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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-PMC="PMC7644702" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-PubMed="33199958" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro &amp; Ablett, Jonathan D." docDate="2020" docId="6D49DDFD5AFB5E4297DB32A82DBDCC8D" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Metalycaeus muciferus" docType="treatment" docVersion="6" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1668169683351" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sajan, Sheikh</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tripathy, Basudev</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asami, Takahiro</mods:namePart>
<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>981</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>220</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.981.53583</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.981.53583</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-981-1</mods:identifier>
<mods:identifier type="ZooBank">5194AAC86B8A473F8A41470A60182A0B</mods:identifier>
<mods:identifier type="Pensoft-UUID">7C44C797C4125A71BAE032A55E6FA5DC</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168715123" LSID="urn:lsid:plazi:treatment:6D49DDFD5AFB5E4297DB32A82DBDCC8D" httpUri="http://treatment.plazi.org/id/6D49DDFD5AFB5E4297DB32A82DBDCC8D" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="6D49DDFD-5AFB-5E42-97DB-32A82DBDCC8D" authority="(Heude, 1885)" baseAuthorityName="Heude" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus muciferus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus">Metalycaeus muciferus (Heude, 1885)</taxonomicName>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Alycaeus expansus Heude, 1890, syntype (HMT- 215 a; synonym of Metalycaeus muciferus (Heude, 1885 )). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure33" httpUri="https://binary.pensoft.net/fig/468678" pageId="0" pageNumber="1">Fig. 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycoeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Alycoeus</taxonomicName>
[sic]
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muciferus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus">muciferus</taxonomicName>
Heude, 1885: 96, pl. 24, figs 1, 1a.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) muciferus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus" subGenus="Orthalycaeus">Alycaeus (Orthalycaeus) muciferus</taxonomicName>
-
<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="156 - 210" publicationUrl="https://www.biodiversitylibrary.org/page/16355855" refId="B134" refString="Moellendorff, OF von, 1886. Materialen zur Fauna for China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 13: 156 - 210, https://www.biodiversitylibrary.org/page/16355855" title="Materialen zur Fauna for China." url="https://www.biodiversitylibrary.org/page/16355855" volume="13" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
: 171.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gredler" authorityYear="1887" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus helicodes" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="helicodes">Alycaeus helicodes</taxonomicName>
Gredler, 1888: 365.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansus">Alycaeus expansus</taxonomicName>
Heude, 1890: 129, pl. 38, fig. 2. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) muciferus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) muciferus</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 149;
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 359.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus muciferus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus">Metalycaeus muciferus</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 88-93, figs 58, 59, 60, 61 (
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="helicodes" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="helicodes">helicodes</taxonomicName>
Gredler is a synonym).
</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;
<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
&quot;
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="expansus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansus">expansus</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 105, fig. 69L.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;in ditione
<normalizedToken originalValue="Tchen-k’eou”">Tchen-k'eou&quot;</normalizedToken>
(
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycoeus muciferus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycoeus muciferus</emphasis>
</taxonomicName>
); Pe-shang, Prov. Hunan (
<taxonomicName lsidName="A. helicodes" pageId="0" pageNumber="1" rank="species" species="helicodes">
<emphasis italics="true" pageId="0" pageNumber="1">A. helicodes</emphasis>
</taxonomicName>
); &quot;Tchen
<normalizedToken originalValue="K’eou”">K'eou&quot;</normalizedToken>
(
<taxonomicName lsidName="A. expansus" pageId="0" pageNumber="1" rank="species" species="expansus">
<emphasis italics="true" pageId="0" pageNumber="1">A. expansus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
China,
<normalizedToken originalValue="Tchen-k’eou">Tchen-k'eou</normalizedToken>
, MCZ 167230 (7 syntypes of
<taxonomicName lsidName="A. muciferus" pageId="0" pageNumber="1" rank="species" species="muciferus">
<emphasis italics="true" pageId="0" pageNumber="1">A. muciferus</emphasis>
</taxonomicName>
);
<normalizedToken originalValue="Tchen-k’eou">Tchen-k'eou</normalizedToken>
, China, Pe-shang, Hunan, leg. Fuchs, coll. Gredler, SMF 192225 (2 paralectotypes of
<taxonomicName lsidName="A. helicodes" pageId="0" pageNumber="1" rank="species" species="helicodes">
<emphasis italics="true" pageId="0" pageNumber="1">A. helicodes</emphasis>
</taxonomicName>
, lectotype in the Franziskanergymnasium Bozen [Bolzano, Italy], designated by
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 228" refId="B233" refString="Zilch, A, 1974. Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol. Archiv fuer Molluskenkunde 104: 171 - 228" title="Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol." volume="104" year="1974">Zilch 1974</bibRefCitation>
); HMT-215a, syntype: labelled as lectotype of
<taxonomicName lsidName="A. expansus" pageId="0" pageNumber="1" rank="species" species="expansus">
<emphasis italics="true" pageId="0" pageNumber="1">A. expansus</emphasis>
</taxonomicName>
deposited in IZCAS:
<normalizedToken originalValue="Tchen-k’eou">Tchen-k'eou</normalizedToken>
(Cheng-kou County, Chong-qing, China) (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Alycaeus expansus Heude, 1890, syntype (HMT- 215 a; synonym of Metalycaeus muciferus (Heude, 1885 )). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure33" httpUri="https://binary.pensoft.net/fig/468678" pageId="0" pageNumber="1">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">Protoconch conspicuously elevated from dorsal surface, its end with inconspicuous spiral striations; R1 with irregular, widely spaced ribs and fine spiral sculpture; R2 with dense, sharp, elevated ribs.</paragraph>
<paragraph pageId="0" pageNumber="1">
No type specimens of
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus expansus</emphasis>
</taxonomicName>
were reported in American museums (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.141074" author="Johnson, RJ" journalOrPublisher="Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University)" pageId="0" pageNumber="1" refId="B92" refString="Johnson, RJ, 1973. Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University), DOI: https://doi.org/10.5962/bhl.title.141074" title="Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude." url="https://doi.org/10.5962/bhl.title.141074" year="1973">Johnson 1973</bibRefCitation>
). We were not able to examine the types deposited in the Beijing Museum before the revision of Chinese
<taxonomicName authorityName="Blanford" authorityYear="1864" family="Alycaeidae" lsidName="" pageId="0" pageNumber="1" rank="family">Alycaeidae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
), but could do so now. The general shell shape, colour, sculpture, and the lengths and proportions of R2 and R3 agree with those of
<taxonomicName lsidName="A. muciferus" pageId="0" pageNumber="1" rank="species" species="muciferus">
<emphasis italics="true" pageId="0" pageNumber="1">A. muciferus</emphasis>
</taxonomicName>
, which was described from the same type locality. Consequently, we move
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus expansus</emphasis>
</taxonomicName>
to the synonymy of
<taxonomicName lsidName="A. muciferus" pageId="0" pageNumber="1" rank="species" species="muciferus">
<emphasis italics="true" pageId="0" pageNumber="1">A. muciferus</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.981.53583.figure33" httpUri="https://binary.pensoft.net/fig/468678" pageId="0" pageNumber="1" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 33.</emphasis>
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="expansus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus expansus</emphasis>
</taxonomicName>
Heude, 1890, syntype (HMT-215a; synonym of
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1885" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus muciferus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muciferus">
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus muciferus</emphasis>
</taxonomicName>
(Heude, 1885)). Photographs: Kaibaryer Meng.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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