" pageId="0" pageNumber="151">21</emphasis>
right paramere of aedeagus
<emphasis bold="true" pageId="0" pageNumber="151">22</emphasis>
median lobe of aedeagus, left lateral view (
<emphasis bold="true" pageId="0" pageNumber="151">22A</emphasis>
), dorsal view (
<emphasis bold="true" pageId="0" pageNumber="151">22B</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="151">23</emphasis>
endophallus of aedeagus, left lateral view (
<emphasis bold="true" pageId="0" pageNumber="151">23A</emphasis>
), dorsal view (
<emphasis bold="true" pageId="0" pageNumber="151">23B</emphasis>
). Scale bar:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">
<emphasis italics="true" pageId="0" pageNumber="151">Female genitalia</emphasis>
:
</emphasis>
Gonocoxite I glabrous; gonocoxite II narrow and slightly curved, blunt-rounded at apex, with two small setae in one fovea at subapical inner margin (Fig.
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">28</figureCitation>
). Reproductive tract with well-marked annular sclerotized ring about
<quantity metricMagnitude="-4" metricUnit="m" metricValue="3.5" unit="mm" value="0.35">0.35 mm</quantity>
in diameter on bursa copulatrix; spermatheca tubular, very small, without spermathecal duct and spermathecal gland (Figs
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">24</figureCitation>
,
<figureCitation captionStart="Figures 24–28" captionStartId="F10" captionText="Figures 24 - 28. Deltomerodes conaensis sp. nov. (paratype female) 24 female reproductive tract 25 female tergite VIII 26 female sternite VIII 27 spermatheca 28 gonocoxa, right lateral view (28 A), dorsal view (28 B). Scale bars: 0.5 mm (a for 24 - 26, b for 27, 28)." figureDoi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151">27</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1023.61553.figures24-28" httpUri="https://binary.pensoft.net/fig/519164" pageId="0" pageNumber="151" start="Figures 24–28" startId="F10">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figures 24-28.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>paratype</typeStatus>
female)
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="151">24</emphasis>
female
</specimenCount>
reproductive tract
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="151">25</emphasis>
female
</specimenCount>
tergite VIII
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="151">26</emphasis>
female
</specimenCount>
sternite VIII
<emphasis bold="true" pageId="0" pageNumber="151">27</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="151">28</emphasis>
gonocoxa, right lateral view (
<emphasis bold="true" pageId="0" pageNumber="151">28A</emphasis>
), dorsal view (
<emphasis bold="true" pageId="0" pageNumber="151">28B</emphasis>
). Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(a for
<emphasis bold="true" pageId="0" pageNumber="151">24-26</emphasis>
, b for
<emphasis bold="true" pageId="0" pageNumber="151">27, 28</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="distribution">
<paragraph pageId="0" pageNumber="151">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="151">
<materialsCitation ID-GBIF-Occurrence="3063627302" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" county="Rama La in Cona County" location="Rama La in Cona County" specimenCount="1" stateProvince="Xizang">
This species is known only from the
<typeStatus>type</typeStatus>
locality of
<collectingCounty>Rama La in Cona County</collectingCounty>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
(Map
<figureCitation captionStart="Map 3" captionStartId="F11" captionText="Map 3. Distribution map for Deltomerodes conaensis sp. nov. (red)." figureDoi="10.3897/zookeys.1023.61553.map3" httpUri="https://binary.pensoft.net/fig/519165" pageId="0" pageNumber="151">3</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1023.61553.map3" httpUri="https://binary.pensoft.net/fig/519165" pageId="0" pageNumber="151" start="Map 3" startId="F11">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Map 3.</emphasis>
Distribution map for
<taxonomicName class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes conaensis" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes conaensis</emphasis>
</taxonomicName>
sp. nov. (red).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="etymology">
<paragraph pageId="0" pageNumber="151">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="151">
The scientific name of the new species refers to its
<typeStatus>type</typeStatus>
locality in Cona County.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="remarks">
<paragraph pageId="0" pageNumber="151">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="151">
According to the key provided by
<bibRefCitation author="Zamotajlov, AS" journalOrPublisher="In: Zamotajlov A, Sciaky R (Eds) Advances in Carabidology. Papers Dedicated to the Memory of Prof. Dr. Oleg L. Kryzhanovskij. Krasnodar: MUISO Publishers" pageId="0" pageNumber="151" refId="B11" refString="Zamotajlov, AS, 1999. Redefinition of the genus Deltomerodes Deuve, 1992, with the description of new species (ColeopteraCarabidaePatrobinae). In: Zamotajlov A, Sciaky R (Eds) Advances in Carabidology. Papers Dedicated to the Memory of Prof. Dr. Oleg L. Kryzhanovskij. Krasnodar: MUISO Publishers" title="Redefinition of the genus Deltomerodes Deuve, 1992, with the description of new species (ColeopteraCarabidaePatrobinae)." year="1999">Zamotajlov (1999)</bibRefCitation>
, the new species belongs to the
<taxonomicName authorityName="Zamotajlov" authorityYear="1999" class="Insecta" family="Carabidae" genus="Deltomerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltomerodes murzini" order="Coleoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="murzini">
<emphasis italics="true" pageId="0" pageNumber="151">Deltomerodes murzini</emphasis>
</taxonomicName>
-group for some characters: (1) head with less than four pairs of setae between eyes and neck constriction; (2) pronotum not cordate, without distinct sinuation before posterior angles, sides with one seta in anterior third; (3) fifth meso- and metatarsomeres with three or four pairs of setae ventrally. Within this species group, the new species seems to be closest to
<taxonomicName lsidName="D. wrasei" pageId="0" pageNumber="151" rank="species" species="wrasei">
<emphasis italics="true" pageId="0" pageNumber="151">D. wrasei</emphasis>
</taxonomicName>
for their adjacent distribution and morphological similarities in chaetotaxy, pronotum shape and in the apical lamella of the aedeagus. However, the armature of the endophallus in
<taxonomicName lsidName="D. conaensis" pageId="0" pageNumber="151" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="151">D. conaensis</emphasis>
</taxonomicName>
sp. nov. has a straight and weakly sclerotized proximal copulatory piece. The new species is different from all other known species of the genus, in which the proximal copulatory piece is well-sclerotized and loop-like.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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