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<mods:title id="D5B15DBDA00CBE155D646C984C849F2B">Review of the mite family Pachylaelapidae (Acari: Mesostigmata)</mods:title>
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<subSubSection id="92BBA8FFFF80FF8BFF236F179F24FAD8" box="[151,816,1333,1359]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="DA1EFB74FF80FF8BFF236F179F24FAD8" blockId="50.[151,816,1333,1359]" box="[151,816,1333,1359]" pageId="50" pageNumber="51">
<heading id="81564C18FF80FF8BFF236F179F24FAD8" box="[151,816,1333,1359]" pageId="50" pageNumber="51">
<emphasis id="E8D52766FF80FF8BFF236F179CF3FAD8" bold="true" box="[151,231,1333,1359]" pageId="50" pageNumber="51">Genus</emphasis>
<taxonomicName id="1DA180F7FF80FF8BFF5A6F179F24FAD8" authority="Oudemans, 1901" authorityName="Oudemans" authorityYear="1901" box="[238,816,1333,1359]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFF5A6F179E72FAD8" bold="true" box="[238,614,1333,1359]" pageId="50" pageNumber="51">
<emphasis id="E8D52766FF80FF8BFF5A6F179D99FAD8" bold="true" box="[238,397,1333,1359]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
<bibRefCitation id="BE308685FF80FF8BFE216F179E72FAD8" author="Oudemans" box="[405,614,1333,1359]" firstAuthor="Oudemans" pageId="50" pageNumber="51" pagination="50 - 87" refId="ref44215" refString="Oudemans, A. C. (1901) Notes on Acari. Third ser. Tijdschrift der Nederlandsche Dierkundige Vereeniging (Second Series), 7, 50 - 87 + Plates I - III." type="journal article" year="1901">Oudemans, 1901</bibRefCitation>
</emphasis>
(Neoparasitidae)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92BBA8FFFF80FF8BFF236F5E9F33FA79" pageId="50" pageNumber="51" type="reference_group">
<paragraph id="DA1EFB74FF80FF8BFF236F5E9F21FA27" blockId="50.[151,1436,1404,1518]" pageId="50" pageNumber="51">
<taxonomicName id="1DA180F7FF80FF8BFF236F5E9E13FA05" authority="Oudemans, 1901: 53" authorityName="Oudemans" authorityPageNumber="53" authorityYear="1901" box="[151,519,1404,1426]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFF236F5E9D0AFA06" box="[151,286,1404,1425]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
<bibRefCitation id="BE308685FF80FF8BFE9D6F5E9E13FA05" author="Oudemans" box="[297,519,1404,1426]" firstAuthor="Oudemans" page="53" pageId="50" pageNumber="51" pagination="50 - 87" refId="ref44215" refString="Oudemans, A. C. (1901) Notes on Acari. Third ser. Tijdschrift der Nederlandsche Dierkundige Vereeniging (Second Series), 7, 50 - 87 + Plates I - III." type="journal article" year="1901">Oudemans, 1901: 53</bibRefCitation>
</taxonomicName>
.
<typeStatus id="051A45D6FF80FF8BFDA16F5E9E53FA06" box="[533,583,1404,1425]" pageId="50" pageNumber="51">Type</typeStatus>
species
<taxonomicName id="1DA180F7FF80FF8BFD126F5E9870FA05" authority="Oudemans, 1901" authorityName="Oudemans" authorityYear="1901" box="[678,1124,1404,1426]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="E8D52766FF80FF8BFD126F5E9FB2FA05" box="[678,934,1404,1426]" italics="true" pageId="50" pageNumber="51">Neoparasitus oudemansi</emphasis>
<bibRefCitation id="BE308685FF80FF8BFC056F5E9870FA05" author="Oudemans" box="[945,1124,1404,1426]" firstAuthor="Oudemans" pageId="50" pageNumber="51" pagination="50 - 87" refId="ref44215" refString="Oudemans, A. C. (1901) Notes on Acari. Third ser. Tijdschrift der Nederlandsche Dierkundige Vereeniging (Second Series), 7, 50 - 87 + Plates I - III." type="journal article" year="1901">Oudemans, 1901</bibRefCitation>
</taxonomicName>
, by monotypy (=
<taxonomicName id="1DA180F7FF80FF8BFA9E6F5E9DAFFA27" authority="Vitzthum, 1926" authorityName="Vitzthum" authorityYear="1926" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="quartus">
<emphasis id="E8D52766FF80FF8BFA9E6F5E9D00FA27" italics="true" pageId="50" pageNumber="51">Pachyseius quartus</emphasis>
<bibRefCitation id="BE308685FF80FF8BFEAF6FB89DAFFA27" author="Vitzthum" box="[283,443,1434,1456]" firstAuthor="Vitzthum" pageId="50" pageNumber="51" pagination="1 - 198" refId="ref45675" refString="Vitzthum, H. (1926) Malayische Acari. Treubia, 8, 1 - 198." type="journal article" year="1926">Vitzthum, 1926</bibRefCitation>
</taxonomicName>
; synonymy by
<bibRefCitation id="BE308685FF80FF8BFDE16FB89F3DFA27" author="Vitzthum" box="[597,809,1434,1456]" firstAuthor="Vitzthum" page="756" pageId="50" pageNumber="51" refId="ref45732" refString="Vitzthum, H. (1941 - 1943) Acarina. In: Bronns, H. G. (Ed.) Klassen und Ordnungen des Tierreichs. 5. Band 5: Arthropoda. IV. Abteilung: Arachnoidea, 5 Buch, Acarina. Becker &amp; Erler, Leipzig, pp. 1011 pp." type="book" year="1941">Vitzthum, 1941: 756</bibRefCitation>
).
</paragraph>
<paragraph id="DA1EFB74FF80FF8BFF236F9B9F33FA79" blockId="50.[151,1436,1404,1518]" pageId="50" pageNumber="51">
<taxonomicName id="1DA180F7FF80FF8BFF236F9B9E69FA58" authority="Gu, Wang &amp; Duan, 1991: 333" authorityName="Gu, Wang &amp; Duan" authorityPageNumber="333" authorityYear="1991" box="[151,637,1465,1487]" class="Arachnida" family="Pachylaelapidae" genus="Bulbogamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFF236F9B9D38FA58" box="[151,300,1465,1487]" italics="true" pageId="50" pageNumber="51">Bulbogamasus</emphasis>
<bibRefCitation id="BE308685FF80FF8BFE826F9B9E69FA58" author="Gu, Wang &amp; Duan" box="[310,637,1465,1487]" firstAuthor="Gu" page="333" pageId="50" pageNumber="51" pagination="333 - 338" refId="ref41072" refString="Gu, Y. - M., Wang, C. - S. &amp; Duan, Q. - X. (1991) Description of a new genus and a new species and proposal of a new family for the gamasides (Acari: Gamasina). Acta Zootaxonomica Sinica, 16, 333 - 338." type="journal article" year="1991">Gu, Wang &amp; Duan, 1991: 333</bibRefCitation>
</taxonomicName>
.
<typeStatus id="051A45D6FF80FF8BFD3F6F989EA9FA58" box="[651,701,1466,1487]" pageId="50" pageNumber="51">Type</typeStatus>
species
<taxonomicName id="1DA180F7FF80FF8BFCA86F9B990FFA58" authority="Gu, Wang &amp; Duan, 1991" authorityName="Gu, Wang &amp; Duan" authorityYear="1991" box="[796,1307,1465,1487]" class="Arachnida" family="Pachylaelapidae" genus="Bulbogamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="E8D52766FF80FF8BFCA86F9B9815FA58" box="[796,1025,1465,1487]" italics="true" pageId="50" pageNumber="51">Bulbogamasus sinicus</emphasis>
<bibRefCitation id="BE308685FF80FF8BFBB86F9B990FFA58" author="Gu, Wang &amp; Duan" box="[1036,1307,1465,1487]" firstAuthor="Gu" pageId="50" pageNumber="51" pagination="333 - 338" refId="ref41072" refString="Gu, Y. - M., Wang, C. - S. &amp; Duan, Q. - X. (1991) Description of a new genus and a new species and proposal of a new family for the gamasides (Acari: Gamasina). Acta Zootaxonomica Sinica, 16, 333 - 338." type="journal article" year="1991">Gu, Wang &amp; Duan, 1991</bibRefCitation>
</taxonomicName>
, by original designation. Synonymy by
<bibRefCitation id="BE308685FF80FF8BFE686FFA9F30FA79" author="Moraza &amp; Johnston" box="[476,804,1496,1518]" firstAuthor="Moraza" page="257" pageId="50" pageNumber="51" pagination="257 - 260" refId="ref43915" refString="Moraza, M. L. &amp; Johnston, D. E. (2002) Observations on Neoparasitus (= Bulbogamasus) associated with scarabaeid beetles in Indonesia (Acari: Mesostigmata). International Journal of Acarology, 28, 257 - 260. http: // dx. doi. org / 10.1080 / 01647950208684300" type="journal article" year="2002">Moraza &amp; Johnston (2002: 257)</bibRefCitation>
.
</paragraph>
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<subSubSection id="92BBA8FFFF80FF8AFF236C389F88FDF7" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="discussion">
<paragraph id="DA1EFB74FF80FF8BFF236C3898F0F955" blockId="50.[151,1436,1562,2019]" pageId="50" pageNumber="51">
<emphasis id="E8D52766FF80FF8BFF236C389CF5F9A4" bold="true" box="[151,225,1562,1587]" pageId="50" pageNumber="51">Notes.</emphasis>
<taxonomicName id="1DA180F7FF80FF8BFF5C6C399D69F9A5" authorityName="Oudemans" authorityYear="1901" box="[232,381,1563,1586]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFF5C6C399D69F9A5" box="[232,381,1563,1586]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
, as
<typeStatus id="051A45D6FF80FF8BFE1F6C399DCFF9A4" box="[427,475,1563,1587]" pageId="50" pageNumber="51">type</typeStatus>
genus of the family
<taxonomicName id="1DA180F7FF80FF8BFD706C389821F9A4" authority="Oudemans, 1939" authorityName="Oudemans" authorityYear="1939" box="[708,1077,1562,1587]" class="Arachnida" family="Neoparasitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">
Neoparasitidae
<bibRefCitation id="BE308685FF80FF8BFCC06C389821F9A4" author="Oudemans" box="[884,1077,1562,1587]" firstAuthor="Oudemans" pageId="50" pageNumber="51" pagination="195 - 201" refId="ref44466" refString="Oudemans, A. C. (1939) Neue Funde auf dem Gebiete der Systematik und Nomenklatur der Acari, IV. Zoologischer Anzeiger, 126, 195 - 201." type="journal article" year="1939">Oudemans, 1939</bibRefCitation>
</taxonomicName>
, has been classified in the same family by several later authors including
<bibRefCitation id="BE308685FF80FF8BFDC06C1C9F24F9C0" author="Vitzthum" box="[628,816,1598,1623]" firstAuthor="Vitzthum" pageId="50" pageNumber="51" refId="ref45732" refString="Vitzthum, H. (1941 - 1943) Acarina. In: Bronns, H. G. (Ed.) Klassen und Ordnungen des Tierreichs. 5. Band 5: Arthropoda. IV. Abteilung: Arachnoidea, 5 Buch, Acarina. Becker &amp; Erler, Leipzig, pp. 1011 pp." type="book" year="1941">Vitzthum (1941)</bibRefCitation>
and
<bibRefCitation id="BE308685FF80FF8BFCD86C1C9885F9C0" author="Baker &amp; Wharton" box="[876,1169,1598,1623]" firstAuthor="Baker" pageId="50" pageNumber="51" refId="ref38536" refString="Baker, E. W. &amp; Wharton, G. W. (1952) An Introduction to Acarology. Macmillan, New York, 465 pp." type="book" year="1952">Baker &amp; Wharton (1952)</bibRefCitation>
. The family was based especially on the presence of a 3-tined palp tarsal apotele and a holodorsal shield. It originally included various genera, most of which have since been transferred to other families, especially
<taxonomicName id="1DA180F7FF80FF8BFBDD6CA49913F908" authorityName="E.O.Evans" authorityYear="1961" box="[1129,1287,1670,1695]" class="Arachnida" family="Ameroseiidae" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Ameroseiidae</taxonomicName>
,
<taxonomicName id="1DA180F7FF80FF8BFAAE6CA4998CF908" baseAuthorityName="Tenorio &amp; Radovsky" baseAuthorityYear="1979" box="[1306,1432,1670,1695]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
,
<taxonomicName id="1DA180F7FF80FF8BFF236C889D29F954" baseAuthorityName="Krantz" baseAuthorityYear="1965" box="[151,317,1706,1731]" class="Arachnida" family="Macrochelidae" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Macrochelidae</taxonomicName>
and
<taxonomicName id="1DA180F7FF80FF8BFEC76C889E02F954" authorityName="Ryke" authorityYear="1962" box="[371,534,1706,1731]" class="Arachnida" family="Ologamasidae" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Ologamasidae</taxonomicName>
, and the family now includes only the
<typeStatus id="051A45D6FF80FF8BFC7C6C899FECF954" box="[968,1016,1707,1731]" pageId="50" pageNumber="51">type</typeStatus>
genus
<taxonomicName id="1DA180F7FF80FF8BFBF36C8998C8F955" authorityName="Oudemans" authorityYear="1901" box="[1095,1244,1707,1730]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFBF36C8998C8F955" box="[1095,1244,1707,1730]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DA1EFB74FF80FF8BFF7D6CED989DF828" blockId="50.[151,1436,1562,2019]" pageId="50" pageNumber="51">
Present knowledge of the
<typeStatus id="051A45D6FF80FF8BFE446CED9E34F970" box="[496,544,1743,1767]" pageId="50" pageNumber="51">type</typeStatus>
genus
<taxonomicName id="1DA180F7FF80FF8BFDC66CED9F1CF971" authorityName="Oudemans" authorityYear="1901" box="[626,776,1743,1766]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFDC66CED9F1CF971" box="[626,776,1743,1766]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
was recently summarised in detail by
<bibRefCitation id="BE308685FF80FF8BFB096CED9CF1F89C" author="Moraza &amp; Johnston" firstAuthor="Moraza" pageId="50" pageNumber="51" pagination="257 - 260" refId="ref43915" refString="Moraza, M. L. &amp; Johnston, D. E. (2002) Observations on Neoparasitus (= Bulbogamasus) associated with scarabaeid beetles in Indonesia (Acari: Mesostigmata). International Journal of Acarology, 28, 257 - 260. http: // dx. doi. org / 10.1080 / 01647950208684300" type="journal article" year="2002">Moraza &amp; Johnston (2002)</bibRefCitation>
. They considered
<taxonomicName id="1DA180F7FF80FF8BFE7F6CD19E74F89D" authorityName="Oudemans" authorityYear="1901" box="[459,608,1779,1802]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFE7F6CD19E74F89D" box="[459,608,1779,1802]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
and
<taxonomicName id="1DA180F7FF80FF8BFD1C6CD09F59F89C" authorityName="Gu, Wang &amp; Duan" authorityYear="1991" box="[680,845,1778,1803]" class="Arachnida" family="Pachylaelapidae" genus="Bulbogamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFD1C6CD09F59F89C" box="[680,845,1778,1803]" italics="true" pageId="50" pageNumber="51">Bulbogamasus</emphasis>
</taxonomicName>
as synonyms, and discussed problems with the classification of the genus. Moraza &amp; Johnston left the systematic position of
<taxonomicName id="1DA180F7FF80FF8BFB876D3598DCF8B9" authorityName="Oudemans" authorityYear="1901" box="[1075,1224,1815,1838]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFB876D3598DCF8B9" box="[1075,1224,1815,1838]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
open, stating that &quot;although the relationships between
<taxonomicName id="1DA180F7FF80FF8BFD8E6D199EDBF8C5" authorityName="Oudemans" authorityYear="1901" box="[570,719,1851,1874]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFD8E6D199EDBF8C5" box="[570,719,1851,1874]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
and other genera of the family
<taxonomicName id="1DA180F7FF80FF8BFBF66D1898ECF8C4" authorityName="Berlese" authorityYear="1913" box="[1090,1272,1850,1875]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
appear strong enough to consider them as members of the same family, this genus has characteristics shared with other eviphidioid families&quot;. It should be noted that the concept of
<taxonomicName id="1DA180F7FF80FF8BFCF86DA19FF5F80D" authorityName="Oudemans" authorityYear="1901" box="[844,993,1923,1946]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF80FF8BFCF86DA19FF5F80D" box="[844,993,1923,1946]" italics="true" pageId="50" pageNumber="51">Neoparasitus</emphasis>
</taxonomicName>
used by
<bibRefCitation id="BE308685FF80FF8BFBF86DA19995F80C" author="Moraza &amp; Johnston" box="[1100,1409,1922,1947]" firstAuthor="Moraza" pageId="50" pageNumber="51" pagination="257 - 260" refId="ref43915" refString="Moraza, M. L. &amp; Johnston, D. E. (2002) Observations on Neoparasitus (= Bulbogamasus) associated with scarabaeid beetles in Indonesia (Acari: Mesostigmata). International Journal of Acarology, 28, 257 - 260. http: // dx. doi. org / 10.1080 / 01647950208684300" type="journal article" year="2002">Moraza &amp; Johnston (2002)</bibRefCitation>
is based on
<taxonomicName id="1DA180F7FF80FF8BFEB46D859D98F828" authorityName="Nikolsky" authorityYear="1982" box="[256,396,1958,1983]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="E8D52766FF80FF8BFEB46D859D07F829" box="[256,275,1959,1982]" italics="true" pageId="50" pageNumber="51">N</emphasis>
.
<emphasis id="E8D52766FF80FF8BFE956D849D98F828" box="[289,396,1958,1983]" italics="true" pageId="50" pageNumber="51">orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="1DA180F7FF80FF8BFE776D859E26F829" authorityName="Gu, Wang &amp; Duan" authorityYear="1991" box="[451,562,1959,1982]" class="Arachnida" family="Pachylaelapidae" genus="Bulbogamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="E8D52766FF80FF8BFE776D859DC2F829" box="[451,470,1959,1982]" italics="true" pageId="50" pageNumber="51">N</emphasis>
.
<emphasis id="E8D52766FF80FF8BFE506D859E26F829" box="[484,562,1959,1982]" italics="true" pageId="50" pageNumber="51">sinicus</emphasis>
</taxonomicName>
, not the
<typeStatus id="051A45D6FF80FF8BFD206D859ED0F828" box="[660,708,1959,1983]" pageId="50" pageNumber="51">type</typeStatus>
species
<taxonomicName id="1DA180F7FF80FF8BFC966D859890F828" authority="Oudemans, 1901" authorityName="Oudemans" authorityYear="1901" box="[802,1156,1958,1983]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="E8D52766FF80FF8BFC966D859F21F829" box="[802,821,1959,1982]" italics="true" pageId="50" pageNumber="51">N</emphasis>
.
<emphasis id="E8D52766FF80FF8BFCF76D849FA8F828" box="[835,956,1958,1983]" italics="true" pageId="50" pageNumber="51">oudemansi</emphasis>
<bibRefCitation id="BE308685FF80FF8BFC706D849890F828" author="Oudemans" box="[964,1156,1958,1983]" firstAuthor="Oudemans" pageId="50" pageNumber="51" pagination="50 - 87" refId="ref44215" refString="Oudemans, A. C. (1901) Notes on Acari. Third ser. Tijdschrift der Nederlandsche Dierkundige Vereeniging (Second Series), 7, 50 - 87 + Plates I - III." type="journal article" year="1901">Oudemans, 1901</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="DA1EFB74FF80FF8AFF7D6DE99EEDFE3A" blockId="50.[151,1436,1562,2019]" lastBlockId="51.[151,1436,151,608]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
We here retain the monotypic family
<taxonomicName id="1DA180F7FF80FF8BFD316DE89F39F874" authorityName="Oudemans" authorityYear="1939" box="[645,813,1994,2019]" class="Arachnida" family="Neoparasitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="50" pageNumber="51" phylum="Arthropoda" rank="family">Neoparasitidae</taxonomicName>
in the superfamily
<taxonomicName id="1DA180F7FF80FF8BFBAD6DE898A5F874" authorityName="A.Berlese" authorityYear="1913" box="[1049,1201,1994,2019]" pageId="50" pageNumber="51" rank="superFamily" superFamily="Eviphidoidea">Eviphidoidea</taxonomicName>
, rather than placing
<taxonomicName id="1DA180F7FF81FF8AFF236ABA9D38FF38" authorityName="Oudemans" authorityYear="1901" box="[151,300,152,175]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFF236ABA9D38FF38" box="[151,300,152,175]" italics="true" pageId="51" pageNumber="52">Neoparasitus</emphasis>
</taxonomicName>
in either the Neoparasitinae or
<taxonomicName id="1DA180F7FF81FF8AFD2D6AB59F3FFF27" box="[665,811,151,176]" pageId="51" pageNumber="41" rank="subFamily" subFamily="Pachyseiinae">Pachyseiinae</taxonomicName>
within the
<taxonomicName id="1DA180F7FF81FF8AFC046AB5987DFF27" authorityName="Berlese" authorityYear="1913" box="[944,1129,151,176]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
, for the following reasons.
<taxonomicName id="1DA180F7FF81FF8AFF236A9F9D38FF43" authorityName="Oudemans" authorityYear="1901" box="[151,300,189,212]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFF236A9F9D38FF43" box="[151,300,189,212]" italics="true" pageId="51" pageNumber="52">Neoparasitus</emphasis>
</taxonomicName>
has in the female (1) a dorsal shield with more than 30 pairs of setae; (2) salivary styli well sclerotised; (3) corniculi weakly sclerotised, adjacent each to other and with hyaline fringe laterally, not horn-like; (4) cheliceral digits robust, each with a coarsely striate major tooth; (5) palp genu with seta al2 spatulate; (6) palp tarsus with elongate distal chemoreceptor; (8) exopodal platelets absent; parapodal platelets free and each well separated from peritrematal shield; (9) tarsi IIIV with minute claws and well developed empodium; (10) trochanters IIIV usually with apical dorsal spur; (11) genital setae sometimes off the epigynal shield; and in the male, (7) spermatodactyl strongly elongate and coiled.
</paragraph>
<paragraph id="DA1EFB74FF81FF8AFF7D6B9A9F88FDF7" blockId="51.[151,1436,151,608]" pageId="51" pageNumber="52">
In addition, species of
<taxonomicName id="1DA180F7FF81FF8AFE736B9A9E48FE58" authorityName="Oudemans" authorityYear="1901" box="[455,604,440,463]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFE736B9A9E48FE58" box="[455,604,440,463]" italics="true" pageId="51" pageNumber="52">Neoparasitus</emphasis>
</taxonomicName>
are mostly insecticolous species associated with coprid beetles, unlike the free-living edaphic species classified in the
<taxonomicName id="1DA180F7FF81FF8AFD376BFE9F0DFE62" box="[643,793,476,501]" pageId="51" pageNumber="41" rank="subFamily" subFamily="Pachyseiinae">Pachyseiinae</taxonomicName>
. The question of the correct classification and systematic placement of
<taxonomicName id="1DA180F7FF81FF8AFE9B68229DD0FD80" authorityName="Oudemans" authorityYear="1901" box="[303,452,512,535]" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFE9B68229DD0FD80" box="[303,452,512,535]" italics="true" pageId="51" pageNumber="52">Neoparasitus</emphasis>
</taxonomicName>
will not be resolved until more detailed morphological analysis of these mites is carried out, including study of their developmental stages and a complete re-description of the
<typeStatus id="051A45D6FF81FF8AFBC2680698B2FDAB" box="[1142,1190,548,572]" pageId="51" pageNumber="52">type</typeStatus>
species
<taxonomicName id="1DA180F7FF81FF8AFAB268079DC1FDF7" authority="Oudemans, 1901" authorityName="Oudemans" authorityYear="1901" class="Arachnida" family="Pachylaelapidae" genus="Neoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="E8D52766FF81FF8AFAB268079D04FDF7" italics="true" pageId="51" pageNumber="52">Neoparasitus oudemansi</emphasis>
<bibRefCitation id="BE308685FF81FF8AFEA368659DC1FDF7" author="Oudemans" box="[279,469,583,608]" firstAuthor="Oudemans" pageId="51" pageNumber="52" pagination="50 - 87" refId="ref44215" refString="Oudemans, A. C. (1901) Notes on Acari. Third ser. Tijdschrift der Nederlandsche Dierkundige Vereeniging (Second Series), 7, 50 - 87 + Plates I - III." type="journal article" year="1901">Oudemans, 1901</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="1DA180F7FF81FF8AFE4868659F84FDF7" authority="Vitzthum, 1926" authorityName="Vitzthum" authorityYear="1926" box="[508,912,583,608]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="quartus">
<emphasis id="E8D52766FF81FF8AFE4868659EC2FDF7" box="[508,726,583,608]" italics="true" pageId="51" pageNumber="52">Pachyseius quartus</emphasis>
<bibRefCitation id="BE308685FF81FF8AFD6968659F84FDF7" author="Vitzthum" box="[733,912,583,608]" firstAuthor="Vitzthum" pageId="51" pageNumber="52" pagination="1 - 198" refId="ref45675" refString="Vitzthum, H. (1926) Malayische Acari. Treubia, 8, 1 - 198." type="journal article" year="1926">Vitzthum, 1926</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>