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<mods:title id="A5B112A6301053CB2F8256BB52C31A0F">Review of the mite family Pachylaelapidae (Acari: Mesostigmata)</mods:title>
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<subSubSection id="92BBA8FFFFB8FFB3FF236E449E27FB17" box="[151,563,1126,1152]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="DA1EFB74FFB8FFB3FF236E449E27FB17" blockId="10.[151,563,1126,1152]" box="[151,563,1126,1152]" pageId="10" pageNumber="11">
<emphasis id="E8D52766FFB8FFB3FF236E449E27FB17" bold="true" box="[151,563,1126,1152]" pageId="10" pageNumber="11">
Genus
<taxonomicName id="1DA180F7FFB8FFB3FF5A6E449E27FB17" ID-CoL="8W4VB" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[238,563,1126,1152]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FF5A6E449D93FB17" bold="true" box="[238,391,1126,1152]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
<bibRefCitation id="BE308685FFB8FFB3FE3A6E449E27FB17" author="Berlese" box="[398,563,1126,1152]" firstAuthor="Berlese" pageId="10" pageNumber="11" pagination="299 - 474" refId="ref38913" refString="Berlese, A. (1904) Illustrazione iconografica degli Acari mirmecofili. Redia, 1, 299 - 474 + Plates VII - XX." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<paragraph id="DA1EFB74FFB8FFB3FF236E8F9E96FB54" blockId="10.[151,986,1197,1250]" box="[151,642,1197,1219]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="FAB1DC5AFFB8FFB3FF236E8F9E96FB54" box="[151,642,1197,1219]" pageId="10" pageNumber="11">
<treatmentCitation id="5B00DD65FFB8FFB3FF236E8F9E6AFB54" author="Berlese, A." box="[151,638,1197,1219]" page="452" pageId="10" pageNumber="11" year="1904">
<taxonomicName id="1DA180F7FFB8FFB3FF236E8F9E6AFB54" authority="Berlese, 1904: 452" authorityName="Berlese" authorityPageNumber="452" authorityYear="1904" box="[151,638,1197,1219]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Onchodellus">
<emphasis id="E8D52766FFB8FFB3FF236E8F9D34FB54" box="[151,288,1197,1219]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
(
<emphasis id="E8D52766FFB8FFB3FE9A6E8F9DBAFB54" box="[302,430,1197,1219]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
)
<bibRefCitation id="BE308685FFB8FFB3FE086E8F9E6AFB54" author="Berlese" box="[444,638,1197,1219]" firstAuthor="Berlese" page="452" pageId="10" pageNumber="11" pagination="299 - 474" refId="ref38913" refString="Berlese, A. (1904) Illustrazione iconografica degli Acari mirmecofili. Redia, 1, 299 - 474 + Plates VII - XX." type="journal article" year="1904">Berlese, 1904: 452</bibRefCitation>
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.
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</paragraph>
<paragraph id="DA1EFB74FFB8FFB3FF236EEE9FCEFB76" blockId="10.[151,986,1197,1250]" box="[151,986,1228,1250]" pageId="10" pageNumber="11">
<typeStatus id="051A45D6FFB8FFB3FF236EEE9CDDFB76" box="[151,201,1228,1249]" pageId="10" pageNumber="11">Type</typeStatus>
species
<taxonomicName id="1DA180F7FFB8FFB3FEAB6EEE9F50FB75" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[287,836,1228,1250]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="reticulatus" subGenus="Onchodellus">
<emphasis id="E8D52766FFB8FFB3FEAB6EEE9DBCFB75" box="[287,424,1228,1250]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
(
<emphasis id="E8D52766FFB8FFB3FE026EEE9E22FB75" box="[438,566,1228,1250]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
)
<emphasis id="E8D52766FFB8FFB3FDF06EEE9EA4FB75" box="[580,688,1228,1250]" italics="true" pageId="10" pageNumber="11">reticulatus</emphasis>
<bibRefCitation id="BE308685FFB8FFB3FD026EEE9F50FB75" author="Berlese" box="[694,836,1228,1250]" firstAuthor="Berlese" pageId="10" pageNumber="11" pagination="299 - 474" refId="ref38913" refString="Berlese, A. (1904) Illustrazione iconografica degli Acari mirmecofili. Redia, 1, 299 - 474 + Plates VII - XX." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
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<paragraph id="DA1EFB74FFB8FFB3FF236F2C9948F9D1" blockId="10.[151,1436,1294,1823]" pageId="10" pageNumber="11">
<bibRefCitation id="BE308685FFB8FFB3FF236F2C9D5AFAB0" author="Berlese's" box="[151,334,1294,1319]" firstAuthor="Berlese's" pageId="10" pageNumber="11" pagination="299 - 474" refId="ref38913" refString="Berlese, A. (1904) Illustrazione iconografica degli Acari mirmecofili. Redia, 1, 299 - 474 + Plates VII - XX." type="journal article" year="1904">Berleses (1904)</bibRefCitation>
attempt to divide the genus
<taxonomicName id="1DA180F7FFB8FFB3FD3D6F2C9F34FAB0" authorityName="Berlese" authorityYear="1888" box="[649,800,1294,1319]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FD3D6F2C9F34FAB0" box="[649,800,1294,1319]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
</taxonomicName>
into subgenera
<taxonomicName id="1DA180F7FFB8FFB3FC646F2C9873FAB0" authorityName="Berlese" authorityYear="1888" box="[976,1127,1294,1319]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E8D52766FFB8FFB3FC646F2C9873FAB0" box="[976,1127,1294,1319]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3E69A1DFFB8FFB3FBD96F2D98BCFAB1" box="[1133,1192,1295,1318]" pageId="10" pageNumber="11" sensu="stricto">s. str.</taxonomicNameLabel>
and
<taxonomicName id="1DA180F7FFB8FFB3FB686F2C997DFAB0" authorityName="Berlese" authorityYear="1904" box="[1244,1385,1294,1319]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FB686F2C997DFAB0" box="[1244,1385,1294,1319]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
</taxonomicName>
was not generally accepted.
<bibRefCitation id="BE308685FFB8FFB3FE1C6F109E67FADC" author="Koroleva" box="[424,627,1330,1355]" firstAuthor="Koroleva" pageId="10" pageNumber="11" pagination="411 - 483" refId="ref42472" refString="Koroleva, E. V. (1977 b) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds) Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 411 - 483." type="book chapter" year="1977" yearSuffix="b">Koroleva (1977b)</bibRefCitation>
and
<bibRefCitation id="BE308685FFB8FFB3FD1A6F119F2EFADC" author="Karg" box="[686,826,1330,1355]" firstAuthor="Karg" pageId="10" pageNumber="11" pagination="1 - 523" refId="ref42302" refString="Karg, W. (1993) Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach. Raubmilben. 2. Uberarbeitete Auflage. Die Tierwelt Deutschlands, 59, 1 - 523." type="journal article" year="1993">Karg (1993)</bibRefCitation>
synonymised
<taxonomicName id="1DA180F7FFB8FFB3FC556F10987AFADC" authorityName="Berlese" authorityYear="1904" box="[993,1134,1330,1355]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FC556F10987AFADC" box="[993,1134,1330,1355]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
</taxonomicName>
with
<taxonomicName id="1DA180F7FFB8FFB3FB006F10995FFADC" authorityName="Berlese" authorityYear="1888" box="[1204,1355,1330,1355]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FB006F10995FFADC" box="[1204,1355,1330,1355]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
</taxonomicName>
. Later, the separate taxonomic status of
<taxonomicName id="1DA180F7FFB8FFB3FDB66F749E9BFAF8" authorityName="Berlese" authorityYear="1904" box="[514,655,1366,1391]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FDB66F749E9BFAF8" box="[514,655,1366,1391]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
</taxonomicName>
was accepted by
<bibRefCitation id="BE308685FFB8FFB3FCE76F759827FAF8" author="Evans &amp; Till" box="[851,1075,1366,1391]" firstAuthor="Evans" pageId="10" pageNumber="11" pagination="145 - 270" refId="ref40437" refString="Evans, G. O. &amp; Till, W. M. (1979) Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari - Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London, 35, 145 - 270. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1979. tb 00059. x" type="journal article" year="1979">Evans &amp; Till (1979)</bibRefCitation>
and
<bibRefCitation id="BE308685FFB8FFB3FBDC6F759983FAF8" author="Moraza &amp; Johnston" box="[1128,1431,1366,1391]" firstAuthor="Moraza" pageId="10" pageNumber="11" pagination="213 - 218" refId="ref43800" refString="Moraza, M. L. &amp; Johnston, D. E. (1990) Pachyseiulus hispanicus n. gen., n. sp., from Navarra (Northern Spain) (Acari: Mesostigmata: Pachylaelapidae). International Journal of Acarology, 16, 213 - 218. http: // dx. doi. org / 10.1080 / 01647959008683870" type="journal article" year="1990">Moraza &amp; Johnston (1990)</bibRefCitation>
, because the original
<taxonomicName id="1DA180F7FFB8FFB3FEC86F589E07FA04" authorityName="Berlese" authorityYear="1888" box="[380,531,1402,1427]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FEC86F589E07FA04" box="[380,531,1402,1427]" italics="true" pageId="10" pageNumber="11">Pachylaelaps</emphasis>
</taxonomicName>
exhibits two main morphological patterns previously noted by
<bibRefCitation id="BE308685FFB8FFB3FB606F589988FA04" author="Koroleva" box="[1236,1436,1402,1427]" firstAuthor="Koroleva" pageId="10" pageNumber="11" pagination="411 - 483" refId="ref42472" refString="Koroleva, E. V. (1977 b) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds) Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 411 - 483." type="book chapter" year="1977" yearSuffix="b">Koroleva (1977b)</bibRefCitation>
and
<bibRefCitation id="BE308685FFB8FFB3FF726FBD9DBDFA20" author="Evans &amp; Till" box="[198,425,1438,1463]" firstAuthor="Evans" pageId="10" pageNumber="11" pagination="145 - 270" refId="ref40437" refString="Evans, G. O. &amp; Till, W. M. (1979) Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari - Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London, 35, 145 - 270. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1979. tb 00059. x" type="journal article" year="1979">Evans &amp; Till (1979)</bibRefCitation>
in their identification keys: (1) species with a slender hypostome and elongated corniculi, a sperm access system associated with coxae IV, sexual dimorphism of the palp tibia (male with palp tibial outgrowths), two spur-like distal setae on female tarsus II, and 13 setae on genu I; and (2) species with a wider and shorter hypostome and normal horn-like corniculi, a sperm access system associated with coxae III, sexual dimorphism of palp tibia not developed, one spur-like distal setae on female tarsus II, and 12 setae on genu I.
</paragraph>
<paragraph id="DA1EFB74FFB8FFB3FF7D6C709CF2F888" blockId="10.[151,1436,1294,1823]" pageId="10" pageNumber="11">
The genus
<taxonomicName id="1DA180F7FFB8FFB3FEFC6C709DC1F9FC" authorityName="Berlese" authorityYear="1904" box="[328,469,1618,1643]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FFB8FFB3FEFC6C709DC1F9FC" box="[328,469,1618,1643]" italics="true" pageId="10" pageNumber="11">Onchodellus</emphasis>
</taxonomicName>
is now based especially on the following character states: (1) spermathecal tubes of sperm access system strongly elongated, fine, and associated with coxae III, (2) clunal setae J5 needle-like and similar in size to the other dorsal shield setae, (3) gland opening gdZ1 mostly not modified and circular (if slit-like, then antiaxial to alignment Z1Z2), (4) female tarsus II with one spur-like distal seta, and genu I with 12 pairs of setae. It is by far the largest genus of
<taxonomicName id="1DA180F7FFB8FFB3FD826CC09EFBF96C" authorityName="Berlese" authorityYear="1913" box="[566,751,1762,1787]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
, with up to 80 species distributed mainly in Europe, Asia and Africa.
</paragraph>
</subSubSection>
</treatment>
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