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<document id="EB063824A490169FA6FA5D733C9B101E" ID-CLB-Dataset="21799" ID-DOI="10.5852/ejt.2020.653" ID-GBIF-Dataset="a2776578-f968-442b-8317-9599e5884e3f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3862477" ID-ZooBank="9E6EB3E1-E20F-48CB-BDD4-6859FC472382" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590670836830" checkinUser="valdenar" docAuthor="Wichard, Wilfried, Espeland, Marianne, Müller, Patrick &amp; Wang, Bo" docDate="2020" docId="8005B657FFCBFF8B6C154148FECEF824" docLanguage="en" docName="ejt-653_wichard_espeland_mueller.pdf.imf" docOrigin="European Journal of Taxonomy 653" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Palaeopsilotreta cretacea Wichard &amp; Espeland &amp; Müller &amp; Wang 2020, sp. nov." docType="treatment" docUuid="DD288746-197D-4F8E-A45E-75D1330D1C45" docUuidSource="ZooBank" docVersion="6" lastPageNumber="9" masterDocId="7C3CCE2FFFC3FF836E444043FA51FFCF" masterDocTitle="New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1698841811242" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5A9CBA84F91EDAB90E3E22544008ED67">New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae)</mods:title>
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<mods:name id="6CA1B3DCC3783EED0D2B386E3558B982" type="personal">
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<mods:namePart id="8474C39A55AAD7A1522302D22DAC5F7E">Wichard, Wilfried</mods:namePart>
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<mods:affiliation id="0575A7C1735E59237174354B20F992BC">Institute of Biology and its Didactics, University of Cologne, Herbert Lewinstrasse 2, 50931 Köln, Germany. &amp; Email: Wichard @ uni-koeln. de &amp; urn: lsid: zoobank. org: author: 1 E 8601 B 2 - A 8 F 8 - 44 A 6 - B 185 - 4 FB 0 B 2 E 5219 E</mods:affiliation>
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<mods:namePart id="C680183C68A4ED214A8F89525514CBDC">Espeland, Marianne</mods:namePart>
<mods:nameIdentifier id="D899070DBEAB27B8E4F0B330337D2671" type="LSID">00D6F9F9-3902-4A8B-846F-720AB32922A6</mods:nameIdentifier>
<mods:affiliation id="E79E8DCE3306D9CEF8C93EC95BEFEC41">Arthropoda Department, Zoological Research Museum Alexander Koenig, Adenauer-Allee 160, 53113 Bonn, Germany. &amp; urn: lsid: zoobank. org: author: 00 D 6 F 9 F 9 - 3902 - 4 A 8 B- 846 F- 720 AB 32922 A 6 &amp; Corresponding author: M. Espeland @ leibniz-zfmk. de</mods:affiliation>
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<mods:namePart id="81DD7304CA28C2CCF0E0D58418A261E8">Müller, Patrick</mods:namePart>
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<mods:affiliation id="BBF8692DAB86FAC2E5BB5CA23446A4D1">Friedhofstrasse 9, 66894 Kashofen, Germany. &amp; Email: pat 14789 @ web. de &amp; urn: lsid: zoobank. org: author: CFA 297 B 2 - 35 D 2 - 4 C 05 - 8 F 60 - D 73 EBF 620 B 5 A</mods:affiliation>
<mods:nameIdentifier id="EC44DCCF6009FFA974E3E478B6240A0D" type="email">pat14789@web.de</mods:nameIdentifier>
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<mods:namePart id="5F08A891902B4E501FB7FAFEF6DE93A3">Wang, Bo</mods:namePart>
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<mods:affiliation id="AAAF2828FB5E6DC8B642655EAB73508D">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. &amp; Email: bowang @ nigpas. ac. cn &amp; urn: lsid: zoobank. org: author: 7 CBE 9 A 76 - 16 C 4 - 4 D 6 D- 9 B 4 D-B 2 BEC 000 ADDF</mods:affiliation>
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<mods:title id="97CB6A4646E40AC994B869A3DFBEB8EB">European Journal of Taxonomy</mods:title>
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<mods:part id="9192F78E0A3453B26697606181242063">
<mods:date id="68EE4C3BDDBC19D3B74001F5B173C451">2020</mods:date>
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<mods:number id="991C54659E95ABB5591E41D23E62C8F9">2020-05-27</mods:number>
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<treatment id="8005B657FFCBFF8B6C154148FECEF824" ID-DOI="http://doi.org/10.5281/zenodo.3864328" ID-GBIF-Taxon="164363914" ID-Zenodo-Dep="3864328" LSID="urn:lsid:zoobank.org:act:DD288746-197D-4F8E-A45E-75D1330D1C45" httpUri="http://treatment.plazi.org/id/8005B657FFCBFF8B6C154148FECEF824" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="40B654CAFFCBFF8B6C154148F9B2FEE9" box="[593,995,267,294]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="08130741FFCBFF8B6C154148F9B2FEE9" blockId="8.[367,1221,267,364]" box="[593,995,267,294]" pageId="8" pageNumber="9">
<heading id="535BB02DFFCBFF8B6C154148F9B2FEE9" box="[593,995,267,294]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="CFAC7CC2FFCBFF8B6C154148F9D0FEE9" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[593,897,267,294]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cretacea" status="sp. nov.">
<emphasis id="3AD8DB53FFCBFF8B6C154148F9D0FEE9" bold="true" box="[593,897,267,294]" italics="true" pageId="8" pageNumber="9">Palaeopsilotreta cretacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCBFF8B6DCC414FF9B2FEE9" box="[904,995,268,294]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="40B654CAFFCBFF8B6F2B416CF96DFEA3" pageId="8" pageNumber="9" type="description">
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<uri id="7C3D0B43FFCBFF8B6F2B416CFE94FE85" box="[367,1221,303,330]" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="7C0A3D94FFCBFF8B6CCA416CFE94FE85" box="[654,1221,303,330]" pageId="8" pageNumber="9">DD288746-197D-4F8E-A45E-75D1330D1C45</uuid>
</uri>
</paragraph>
<paragraph id="08130741FFCBFF8B6CB34111F96DFEA3" blockId="8.[367,1221,267,364]" box="[759,828,338,364]" pageId="8" pageNumber="9">
<figureCitation id="90971BC4FFCBFF8B6CB34111F96DFEA3" box="[759,828,338,364]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[219,1369,265,1891]" captionTargetId="figure@9.[219,1369,265,1891]" captionTargetPageId="9" captionText="Fig. 3. Palaeopsilotreta cretacea sp. nov. (ZFMK-TRI000814) A. Holotype male in dorsal view. B. Head with bipectinate antennae, maxillary palps and labial palps. C. Drawing of a forewing. D. Drawing of the male genitalia. E. Male genitalia in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3862487" httpUri="https://zenodo.org/record/3862487/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCBFF8B6EF941D7F899FD41" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="08130741FFCBFF8B6EF941D7FB67FE61" blockId="8.[189,310,404,430]" box="[189,310,404,430]" pageId="8" pageNumber="9">
<heading id="535BB02DFFCBFF8B6EF941D7FB67FE61" bold="true" box="[189,310,404,430]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="3AD8DB53FFCBFF8B6EF941D7FB67FE61" bold="true" box="[189,310,404,430]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCBFF8B6EF94180F899FD41" blockId="8.[189,1399,450,654]" pageId="8" pageNumber="9">
Antennae of males with 12 bipectinate flagellomeres and last two flagellomeres simple, not bipectinate. In male forewing, media branched in M1+2 and M3+4. In male genitalia, harpago apically with two clear thorn-like black spines.
<taxonomicName id="CFAC7CC2FFCBFF8B6C60424AF904FDEC" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[548,853,521,547]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cretacea" status="sp. nov.">
<emphasis id="3AD8DB53FFCBFF8B6C60424AF904FDEC" box="[548,853,521,547]" italics="true" pageId="8" pageNumber="9">Palaeopsilotreta cretacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCBFF8B6D1B4249F9ECFDEB" box="[863,957,522,548]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is clearly distinguished from
<taxonomicName id="CFAC7CC2FFCBFF8B6B6E4249FF27FDEC" authorityName="Wichard &amp; Wang" authorityYear="2017" box="[1322,1398,522,547]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xiai">
<emphasis id="3AD8DB53FFCBFF8B6B6E4249FF27FDEC" box="[1322,1398,522,547]" italics="true" pageId="8" pageNumber="9">P. xiai</emphasis>
</taxonomicName>
and
<taxonomicName id="CFAC7CC2FFCBFF8B6EB4426DFBDFFD88" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[240,398,557,583]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="burmanica" status="sp. nov.">
<emphasis id="3AD8DB53FFCBFF8B6EB4426DFBDFFD88" box="[240,398,557,583]" italics="true" pageId="8" pageNumber="9">P. burmanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCBFF8B6FD3426EFBA2FD88" box="[407,499,557,583]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
by the branched media in forewings and by the harpago at an angle of 90° to the coxopodite.
<taxonomicName id="CFAC7CC2FFCBFF8B6FDC4213F8DEFDA5" authorityName="Wichard &amp; Wang" authorityYear="2017" box="[408,655,592,618]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xiai">
<emphasis id="3AD8DB53FFCBFF8B6FDC4213F8DEFDA5" box="[408,655,592,618]" italics="true" pageId="8" pageNumber="9">Palaeopsilotreta xiai</emphasis>
</taxonomicName>
and
<taxonomicName id="CFAC7CC2FFCBFF8B6C8C4212F934FDA5" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[712,869,592,618]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="burmanica" status="sp. nov.">
<emphasis id="3AD8DB53FFCBFF8B6C8C4212F934FDA5" box="[712,869,592,618]" italics="true" pageId="8" pageNumber="9">P. burmanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCBFF8B6D284212F996FDA4" box="[876,967,593,619]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
have the media unbranched, and the harpago and coxopodite form a straight unit.
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCBFF8B6EF942F5FE34FD31" box="[189,1125,694,767]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="08130741FFCBFF8B6EF942F5FB15FD1F" blockId="8.[189,324,694,720]" box="[189,324,694,720]" pageId="8" pageNumber="9">
<heading id="535BB02DFFCBFF8B6EF942F5FB15FD1F" bold="true" box="[189,324,694,720]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="3AD8DB53FFCBFF8B6EF942F5FB15FD1F" bold="true" box="[189,324,694,720]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCBFF8B6EF942A7FE34FD31" blockId="8.[189,1125,740,767]" box="[189,1125,740,767]" pageId="8" pageNumber="9">
<taxonomicName id="CFAC7CC2FFCBFF8B6EF942A7FBBAFD31" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[189,491,740,766]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cretacea" status="sp. nov.">
<emphasis id="3AD8DB53FFCBFF8B6EF942A7FBBAFD31" box="[189,491,740,766]" italics="true" pageId="8" pageNumber="9">Palaeopsilotreta cretacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCBFF8B6FB642A6F81CFD30" box="[498,589,741,767]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is named for the geological age of the amber.
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCBFF8B6EF94365FB22FC33" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="08130741FFCBFF8B6EF94365FBFBFC8F" blockId="8.[189,426,806,832]" box="[189,426,806,832]" pageId="8" pageNumber="9">
<heading id="535BB02DFFCBFF8B6EF94365FBFBFC8F" bold="true" box="[189,426,806,832]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="3AD8DB53FFCBFF8B6EF94365FBFBFC8F" bold="true" box="[189,426,806,832]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCBFF8B6EAE4316FB0DFCA0" blockId="8.[189,1399,853,1020]" box="[234,348,853,879]" pageId="8" pageNumber="9">
<emphasis id="3AD8DB53FFCBFF8B6EAE4316FB0DFCA0" bold="true" box="[234,348,853,879]" pageId="8" pageNumber="9">
<typeStatus id="D717B9E3FFCBFF8B6EAE4316FB0DFCA0" box="[234,348,853,879]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="08130741FFCBFF8B6EF9433BFB22FC33" blockId="8.[189,1399,853,1020]" pageId="8" pageNumber="9">
<materialsCitation id="B8C40D1CFFCBFF8B6EF9433BFB22FC33" ID-GBIF-Occurrence="2628167304" collectionCode="BUB" country="Myanmar" location="Kachin State" pageId="8" pageNumber="9" specimenCode="BUB 2883, ZFMK-TRI000814" specimenCount="1" stateProvince="Kachin State" typeStatus="holotype">
<collectingCountry id="70BB47D1FFCBFF8B6EF9433BFB07FC5D" box="[189,342,888,914]" name="Myanmar" pageId="8" pageNumber="9">MYANMAR</collectingCountry>
<emphasis id="3AD8DB53FFCBFF8B6F31433BF87BFC5D" bold="true" box="[373,554,888,914]" pageId="8" pageNumber="9">
<collectingRegion id="CA68C9A3FFCBFF8B6F31433BF849FC5D" box="[373,536,888,914]" country="Myanmar" name="Kachin" pageId="8" pageNumber="9">Kachin State</collectingRegion>
</emphasis>
<specimenCount id="1EAACCC8FFCBFF8B6C76433BF81BFC5D" box="[562,586,888,914]" pageId="8" pageNumber="9" type="male"></specimenCount>
; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; former Patrick Müller collection,
<specimenCode id="580AAF3AFFCBFF8B6C1243DFF88AFC7A" box="[598,731,923,950]" pageId="8" pageNumber="9">BUB 2883</specimenCode>
;
<specimenCode id="580AAF3AFFCBFF8B6CAE43D8F99CFC7A" box="[746,973,923,949]" pageId="8" pageNumber="9">ZFMK-TRI000814</specimenCode>
. Body well preserved; forewing in dorsal view, hind wing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCBFF8B6EF94467FECEF824" pageId="8" pageNumber="9" type="description">
<paragraph id="08130741FFCBFF8B6EF94467FB1FFBF1" blockId="8.[189,334,1060,1086]" box="[189,334,1060,1086]" pageId="8" pageNumber="9">
<heading id="535BB02DFFCBFF8B6EF94467FB1FFBF1" bold="true" box="[189,334,1060,1086]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="3AD8DB53FFCBFF8B6EF94467FB1FFBF1" bold="true" box="[189,334,1060,1086]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCBFF8B6EF94410F8D5FAF0" blockId="8.[189,1399,1106,1343]" pageId="8" pageNumber="9">
HEAD. Light brown. Antennae partially bipectinate, consisting of long scapus, short pedicellus, and first flagellomere as long as pedicellus, both together about as long as scapus, followed by 12 successive bipectinate flagellomeres, and, finally, 2 simple flagellomeres without rami (
<figureCitation id="90971BC4FFCBFF8B6A7644DAFEDCFB7B" box="[1074,1165,1177,1204]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[219,1369,265,1891]" captionTargetId="figure@9.[219,1369,265,1891]" captionTargetPageId="9" captionText="Fig. 3. Palaeopsilotreta cretacea sp. nov. (ZFMK-TRI000814) A. Holotype male in dorsal view. B. Head with bipectinate antennae, maxillary palps and labial palps. C. Drawing of a forewing. D. Drawing of the male genitalia. E. Male genitalia in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3862487" httpUri="https://zenodo.org/record/3862487/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
). Antennae about as long as forewings. Maxillary palps 5-segmented and labial palps 3-segmented, their terminal segments not flexible or annulate. Head dorsal ocelli absent, probably with interantennal, postantennal and posterior setose warts, but not well preserved and hard to recognize; scutum slightly macerated, scutellum shows single dome-shaped, large, ovoid wart.
</paragraph>
<paragraph id="08130741FFCBFF8B6EF94524FB62F975" blockId="8.[189,1399,1382,1723]" pageId="8" pageNumber="9">
FOREWINGS (
<figureCitation id="90971BC4FFCBFF8B6F0F4525FBF7FA4F" box="[331,422,1382,1409]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[219,1369,265,1891]" captionTargetId="figure@9.[219,1369,265,1891]" captionTargetPageId="9" captionText="Fig. 3. Palaeopsilotreta cretacea sp. nov. (ZFMK-TRI000814) A. Holotype male in dorsal view. B. Head with bipectinate antennae, maxillary palps and labial palps. C. Drawing of a forewing. D. Drawing of the male genitalia. E. Male genitalia in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3862487" httpUri="https://zenodo.org/record/3862487/files/figure.png" pageId="8" pageNumber="9">Fig. 3C</figureCitation>
). Light brown, translucent,
<quantity id="CF54AAA4FFCBFF8B6CAE4525F915FA4E" box="[746,836,1382,1409]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="9" unit="mm" value="2.6">2.6 mm</quantity>
long. Radius R1 running straight to apical wing margin. Forks I and II of Rs present, sessile, branching at same level. Discoidal cell closed by crossveins, DC smaller, about half as long as fork I, fork II. Media branching into M1+2 and M3+4. Fork M longer than fork I and fork II, reaching up to half length of discoidal cell. Medial and thyridial cells absent. Cu1 running straight, apically bifurcated into fork V (Cu1a Cu1b); Cu2 simple, not forked, so far as visible, probably with crossvein to Cu1b. Anal veins probably complete. Hindwings light brown, translucent, relatively narrow, smaller than forewings, about
<quantity id="CF54AAA4FFCBFF8B6CA94674F917F99D" box="[749,838,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="9" unit="mm" value="2.1">2.1 mm</quantity>
long, so far as visible, venation probably similar to forewing venation. Radius R1 straight, forks I, II present; M probably simple, without fork; Cu1 with relatively long fork V, Cu2 simple. [Generally, crossveins in hyaline wings are difficult or impossible to detect].
</paragraph>
<paragraph id="08130741FFCBFF8B6EF946A1FBCFF934" blockId="8.[189,414,1761,1788]" box="[189,414,1761,1788]" pageId="8" pageNumber="9">TIBIAL SPURS. 2/4/4.</paragraph>
<paragraph id="08130741FFCBFF8B6EF94760FECEF824" blockId="8.[189,1399,1826,2027]" pageId="8" pageNumber="9">
MALE GENITALIA.Visible only in ventral view (
<figureCitation id="90971BC4FFCBFF8B6CF94761F8AFF8F3" box="[701,766,1826,1852]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[219,1369,265,1891]" captionTargetId="figure@9.[219,1369,265,1891]" captionTargetPageId="9" captionText="Fig. 3. Palaeopsilotreta cretacea sp. nov. (ZFMK-TRI000814) A. Holotype male in dorsal view. B. Head with bipectinate antennae, maxillary palps and labial palps. C. Drawing of a forewing. D. Drawing of the male genitalia. E. Male genitalia in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3862487" httpUri="https://zenodo.org/record/3862487/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
DE): Inferior appendages(gonocoxites)2-segmented. Coxopodites strong and slightly angular, broad at base, tapering towards apex, slightly curved towards genital centre, flanks more curved mesally than laterally. Harpago almost at right angle to coxopodites, directed centrad, rod-shaped, black, less than ¼ as long as coxopodites and much smaller in volume; apical end of harpago not clearly visible. Segment X apparently covering central phallic apparatus dorsally with 2 oblong scales. Lateral preanal appendages long, slightly bent inward.
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</subSubSection>
</treatment>
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