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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1132.87364" ID-GBIF-Dataset="eab928e7-32a6-4584-a351-7457a103c5b0" ID-Pensoft-Pub="1313-2970-1132-189" ID-Pensoft-UUID="F005EAA1CA1959BFBC05BB150D885BD6" ID-ZooBank="B4388BBDBD5140F68341ADA7CF276E4B" ModsDocID="1313-2970-1132-189" checkinTime="1669899219071" checkinUser="pensoft" docAuthor="Lv, Heyu &amp; Di, Zhiyong" docDate="2022" docId="6468ABCA5F455BF9AF31BF2558DAC162" docLanguage="en" docName="ZooKeys 1132: 189-214" docOrigin="ZooKeys 1132" docPubDate="2022-11-30" docSource="http://dx.doi.org/10.3897/zookeys.1132.87364" docTitle="Scorpiops tibetanus Hirst 1911" docType="treatment" docVersion="3" id="F005EAA1CA1959BFBC05BB150D885BD6" lastPageNumber="189" masterDocId="F005EAA1CA1959BFBC05BB150D885BD6" masterDocTitle="Scorpiops lourencoi sp. nov., the revalidation of Scorpiops atomatus Qi, Zhu &amp; Lourenco, 2005, and the redescription of Scorpiops tibetanus Hirst, 1911 (Scorpiones, Scorpiopidae) from China" masterLastPageNumber="214" masterPageNumber="189" pageNumber="189" updateTime="1669900126619" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Scorpiops lourencoi sp. nov., the revalidation of Scorpiops atomatus Qi, Zhu &amp; Lourenco, 2005, and the redescription of Scorpiops tibetanus Hirst, 1911 (Scorpiones, Scorpiopidae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lv, Heyu</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Di, Zhiyong</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
<mods:nameIdentifier type="email">zydi@ustc.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1132</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>189</mods:start>
<mods:end>214</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1132.87364</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1132.87364</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1132-189</mods:identifier>
<mods:identifier type="ZooBank">B4388BBDBD5140F68341ADA7CF276E4B</mods:identifier>
<mods:identifier type="Pensoft-UUID">F005EAA1CA1959BFBC05BB150D885BD6</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162" httpUri="http://treatment.plazi.org/id/6468ABCA5F455BF9AF31BF2558DAC162" lastPageNumber="189" pageId="0" pageNumber="189">
<subSubSection pageId="0" pageNumber="189" type="nomenclature">
<paragraph pageId="0" pageNumber="189">
<taxonomicName LSID="6468ABCA-5F45-5BF9-AF31-BF2558DAC162" authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">Scorpiops tibetanus Hirst, 1911</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="description">
<paragraph pageId="0" pageNumber="189">
<figureCitation captionStart="Figures 33–36" captionStartId="F5" captionText="Figures 33 - 36. Scorpiops tibetanus from Qushui County 33, 34 male (Ar. - MHBU-ScXZQS 1907200101), dorsal and ventral views 35, 36 female (Ar. - MHBU-ScXZQS 1907200102), dorsal and ventral views. Scale bars: 12.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189">Figs 33-36</figureCitation>
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">, 37-46</figureCitation>
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">, 47-56</figureCitation>
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">, 57-64</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="reference_group">
<paragraph pageId="0" pageNumber="189">
<taxonomicName authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">Scorpiops tibetanus</taxonomicName>
Hirst, 1911: 472-473;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="189" pagination="153 - 201" refId="B7" refString="Kovarik, F, 2000. Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species. Acta Societatis Zoologicae Bohemicae 64: 153 - 201" title="Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species." volume="64" year="2000">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
2000
</bibRefCitation>
: 196, figs 47, 68, 69, tab. 1-3; Fet 2000: 495;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="0" pageNumber="189" pagination="1 - 140" publicationUrl="https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" refId="B9" refString="Kovarik, F, Lowe, G, Stockmann, M, Stahlavsky, F, 2020. Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida: Scorpiones). Euscorpius 325: 1 - 140, https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" title="Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida: Scorpiones)." url="https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" volume="325" year="2020">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
et al. 2020
</bibRefCitation>
: 126, figs 46, 143, 239-240, 799, tab. 9.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<taxonomicName authorityName="Qi, Zhu &amp; Lourenco" authorityYear="2005" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">Scorpiops pococki</taxonomicName>
Qi, Zhu &amp;
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
, 2005: 14, figs 47-61;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.301.4608" author="Di, ZY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="189" pagination="51 - 99" refId="B3" refString="Di, ZY, Xu, XB, Cao, ZJ, Wu, YL, Li, WX, 2013. Notes on the scorpions (Arachnida, Scorpiones) from Tibet with the redescription of Scorpiops jendeki Kovarik, 2000 (Scorpiones, Euscorpiidae) from Yunnan (China). ZooKeys 301: 51 - 99, DOI: https://doi.org/10.3897/zookeys.301.4608" title="Notes on the scorpions (Arachnida, Scorpiones) from Tibet with the redescription of Scorpiops jendeki Kovarik, 2000 (Scorpiones, Euscorpiidae) from Yunnan (China)." url="https://doi.org/10.3897/zookeys.301.4608" volume="301" year="2013">Di et al. 2013</bibRefCitation>
: 72, 75, figs 64-84, tab. 3;
<bibRefCitation author="Di, ZY" journalOrPublisher="Zoological Research" pageId="0" pageNumber="189" pagination="3 - 19" publicationUrl="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5042949/pdf/ZoolRes-35-1-3.pdf" refId="B4" refString="Di, ZY, Yang, ZZ, Yin, SJ, Cao, ZJ, Li, WX, 2014. History of study, updated checklist, distribution and key of scorpions (Arachnida: Scorpiones) from China. Zoological Research 35 (1): 3 - 19, https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5042949/pdf/ZoolRes-35-1-3.pdf" title="History of study, updated checklist, distribution and key of scorpions (Arachnida: Scorpiones) from China." url="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5042949/pdf/ZoolRes-35-1-3.pdf" volume="35" year="2014">Di et al. 2014</bibRefCitation>
: 12.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="type locality">
<paragraph pageId="0" pageNumber="189">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="189">China, Xizang, Tsangpo Valley, Chaksam Ferry.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="materials_examined">
<paragraph pageId="0" pageNumber="189">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="189">
<materialsCitation collectingDate="2019-07-20" collectorName="Zhiyong Di, Ar." country="China" county="Qushui County" location="Lhasa City" municipality="Lasa City" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xizang">
<specimenCount type="male">1 male</specimenCount>
and
<specimenCount type="female">1 female</specimenCount>
,
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
,
<collectingMunicipality>Lasa City</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:0A3520D33B6CA31139FEF9FC18FDDBF3" country="China" county="Qushui County" municipality="Lasa City" name="Lhasa City" stateProvince="Xizang">Lhasa City</location>
),
<collectingCounty>Qushui County</collectingCounty>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:D09972ED07626FC59986809BE7B33BF4" country="China" county="Qushui County" municipality="Lasa City" name="Chushur County" stateProvince="Xizang">Chushur County</location>
),
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:EB94D353D76201B3F69062B0F28A9563" country="China" county="Qushui County" municipality="Lasa City" name="Caina Town" stateProvince="Xizang">Caina Town</location>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:2115F4C296FB1C2D289BC2B4BCF725EB" country="China" county="Qushui County" municipality="Lasa City" name="Saena Town" stateProvince="Xizang">Saena Town</location>
),
<collectingDate value="2019-07-20">20/7/2019</collectingDate>
,
<collectorName>Zhiyong Di</collectorName>
leg, (
<collectorName>Ar.</collectorName>
-MHBU-ScXZQS1907200101,
<collectorName>Ar.</collectorName>
-MHBU-ScXZQS1907200102)
</materialsCitation>
;
<materialsCitation collectingDate="2021-08-12" collectorName="Zhiyong Di, Ar." country="China" county="Jiacha County" location="Lhoka City" municipality="Shannan City" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Xizang">
<specimenCount type="male">1 male</specimenCount>
and
<specimenCount type="female">2 females</specimenCount>
,
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
,
<collectingMunicipality>Shannan City</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:7380A67439B84640DFD056CAC52BFFF5" country="China" county="Jiacha County" municipality="Shannan City" name="Lhoka City" stateProvince="Xizang">Lhoka City</location>
),
<collectingCounty>Jiacha County</collectingCounty>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:01A6DCFB09622FA8DCAC423212410723" country="China" county="Jiacha County" municipality="Shannan City" name="Gyaca County" stateProvince="Xizang">Gyaca County</location>
),
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:D8E3443D5F951637D5F202A13D5BA39C" country="China" county="Jiacha County" municipality="Shannan City" name="Jiacha Town" stateProvince="Xizang">Jiacha Town</location>
(
<location LSID="urn:lsid:plazi:treatment:6468ABCA5F455BF9AF31BF2558DAC162:D18125BA2A89FB185FE20ECCD068F4F2" country="China" county="Jiacha County" municipality="Shannan City" name="Gyaca Town" stateProvince="Xizang">Gyaca Town</location>
),
<collectingDate value="2021-08-12">12/8/2021</collectingDate>
,
<collectorName>Zhiyong Di</collectorName>
leg, (
<collectorName>Ar.</collectorName>
-MHBU-ScXZJC
<collectingDate value="1206-08-21">21081206</collectingDate>
, 01-03)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="diagnosis">
<paragraph pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="189">
Adult body length 50-57 mm. Base color uniformly reddish black. Patella of pedipalp with 17 (5
<emphasis italics="true" pageId="0" pageNumber="189">eb</emphasis>
, 2
<emphasis italics="true" pageId="0" pageNumber="189">esb</emphasis>
, 2
<emphasis italics="true" pageId="0" pageNumber="189">em</emphasis>
, 4
<emphasis italics="true" pageId="0" pageNumber="189">est</emphasis>
, 4
<emphasis italics="true" pageId="0" pageNumber="189">et</emphasis>
) external and 6-8 (usually seven) ventral trichobothria. Chelal trichobothria
<emphasis italics="true" pageId="0" pageNumber="189">Eb3</emphasis>
is located in proximal half of manus between trichobothria
<emphasis italics="true" pageId="0" pageNumber="189">Dt</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="189">Db.</emphasis>
Chela with four ventral trichobothria. Chela with an average length/width ratio of 2.0 in both sexes, pedipalp movable finger with ca. four or five ID, 10-25 IAD, 45-62 MD, and eight or nine OD present, chela fingers on adult males and females scalloped, usually more strongly in males. Pectinal teeth count 4-7, fulcra absent. Pectinal with two marginal and one middle lamellae. Telson bulbous and granulate, annular ring present.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="description">
<paragraph pageId="0" pageNumber="189">Description</paragraph>
<paragraph pageId="0" pageNumber="189">(based on male specimen: Ar.-MHBU-ScXZQS1907200101).</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Coloration</emphasis>
(Figs
<figureCitation captionStart="Figures 33–36" captionStartId="F5" captionText="Figures 33 - 36. Scorpiops tibetanus from Qushui County 33, 34 male (Ar. - MHBU-ScXZQS 1907200101), dorsal and ventral views 35, 36 female (Ar. - MHBU-ScXZQS 1907200102), dorsal and ventral views. Scale bars: 12.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189">33</figureCitation>
,
<figureCitation captionStart="Figures 33–36" captionStartId="F5" captionText="Figures 33 - 36. Scorpiops tibetanus from Qushui County 33, 34 male (Ar. - MHBU-ScXZQS 1907200101), dorsal and ventral views 35, 36 female (Ar. - MHBU-ScXZQS 1907200102), dorsal and ventral views. Scale bars: 12.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189">34</figureCitation>
; after three years of preservation in alcohol): Carapace reddish black. Median and lateral ocular tubercles dark brown. Tergites and metasomal segments dark brown. Vesicle dark brown, with dark brown aculeus. Chelicerae unevenly dark brown and fingers uniformly dark reddish. Pedipalps dark reddish brown. Legs dark brown. Tarsal claws brown. Sternum reddish brown. Genital operculum and sternites brown. Pectinal teeth light brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="morphology">
<paragraph pageId="0" pageNumber="189">Morphology.</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Prosoma</emphasis>
(Figs
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">37</figureCitation>
,
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">38</figureCitation>
): Integument coarse, carapace with dense, fine granules; anterior median furrow broad and deep; lateral furrow broad; posterior median furrow broad and deep. Median eyes situated anteriorly compared to center of carapace; three pairs of lateral ocelli with posterior-most the smallest. Median ocular tubercle with granules and median furrow. Lateral ocular tubercle with some coarse granules around lateral eyes.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Mesosoma</emphasis>
: Tergites densely covered with fine granules, tergites II-VII with median carina barely visible at first and gradually becomes distinct; tergite VII with two pairs of lateral carinae with large granules present only on posterior half (Fig.
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">44</figureCitation>
). Pectinal teeth count 7/7, fulcra absent (Fig.
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">43</figureCitation>
). Genital operculum subtriangular with genital papillae protruding (Fig.
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">43</figureCitation>
). Sternum pentagonal (Fig.
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">43</figureCitation>
). Sternite segments III-VI entirely smooth and shiny with few setae; segment VII with four ventral carinae and few setae.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Metasoma</emphasis>
: Integument coarse, segments II-V longer than wide; segments I-V with respectively 10-8-8-8-8 granular carinae; segment V with pair of vestigial lateral carinae; all ventral carinae crenulated, gradually becoming more strongly crenulated. Vesicle with dense granules and few setae (Fig.
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">47</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Pedipalps</emphasis>
: Integument smooth with smooth granules and few setae. Femur with dorsointernal, dorsoexternal, external, ventroexternal, ventrointernal carinae granulated, and internal carinae crenulated (Fig.
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">48</figureCitation>
). Patella with large granules on dorsointernal, dorsoexternal, ventrointernal, ventroexternal, and smooth external carinae; two spinoid granules present on internal surface (Figs
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">49-51</figureCitation>
). Trichobothrial pattern C, neobothriotaxic (Vachon, 1974); patella with 17 external trichobothria (5
<emphasis italics="true" pageId="0" pageNumber="189">eb</emphasis>
, 2
<emphasis italics="true" pageId="0" pageNumber="189">esb</emphasis>
, 2
<emphasis italics="true" pageId="0" pageNumber="189">em</emphasis>
, 4
<emphasis italics="true" pageId="0" pageNumber="189">est</emphasis>
, 4
<emphasis italics="true" pageId="0" pageNumber="189">et</emphasis>
), 4 (right, dysplastic) and 7 (left) ventral trichobothria (Figs
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">50</figureCitation>
,
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">51</figureCitation>
). Chela with granules forming the indistinct reticulated pattern, ventral with four trichobothria, all carinae are granular and coalesced except the subdigital, dorsal internal, interomedian, and ventromedian carinae vestigial; movable and fixed fingers with scalloped margins, a pronounced lobe in movable finger and corresponding notch in fixed finger (Figs
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">57-60</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189" start="Figures 33–36" startId="F5">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 33-36.</emphasis>
<taxonomicName authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops tibetanus</emphasis>
</taxonomicName>
from Qushui County
<emphasis bold="true" pageId="0" pageNumber="189">33, 34</emphasis>
male (Ar.-MHBU-ScXZQS1907200101), dorsal and ventral views
<emphasis bold="true" pageId="0" pageNumber="189">35, 36</emphasis>
female (Ar.-MHBU-ScXZQS1907200102), dorsal and ventral views. Scale bars: 12.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189" start="Figures 37–46" startId="F6">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 37-46.</emphasis>
<taxonomicName authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops tibetanus</emphasis>
</taxonomicName>
from Qushui County
<emphasis bold="true" pageId="0" pageNumber="189">37, 38, 39, 43, 44</emphasis>
male (Ar.-MHBU-ScXZQS1907200101)
<emphasis bold="true" pageId="0" pageNumber="189">40, 41, 42, 45, 46</emphasis>
female (Ar.-MHBU-ScXZQS1907200102)
<emphasis bold="true" pageId="0" pageNumber="189">37, 40</emphasis>
carapace
<emphasis bold="true" pageId="0" pageNumber="189">38, 41</emphasis>
eyes and nearby area
<emphasis bold="true" pageId="0" pageNumber="189">39, 42</emphasis>
chelicera dorsal surface
<emphasis bold="true" pageId="0" pageNumber="189">43, 45</emphasis>
sternum, genital operculum and pectines
<emphasis bold="true" pageId="0" pageNumber="189">44, 46</emphasis>
tergites. Scale bars: 2.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189" start="Figures 47–56" startId="F7">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 47-56.</emphasis>
<taxonomicName authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops tibetanus</emphasis>
</taxonomicName>
from Qushui County
<emphasis bold="true" pageId="0" pageNumber="189">47-51</emphasis>
male (Ar.-MHBU-ScXZQS1907200101)
<emphasis bold="true" pageId="0" pageNumber="189">52-56</emphasis>
female (Ar.-MHBU-ScXZQS1907200102)
<emphasis bold="true" pageId="0" pageNumber="189">47, 52</emphasis>
telson, lateral surface
<emphasis bold="true" pageId="0" pageNumber="189">48, 53</emphasis>
femur dorsal surface
<emphasis bold="true" pageId="0" pageNumber="189">49-51, 54-56</emphasis>
patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189" start="Figures 57–64" startId="F8">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 57-64.</emphasis>
<taxonomicName authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops tibetanus</emphasis>
</taxonomicName>
from Qushui County and Jiacha County
<emphasis bold="true" pageId="0" pageNumber="189">57-59, 64</emphasis>
male (Ar.-MHBU-ScXZQS1907200101)
<emphasis bold="true" pageId="0" pageNumber="189">61-63</emphasis>
female (Ar.-MHBU-ScXZQS1907200102)
<emphasis bold="true" pageId="0" pageNumber="189">60</emphasis>
male (Ar.-MHBU-ScXZJC2108120601, from Jiacha County)
<emphasis bold="true" pageId="0" pageNumber="189">57-59, 61-63</emphasis>
dorsal, external, and ventral surfaces of chela
<emphasis bold="true" pageId="0" pageNumber="189">60</emphasis>
dentate margin of movable finger under UV light, showing rows of granules
<emphasis bold="true" pageId="0" pageNumber="189">64</emphasis>
right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Legs</emphasis>
: Integument coarse with few setae. Trochanter dorsal surface with few granules. Femur dorsal surface densely granular. Patella dorsal surface densely granular, with dorsoexternal and dorsal granular carinae. Tibiae without spurs (Fig.
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">64</figureCitation>
). Basitarsus with more setae, spurs, and two lateral pedal spurs (Fig.
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">64</figureCitation>
). Tarsus ventrally with single row of spinules (Fig.
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">64</figureCitation>
). Tarsal ungues curved and hook-like (Fig.
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">64</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Chelicerae</emphasis>
(Fig.
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">39</figureCitation>
): Dorsally with irregular pattern, ventrally with long hairs. Fixed finger of chelicera with three large triangular teeth on inner margin; ventral of movable finger with five teeth on inner margin, dorsal of movable finger with four teeth on inner margin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="variation">
<paragraph pageId="0" pageNumber="189">Variation.</paragraph>
<paragraph pageId="0" pageNumber="189">
Figures of adult females are provided (Figs
<figureCitation captionStart="Figures 33–36" captionStartId="F5" captionText="Figures 33 - 36. Scorpiops tibetanus from Qushui County 33, 34 male (Ar. - MHBU-ScXZQS 1907200101), dorsal and ventral views 35, 36 female (Ar. - MHBU-ScXZQS 1907200102), dorsal and ventral views. Scale bars: 12.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189">35</figureCitation>
,
<figureCitation captionStart="Figures 33–36" captionStartId="F5" captionText="Figures 33 - 36. Scorpiops tibetanus from Qushui County 33, 34 male (Ar. - MHBU-ScXZQS 1907200101), dorsal and ventral views 35, 36 female (Ar. - MHBU-ScXZQS 1907200102), dorsal and ventral views. Scale bars: 12.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures33-36" httpUri="https://binary.pensoft.net/fig/775974" pageId="0" pageNumber="189">36</figureCitation>
,
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">40-42</figureCitation>
,
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">45</figureCitation>
,
<figureCitation captionStart="Figures 37–46" captionStartId="F6" captionText="Figures 37 - 46. Scorpiops tibetanus from Qushui County 37, 38, 39, 43, 44 male (Ar. - MHBU-ScXZQS 1907200101) 40, 41, 42, 45, 46 female (Ar. - MHBU-ScXZQS 1907200102) 37, 40 carapace 38, 41 eyes and nearby area 39, 42 chelicera dorsal surface 43, 45 sternum, genital operculum and pectines 44, 46 tergites. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures37-46" httpUri="https://binary.pensoft.net/fig/775975" pageId="0" pageNumber="189">46</figureCitation>
,
<figureCitation captionStart="Figures 47–56" captionStartId="F7" captionText="Figures 47 - 56. Scorpiops tibetanus from Qushui County 47 - 51 male (Ar. - MHBU-ScXZQS 1907200101) 52 - 56 female (Ar. - MHBU-ScXZQS 1907200102) 47, 52 telson, lateral surface 48, 53 femur dorsal surface 49 - 51, 54 - 56 patella dorsal, external, and ventral surfaces. Green dots showing trichobothrial patterns of pedipalps (the white dots representing dysplasia caused trichobothrial absence). Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures47-56" httpUri="https://binary.pensoft.net/fig/775976" pageId="0" pageNumber="189">52-56</figureCitation>
,
<figureCitation captionStart="Figures 57–64" captionStartId="F8" captionText="Figures 57 - 64. Scorpiops tibetanus from Qushui County and Jiacha County 57 - 59, 64 male (Ar. - MHBU-ScXZQS 1907200101) 61 - 63 female (Ar. - MHBU-ScXZQS 1907200102) 60 male (Ar. - MHBU-ScXZJC 2108120601, from Jiacha County) 57 - 59, 61 - 63 dorsal, external, and ventral surfaces of chela 60 dentate margin of movable finger under UV light, showing rows of granules 64 right leg I retrolateral surface. Green dots showing trichobothrial patterns of pedipalps. Scale bars: 2.0 mm." figureDoi="10.3897/zookeys.1132.87364.figures57-64" httpUri="https://binary.pensoft.net/fig/775977" pageId="0" pageNumber="189">61-63</figureCitation>
). Number (left/right) of trichobothria on the ventral surface of the pedipalp patellae: females with 8/8 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1), 7/7 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1), and 6/6 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1), males with?/7 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1) and 7/7 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1). Number of pectinal teeth: females with 5/4 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 2) and 6/5 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 1), males with 7/7 (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 2). Chela with an average length/width ratio of 2.0 in males (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 2) and 2.0 in females (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
= 3), male pedipalp chela fingers stronger curved than females. The measurements are provided in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm) of S. atomatus, S. tibetanus, and S. lourencoi sp. nov." httpUri="http://table.plazi.org/id/89F0AD8D9541CCA87B3BEA4D7B05234F" pageId="0" pageNumber="189" tableUuid="89F0AD8D9541CCA87B3BEA4D7B05234F">1</tableCitation>
. Holotype (male, not examined;
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
, 2000): body length 60.4 mm, patella with eight ventral trichobothria, pectinal teeth count seven or eight. Holotype (male) of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
(=
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
): patella with 17 external and eight ventral trichobothria, pectinal teeth count 8/7; paratype (female) of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
(=
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
): pectinal teeth count 6/6, telson is smaller than that of male in
<bibRefCitation DOI="https://doi.org/10.18590/euscorpius.2005.vol2005.iss32.1" author="Qi, JX" journalOrPublisher="Euscorpius" pageId="0" pageNumber="189" pagination="1 - 40" refId="B13" refString="Qi, JX, Zhu, MS, Lourenco, WR, 2005. Eight new species of the genera Scorpiops Peters, Euscorpiops Vachon, and Chaerilus Simon (Scorpiones: Euscorpiidae, Chaerilidae) from Tibet and Yunnan, China. Euscorpius 32 (32): 1 - 40, DOI: https://doi.org/10.18590/euscorpius.2005.vol2005.iss32.1" title="Eight new species of the genera Scorpiops Peters, Euscorpiops Vachon, and Chaerilus Simon (Scorpiones: Euscorpiidae, Chaerilidae) from Tibet and Yunnan, China." url="https://doi.org/10.18590/euscorpius.2005.vol2005.iss32.1" volume="32" year="2005">Qi et al. (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="distribution">
<paragraph pageId="0" pageNumber="189">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="189">
China (Xizang) (Fig.
<figureCitation captionStart="Figure 97" captionStartId="F13" captionText="Figure 97. Distribution of S. atomatus (circle and diamond), S. tibetanus (pentagon and square), S. lourencoi sp. nov. (triangle), and the location of Qushui Bridge and Chaksam Ferry (star) in Xizang, China." figureDoi="10.3897/zookeys.1132.87364.figure97" httpUri="https://binary.pensoft.net/fig/775982" pageId="0" pageNumber="189">97</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="remarks">
<paragraph pageId="0" pageNumber="189">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="189">
<bibRefCitation DOI="https://doi.org/10.1080/00222931108693056" author="Hirst, S" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="189" pagination="462 - 473" refId="B6" refString="Hirst, S, 1911. Descriptions of new scorpions. Annals and Magazine of Natural History 8 (8): 462 - 473, DOI: https://doi.org/10.1080/00222931108693056" title="Descriptions of new scorpions." url="https://doi.org/10.1080/00222931108693056" volume="8" year="1911">Hirst (1911)</bibRefCitation>
erected
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
based on a male specimen from &quot;Tsangpo Valley, Chaksam Ferry&quot;, for which the species information is brief, and no figures provided.
</paragraph>
<paragraph pageId="0" pageNumber="189">
The type locality is therefore the most crucial information on the species. Followed the Internet and literature information for &quot;Chaksam Ferry&quot;:
<normalizedToken originalValue="“Chaksam”">&quot;Chaksam&quot;</normalizedToken>
means &quot;iron bridge&quot;, located in Dagar village and under the Quwori Mountain, Qushui County. It was created by the famous bridge designer, Tangdongjiebu, and is the first cable bridge across the Brahmaputra River, completed in 1420 or 1430. &quot;Chaksam Ferry&quot; subsequently became a famous place, also called &quot;Qushui Ferry&quot; or &quot;Daga Ferry&quot; in ancient times (Fig.
<figureCitation captionStart="Figure 97" captionStartId="F13" captionText="Figure 97. Distribution of S. atomatus (circle and diamond), S. tibetanus (pentagon and square), S. lourencoi sp. nov. (triangle), and the location of Qushui Bridge and Chaksam Ferry (star) in Xizang, China." figureDoi="10.3897/zookeys.1132.87364.figure97" httpUri="https://binary.pensoft.net/fig/775982" pageId="0" pageNumber="189">97</figureCitation>
). However, with the disappearance of the &quot;iron bridge&quot; and the construction of Qushui Bridge (Qushuidaqiao) near where the ancient &quot;iron bridge&quot; used to be in 1966, &quot;Chaksam Ferry&quot; has now been replaced by Qushui Bridge and is the probable type locality of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="189">
In recent years, we found
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. atomatus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="atomatus">
<emphasis italics="true" pageId="0" pageNumber="189">S. atomatus</emphasis>
</taxonomicName>
distributed near the &quot;iron bridge&quot;, and found
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
28 km away from this &quot;iron bridge&quot; (Fig.
<figureCitation captionStart="Figure 97" captionStartId="F13" captionText="Figure 97. Distribution of S. atomatus (circle and diamond), S. tibetanus (pentagon and square), S. lourencoi sp. nov. (triangle), and the location of Qushui Bridge and Chaksam Ferry (star) in Xizang, China." figureDoi="10.3897/zookeys.1132.87364.figure97" httpUri="https://binary.pensoft.net/fig/775982" pageId="0" pageNumber="189">97</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="189">
The list of taxa included in the
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. hardwickii" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="hardwickii">
<emphasis italics="true" pageId="0" pageNumber="189">S. hardwickii</emphasis>
</taxonomicName>
<normalizedToken originalValue="“complex”">&quot;complex&quot;</normalizedToken>
proposed by
<bibRefCitation DOI="https://doi.org/10.18590/euscorpius.2009.vol2009.iss88.1" author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="0" pageNumber="189" pagination="1 - 11" refId="B8" refString="Kovarik, F, Ahmed, Z, 2009. Three new species of Scorpiops Peters, 1861 (Scorpiones: Euscorpiidae: Scorpiopinae) from Pakistan. Euscorpius 88 (88): 1 - 11, DOI: https://doi.org/10.18590/euscorpius.2009.vol2009.iss88.1" title="Three new species of Scorpiops Peters, 1861 (Scorpiones: Euscorpiidae: Scorpiopinae) from Pakistan." url="https://doi.org/10.18590/euscorpius.2009.vol2009.iss88.1" volume="88" year="2009">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
and Ahmed (2009)
</bibRefCitation>
included
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Kovarik, F" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="189" pagination="153 - 201" refId="B7" refString="Kovarik, F, 2000. Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species. Acta Societatis Zoologicae Bohemicae 64: 153 - 201" title="Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species." volume="64" year="2000">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
(2000)
</bibRefCitation>
examined the holotype (male) of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
and recorded some important information: &quot;length 60.4 mm, ventral trichobothria on the patella number 8, and pectinal teeth number 7 or 8&quot;.
<bibRefCitation author="Kovarik, F" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="189" pagination="153 - 201" refId="B7" refString="Kovarik, F, 2000. Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species. Acta Societatis Zoologicae Bohemicae 64: 153 - 201" title="Revision of family Scorpiopidae (Scorpiones), with descriptions of six new species." volume="64" year="2000">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
(2000)
</bibRefCitation>
concluded that the characters of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
were &quot;length 50-65 mm, ventral trichobothria on the patella number 7-10 (usually 9, in one young of 37 specimens, 7 on one side), pectinal teeth number 5-11&quot;. In his revision, the new localities of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
included Lhasa, Shigatse, and Kambu batsi.
<bibRefCitation author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="0" pageNumber="189" pagination="1 - 140" publicationUrl="https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" refId="B9" refString="Kovarik, F, Lowe, G, Stockmann, M, Stahlavsky, F, 2020. Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida: Scorpiones). Euscorpius 325: 1 - 140, https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" title="Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida: Scorpiones)." url="https://mds.marshall.edu/cgi/viewcontent.cgi?article=1356&amp;context=euscorpius" volume="325" year="2020">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
et al. (2020)
</bibRefCitation>
proposed
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
and provided figures of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
(one male identified as &quot;
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
&quot; and donated by Di).
</paragraph>
<paragraph pageId="0" pageNumber="189">
During several surveys in Shigatse and the surrounding counties, we found
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. lourencoi" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="lourencoi">
<emphasis italics="true" pageId="0" pageNumber="189">S. lourencoi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. luridus." order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="luridus.">
<emphasis italics="true" pageId="0" pageNumber="189">S. luridus.</emphasis>
</taxonomicName>
The body length of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. lourencoi" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="lourencoi">
<emphasis italics="true" pageId="0" pageNumber="189">S. lourencoi</emphasis>
</taxonomicName>
sp. nov. is no more than 50 mm, while
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. luridus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="luridus">
<emphasis italics="true" pageId="0" pageNumber="189">S. luridus</emphasis>
</taxonomicName>
is a distinctive species.
<taxonomicName authorityName="Qi, Zhu &amp; Lourenco" authorityYear="2005" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops pococki</emphasis>
</taxonomicName>
(from Chaksam Ferry 28 km away) has the diagnosis as followed: fulcra absent, body length ~ 55 mm, the ventral trichobothria on the patella number seven or eight, and the pectinal teeth number 4-7.
<taxonomicName authorityName="Qi, Zhu &amp; Lourenco" authorityYear="2005" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpiops pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">Scorpiops pococki</emphasis>
</taxonomicName>
fits the characteristics of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
, and we agree that
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. pococki" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pococki">
<emphasis italics="true" pageId="0" pageNumber="189">S. pococki</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. tibetanus" order="Scorpiones" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis italics="true" pageId="0" pageNumber="189">S. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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