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<document id="E2DE035B6B2FA35E9F36FD48B36B784A" ID-CLB-Dataset="21994" ID-DOI="10.5852/ejt.2017.352" ID-GBIF-Dataset="7fc7ebcb-2981-4a44-820c-1d8a47569a60" ID-ISSN="2118-9773" ID-Zenodo-Dep="3836261" ID-ZooBank="6328E049-24F3-42BE-A405-A79BF2F16EA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589997618402" checkinUser="carolina" docAuthor="Johanson, Kjell Arne" docDate="2017" docId="502D3B51FFEF1127FD87BDE1FB0DFEB4" docLanguage="en" docName="ejt-2017-352.pdf.imf" docOrigin="European Journal of Taxonomy 352" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Caledomina paniensis Johanson 2017, sp. nov." docType="treatment" docUuid="6713A2D6-B4FF-4329-BEA9-952BB910F8D8" docUuidSource="ZooBank" docVersion="6" lastPageNumber="4" masterDocId="AC144329FFEE1124FFE4BE77FFC9FFE2" masterDocTitle="Description of three new species of Caledomina (Insecta, Trichoptera, Ecnomidae) from New Caledonia" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1698840488652" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E73771330A2B49DEF256C192F79C97D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="26A2AD06E95F929A507985D07508DC7A">
<mods:title id="3DE8D8A0F255103BC3A46237802F9365">Description of three new species of Caledomina (Insecta, Trichoptera, Ecnomidae) from New Caledonia</mods:title>
</mods:titleInfo>
<mods:name id="6975F7A9D12573D87B8C090ED6A1CBA5" type="personal">
<mods:role id="4CD680544680C4188F38E7FD12A6F590">
<mods:roleTerm id="B8C05DF5DE93B5881BEEDFB9C1465310">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3C03DD16BB0D3CA0C86ABFC414984455">Johanson, Kjell Arne</mods:namePart>
</mods:name>
<mods:typeOfResource id="CAE9A6B6CFE169EEE6740612B191B894">text</mods:typeOfResource>
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<mods:titleInfo id="C6DD780965CCC4F362C3B265E77F7EAC">
<mods:title id="7E9F362935853E80FF30B2FFFCEE3649">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3542643786D3E63195BADB7B926FB4F8">
<mods:date id="D304ED275ABC2B7D610999D40D9C909A">2017</mods:date>
<mods:detail id="F0692135E243884961D86FE52627BE4F" type="pubDate">
<mods:number id="EEACF9CDCC4CB95AD30C184667AEAC21">2017-09-22</mods:number>
</mods:detail>
<mods:detail id="27EADBC0B0186E7D028C3F2F45F3F54C" type="issue">
<mods:number id="83D505B5F72A654C9E2E9B7139D3BF16">352</mods:number>
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<mods:start id="D9DAA27C31CD46802C4E5874A63F0AB5">1</mods:start>
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<mods:classification id="31EF8460A6F730B494D0DAC055E23B83">journal article</mods:classification>
<mods:identifier id="3B91ADD832A16CA4ED7D52B75F409556" type="CLB-Dataset">21994</mods:identifier>
<mods:identifier id="2120DBF9DAE504AABDA8FFC977268F84" type="DOI">10.5852/ejt.2017.352</mods:identifier>
<mods:identifier id="9C63E387B59A7F130BF1DC91843C095E" type="GBIF-Dataset">7fc7ebcb-2981-4a44-820c-1d8a47569a60</mods:identifier>
<mods:identifier id="9472E02707C4274515D6811835474B88" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="1EEBF6F1A8D077B9CFCA6224832B7AC0" type="Zenodo-Dep">3836261</mods:identifier>
<mods:identifier id="7449D911CE8258852E5C6DC7F5434869" type="ZooBank">6328E049-24F3-42BE-A405-A79BF2F16EA8</mods:identifier>
</mods:mods>
<treatment id="502D3B51FFEF1127FD87BDE1FB0DFEB4" ID-DOI="http://doi.org/10.5281/zenodo.3851717" ID-GBIF-Taxon="164243563" ID-Zenodo-Dep="3851717" LSID="urn:lsid:zoobank.org:act:6713A2D6-B4FF-4329-BEA9-952BB910F8D8" httpUri="http://treatment.plazi.org/id/502D3B51FFEF1127FD87BDE1FB0DFEB4" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="909ED9CCFFEF1125FD87BDE1FC18FC53" box="[611,977,918,945]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="D83B8A47FFEF1125FD87BDE1FC18FC53" blockId="1.[366,1222,918,1017]" box="[611,977,918,945]" pageId="1" pageNumber="2">
<heading id="83733D2BFFEF1125FD87BDE1FC18FC53" box="[611,977,918,945]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="1F84F1C4FFEF1125FD87BDE1FCA6FC52" authority="Johanson, 2017" authorityName="Johanson" authorityYear="2017" box="[611,879,918,945]" class="Insecta" family="Ecnomidae" genus="Caledomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="paniensis" status="sp. nov.">
<emphasis id="EAF05655FFEF1125FD87BDE1FCA6FC52" bold="true" box="[611,879,918,945]" italics="true" pageId="1" pageNumber="2">Caledomina paniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1C3EB2EFFEF1125FC92BDE0FC18FC53" box="[886,977,919,945]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="909ED9CCFFEF1125FE8ABDCDFCB1FC1A" pageId="1" pageNumber="2" type="description">
<paragraph id="D83B8A47FFEF1125FE8ABDCDFB0FFC37" blockId="1.[366,1222,918,1017]" box="[366,1222,954,981]" pageId="1" pageNumber="2">
<uri id="AC158645FFEF1125FE8ABDCDFB0FFC37" box="[366,1222,954,981]" pageId="1" pageNumber="2">
urn:lsid:zoobank.org:act:
<uuid id="AC22B092FFEF1125FD6ABDCDFB0FFC37" box="[654,1222,954,981]" pageId="1" pageNumber="2">6713A2D6-B4FF-4329-BEA9-952BB910F8D8</uuid>
</uri>
</paragraph>
<paragraph id="D83B8A47FFEF1125FD5FBDA9FCB1FC1A" blockId="1.[366,1222,918,1017]" box="[699,888,990,1017]" pageId="1" pageNumber="2">
<figureCitation id="40BF96C2FFEF1125FD5FBDA9FCEAFC1B" box="[699,803,990,1017]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Figs 1–5</figureCitation>
,
<figureCitation id="40BF96C2FFEF1125FCCBBDA9FC86FC1A" box="[815,847,990,1016]" captionStart="Fig" captionStartId="7.[190,233,1930,1956]" captionTargetId="figure@7.[187,1402,559,1687]" captionTargetPageId="7" captionText="Fig. 16. Map of New Caledonia with collecting sites of Caledomina indicated. Caledomina paniensis sp. nov. (black square), C. dorsospina sp. nov. (black triangle), C. kohensis sp. nov. (black filled circle), C. noumea Johanson, 2011 (red stars)." figureDoi="http://doi.org/10.5281/zenodo.3836271" httpUri="https://zenodo.org/record/3836271/files/figure.png" pageId="1" pageNumber="2">16</figureCitation>
,
<figureCitation id="40BF96C2FFEF1125FCBFBDA9FCB1FC1A" box="[859,888,990,1016]" captionStart="Fig" captionStartId="9.[189,232,1965,1991]" captionTargetBox="[189,1394,1168,1919]" captionTargetId="figure@9.[189,1399,1168,1924]" captionTargetPageId="9" captionText="Fig. 19. Phylogenetic relationship among species of Caledomina, with morphology of male genitalia in lateral view to the right." figureDoi="http://doi.org/10.5281/zenodo.3836275" httpUri="https://zenodo.org/record/3836275/files/figure.png" pageId="1" pageNumber="2">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="909ED9CCFFEF1125FF59BA57FCA3FBD8" box="[189,874,1056,1082]" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="D83B8A47FFEF1125FF59BA57FCA3FBD8" blockId="1.[189,874,1056,1082]" box="[189,874,1056,1082]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="F894AD69FFEF1125FF59BA57FCA3FBD8" box="[189,874,1056,1082]" pageId="1" pageNumber="2">
FI4
<taxonomicName id="1F84F1C4FFEF1125FF09BA57FEB5FBD8" authorityName="Johanson" authorityYear="2011" box="[237,380,1056,1082]" class="Insecta" family="Ecnomidae" genus="Caledomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="EAF05655FFEF1125FF09BA57FEB5FBD8" box="[237,380,1056,1082]" italics="true" pageId="1" pageNumber="2">Caledomina</emphasis>
</taxonomicName>
sp. 2 –
<treatmentCitation id="5925AC56FFEF1125FE3DBA57FCACFBD8" box="[473,869,1056,1082]" pageId="1" pageNumber="2">
<bibRefCitation id="BC15F7B6FFEF1125FE3DBA57FCACFBD8" author="Johanson K. A. &amp; Espeland M." box="[473,869,1056,1082]" pageId="1" pageNumber="2" pagination="36 - 48" refId="ref3441" refString="Johanson K. A. &amp; Espeland M. 2010. Phylogeny of the Ecnomidae (Insecta: Trichoptera). Cladistics 26: 36 - 48. https: // doi. org / 10.1111 / j. 1096 - 0031.2009.00276. x" type="journal article" year="2010">Espeland &amp; Johanson 2010: 2116</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="909ED9CCFFEF1125FF59BA16FB36FB2C" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="D83B8A47FFEF1125FF59BA16FEFFFB99" blockId="1.[189,310,1121,1147]" box="[189,310,1121,1147]" pageId="1" pageNumber="2">
<heading id="83733D2BFFEF1125FF59BA16FEFFFB99" bold="true" box="[189,310,1121,1147]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="EAF05655FFEF1125FF59BA16FEFFFB99" bold="true" box="[189,310,1121,1147]" pageId="1" pageNumber="2">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D83B8A47FFEF1125FF59BAE7FB36FB2C" blockId="1.[189,1398,1168,1231]" pageId="1" pageNumber="2">
<taxonomicName id="1F84F1C4FFEF1125FF59BAE7FE0FFB48" authorityName="Johanson" authorityYear="2017" box="[189,454,1168,1194]" class="Insecta" family="Ecnomidae" genus="Caledomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="paniensis">
<emphasis id="EAF05655FFEF1125FF59BAE7FE0FFB48" box="[189,454,1168,1194]" italics="true" pageId="1" pageNumber="2">Caledomina paniensis</emphasis>
</taxonomicName>
is distinguished from other species in the genus by the presence of a pair of sternal processes of sternite IX reaching about as long posteriorly as the inferior appendages.
</paragraph>
</subSubSection>
<subSubSection id="909ED9CCFFEF1125FF59BA81FC04FADD" box="[189,973,1270,1344]" pageId="1" pageNumber="2" type="etymology">
<paragraph id="D83B8A47FFEF1125FF59BA81FE8DFAF2" blockId="1.[189,324,1270,1296]" box="[189,324,1270,1296]" pageId="1" pageNumber="2">
<heading id="83733D2BFFEF1125FF59BA81FE8DFAF2" bold="true" box="[189,324,1270,1296]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="EAF05655FFEF1125FF59BA81FE8DFAF2" bold="true" box="[189,324,1270,1296]" pageId="1" pageNumber="2">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="D83B8A47FFEF1125FF59BB52FC04FADD" blockId="1.[189,973,1317,1344]" box="[189,973,1317,1344]" pageId="1" pageNumber="2">
The specific epithet is derived from the
<typeStatus id="073F34E5FFEF1125FD74BB51FD0AFAA2" box="[656,707,1318,1344]" pageId="1" pageNumber="2">type</typeStatus>
locality, Mount Panié.
</paragraph>
</subSubSection>
<subSubSection id="909ED9CCFFEF1125FF59BB10FE74F9F9" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="D83B8A47FFEF1125FF59BB10FEA5FA63" blockId="1.[189,364,1383,1409]" box="[189,364,1383,1409]" pageId="1" pageNumber="2">
<heading id="83733D2BFFEF1125FF59BB10FEA5FA63" bold="true" box="[189,364,1383,1409]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="EAF05655FFEF1125FF59BB10FEA5FA63" bold="true" box="[189,364,1383,1409]" pageId="1" pageNumber="2">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="68EC801AFFEF1125FF0EBBE1FE74F9F9" ID-GBIF-Occurrence="2621111303" collectingDate="2001-11-22" collectingDateMax="2001-11-28" collectingDateMin="2001-11-22" collectingMethod="Malaise trap" collectionCode="MNHN" collectorName="Johanson &amp; Pape &amp; Viklund" country="New Caledonia" elevation="350" latitude="-20.597733" location="Province Nord" longLatPrecision="1" longitude="164.82967" municipality="Mt Panie" pageId="1" pageNumber="2" specimenCode="NHRS-FI4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="D83B8A47FFEF1125FF0EBBE1FE95FA52" blockId="1.[189,1399,1430,1563]" box="[234,348,1430,1456]" pageId="1" pageNumber="2">
<emphasis id="EAF05655FFEF1125FF0EBBE1FE95FA52" bold="true" box="[234,348,1430,1456]" pageId="1" pageNumber="2">
<typeStatus id="073F34E5FFEF1125FF0EBBE1FE95FA52" box="[234,348,1430,1456]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D83B8A47FFEF1125FF59BBCDFE74F9F9" blockId="1.[189,1399,1430,1563]" pageId="1" pageNumber="2">
<collectingCountry id="A093CAD7FFEF1125FF59BBCDFE7BFA36" box="[189,434,1466,1492]" name="New Caledonia" pageId="1" pageNumber="2">NEW CALEDONIA</collectingCountry>
:
<specimenCount id="CE8241CEFFEF1125FE26BBCDFE13FA36" box="[450,474,1466,1492]" pageId="1" pageNumber="2" type="male">♂</specimenCount>
,
<location id="DD5BDC9CFFEF1125FE03BBCEFD5FFA31" LSID="urn:lsid:plazi:treatment:502D3B51FFEF1127FD87BDE1FB0DFEB4:DD5BDC9CFFEF1125FE03BBCEFD5FFA31" box="[487,662,1465,1491]" country="New Caledonia" latitude="-20.597733" longLatPrecision="1" longitude="164.82967" municipality="Mt Panie" name="Province Nord" pageId="1" pageNumber="2">Province Nord</location>
,
<collectingMunicipality id="385F103DFFEF1125FD46BBCDFCD9FA31" box="[674,784,1465,1492]" pageId="1" pageNumber="2">Mt Panié</collectingMunicipality>
, river below waterfall,
<geoCoordinate id="BDB0EC80FFEF1125FBC6BBCEFB73FA36" box="[1058,1210,1465,1492]" degrees="20" direction="south" minutes="35.864" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="-20.597733">20°35.864' S</geoCoordinate>
,
<geoCoordinate id="BDB0EC80FFEF1125FB2CBBCEFAB8FA36" box="[1224,1393,1465,1492]" degrees="164" direction="east" minutes="49.780" orientation="longitude" pageId="1" pageNumber="2" precision="1" value="164.82967">164°49.780' E</geoCoordinate>
,
<quantity id="1F7C27A2FFEF1125FF59BBAAFEC3FA1A" box="[189,266,1501,1528]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="2" unit="m" value="350.0">
<elevation id="53A96D74FFEF1125FF59BBAAFEC3FA1A" box="[189,266,1501,1528]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="2" unit="m" value="350.0">350 m</elevation>
</quantity>
, loc #16-2001,
<collectingMethod id="01C5F250FFEF1125FE26BBAAFD91FA1A" box="[450,600,1501,1528]" pageId="1" pageNumber="2">Malaise trap</collectingMethod>
,
<date id="AC3AAC87FFEF1125FD87BBAAFCF8FA15" box="[611,817,1501,1528]" pageId="1" pageNumber="2" value="2001-11-22" valueMax="2001-11-28" valueMin="2001-11-22">
<collectingDate id="BC7E556FFFEF1125FD87BBAAFCF8FA15" box="[611,817,1501,1528]" pageId="1" pageNumber="2" value="2001-11-22" valueMax="2001-11-28" valueMin="2001-11-22">22–28 Nov. 2001</collectingDate>
</date>
,
<collectorName id="7571EF91FFEF1125FCDABBAAFC67FA15" box="[830,942,1501,1527]" pageId="1" pageNumber="2">Johanson</collectorName>
,
<collectorName id="7571EF91FFEF1125FC5EBBA9FC3AFA1A" box="[954,1011,1502,1528]" pageId="1" pageNumber="2">Pape</collectorName>
and
<collectorName id="7571EF91FFEF1125FBC9BBAAFB42FA15" box="[1069,1163,1501,1527]" pageId="1" pageNumber="2">Viklund</collectorName>
leg. (
<collectionCode id="BE951282FFEF1125FB2ABBAAFAE5FA15" box="[1230,1324,1501,1527]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="2">MNHN</collectionCode>
, DNA voucher:
<specimenCode id="8822223CFFEF1125FECFB876FE78F9F9" box="[299,433,1537,1563]" collectionCode="NHRS-FI" pageId="1" pageNumber="2">NHRS-FI4</specimenCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="909ED9CCFFEF1126FF59B835FBF2FD42" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="D83B8A47FFEF1125FF59B835FE87F9BE" blockId="1.[189,334,1602,1628]" box="[189,334,1602,1628]" pageId="1" pageNumber="2">
<heading id="83733D2BFFEF1125FF59B835FE87F9BE" bold="true" box="[189,334,1602,1628]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="EAF05655FFEF1125FF59B835FE87F9BE" bold="true" box="[189,334,1602,1628]" pageId="1" pageNumber="2">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D83B8A47FFEF1125FF59B805FCB6F96E" blockId="1.[189,895,1650,1676]" box="[189,895,1650,1676]" pageId="1" pageNumber="2">
WINGS. Fore wing length
<quantity id="1F7C27A2FFEF1125FE03B805FD8DF96E" box="[487,580,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="2" unit="mm" value="2.2">2.2 mm</quantity>
, hind wing length
<quantity id="1F7C27A2FFEF1125FCF9B805FCB3F96E" box="[797,890,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="2" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
<paragraph id="D83B8A47FFEF1126FF59B8C3FBF2FD42" blockId="1.[189,1399,1715,2027]" lastBlockId="2.[189,1399,267,673]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
GENITALIA. Total length about
<quantity id="1F7C27A2FFEF1125FDFEB8C4FDBEF92C" box="[538,631,1715,1742]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="mm" value="0.5">0.5 mm</quantity>
. Segment IX divided into pair of strong, anteriorly pointed ridges in lateral view (
<figureCitation id="40BF96C2FFEF1125FE9DB8A0FE09F913" box="[377,448,1751,1777]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
); sternite IX originating from anterior ridge at mid-height; broadening posterad into pair of nearly parallel-sided plates with postero-dorsal sternal process (
<figureCitation id="40BF96C2FFEF1125FBBEB88CFB6AF8F7" box="[1114,1187,1787,1813]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
); smooth, except few ventral setae. Sternal processes short, slender, slightly curving dorsally, each with large apical and subapical spine nearly as long as sternal process (
<figureCitation id="40BF96C2FFEF1125FCF3B935FC4CF8BE" box="[791,901,1858,1884]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Figs 1, 3</figureCitation>
); orienting posterad along their length in ventral view (
<figureCitation id="40BF96C2FFEF1125FE86B911FE63F862" box="[354,426,1894,1920]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
); almost reaching point of bifurcation of superior appendages (
<figureCitation id="40BF96C2FFEF1125FB78B911FB2AF862" box="[1180,1251,1894,1920]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Tergite IX Y-shaped in lateral view (
<figureCitation id="40BF96C2FFEF1125FE03B9FEFDE3F846" box="[487,554,1929,1956]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
), anterodorsal corners strongly curving anterad before bending posterad at apex; with long setae on lateral faces; in dorsal view (
<figureCitation id="40BF96C2FFEF1125FCBFB9DAFC6AF825" box="[859,931,1965,1991]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
) with anterior-most part fused mesally; posterior part divided into tapering branches. Tergite X membranous, present above posterior apex of tergite IX in lateral view (
<figureCitation id="40BF96C2FFEC1126FE1EBF7CFD8BFEC7" box="[506,578,267,293]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
); in dorsal view divided into two lobes associated with branches of tergite IX; with minute setae along mesal margins (
<figureCitation id="40BF96C2FFEC1126FCC8BF46FCBEFEA9" box="[812,887,305,331]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Superior appendages large; each divided into posterior dorsal and ventral lobes (
<figureCitation id="40BF96C2FFEC1126FD61BF20FD01FE93" box="[645,712,343,369]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
), with setae basally of bifurcation (
<figureCitation id="40BF96C2FFEC1126FB81BF20FB06FE93" box="[1125,1231,343,369]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
). Dorsal lobes smooth, except with long row of hook-like spines oriented ventro-mesad (
<figureCitation id="40BF96C2FFEC1126FBF6BF0AFBB2FE75" box="[1042,1147,381,407]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
). Ventral lobes nearly straight along their length; ventral margin with minute spines; posterior apex narrowly rounded, covered with small setae; slightly exceeding dorsal lobe in length (
<figureCitation id="40BF96C2FFEC1126FC85BFBEFC6FFE01" box="[865,934,457,483]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Inferior appendages completely fused into small, mesal process, connected to central part of sternite IX by vaguely defined sclerotized ridges; dorsal margin slightly sigmoid; ventral margin convex except curving at apex; apex long, narrow in lateral view (
<figureCitation id="40BF96C2FFEC1126FEB9BC4DFE6CFDB7" box="[349,421,570,597]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
); in ventral view (
<figureCitation id="40BF96C2FFEC1126FD68BC4DFD1CFDB7" box="[652,725,570,597]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
) with broad basis; narrowing slightly at one quarter its length, apex smooth, bifurcated. Phallic apparatus without processes (
<figureCitation id="40BF96C2FFEC1126FBECBC17FBBFFD99" box="[1032,1142,608,635]" captionStart="Figs 1–5" captionStartId="2.[189,241,1965,1991]" captionTargetBox="[230,1373,725,1922]" captionTargetId="figure@2.[189,1398,715,1928]" captionTargetPageId="2" captionText="Figs 1–5. Caledomina paniensis sp. nov., holotype, adult ♂. 1. Genitalia, lateral view. 2. Genitalia dorsal view. 3. Genitalia, ventral view. 4. Phallus, lateral view. 5. Phallus ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3">Figs 4–5</figureCitation>
); tube-shaped; dorsal margin weakly narrowing at anterior one-third; posterior apex membranous.
</paragraph>
</subSubSection>
<caption id="8CFBDACFFFEC1126FF59B9DAFA91F809" ID-DOI="http://doi.org/10.5281/zenodo.3836263" ID-Zenodo-Dep="3836263" httpUri="https://zenodo.org/record/3836263/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,241,1965,1991]" targetBox="[230,1373,725,1922]" targetPageId="2">
<paragraph id="D83B8A47FFEC1126FF59B9DAFA91F809" blockId="2.[189,1399,1965,2027]" pageId="2" pageNumber="3">
<emphasis id="EAF05655FFEC1126FF59B9DAFEE2F825" bold="true" box="[189,299,1965,1991]" pageId="2" pageNumber="3">Figs 1–5.</emphasis>
<taxonomicName id="1F84F1C4FFEC1126FED4B9DAFDFCF825" authority="Johanson, 2017" authorityName="Johanson" authorityYear="2017" box="[304,565,1965,1991]" class="Insecta" family="Ecnomidae" genus="Caledomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="paniensis" status="sp. nov.">
<emphasis id="EAF05655FFEC1126FED4B9DAFDFCF825" box="[304,565,1965,1991]" italics="true" pageId="2" pageNumber="3">Caledomina paniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1C3EB2EFFEC1126FDDFB9D9FD5FF82A" box="[571,662,1966,1992]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, adult ♂.
<emphasis id="EAF05655FFEC1126FC9DB9DAFC41F825" bold="true" box="[889,904,1965,1991]" pageId="2" pageNumber="3">1</emphasis>
. Genitalia, lateral view.
<emphasis id="EAF05655FFEC1126FB45B9DAFB79F825" bold="true" box="[1185,1200,1965,1991]" pageId="2" pageNumber="3">2</emphasis>
. Genitalia dorsal view.
<emphasis id="EAF05655FFEC1126FEE7B9A6FEDBF809" bold="true" box="[259,274,2001,2027]" pageId="2" pageNumber="3">3</emphasis>
. Genitalia, ventral view.
<emphasis id="EAF05655FFEC1126FDDDB9A6FD81F809" bold="true" box="[569,584,2001,2027]" pageId="2" pageNumber="3">4</emphasis>
. Phallus, lateral view.
<emphasis id="EAF05655FFEC1126FCB6B9A6FCA8F809" bold="true" box="[850,865,2001,2027]" pageId="2" pageNumber="3">5</emphasis>
. Phallus ventral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="909ED9CCFFED1127FF59BF7CFB0DFEB4" box="[189,1220,267,342]" pageId="3" pageNumber="4" type="discussion">
<paragraph id="D83B8A47FFED1127FF59BF7CFEE6FEC7" blockId="3.[189,303,267,293]" box="[189,303,267,293]" pageId="3" pageNumber="4">
<heading id="83733D2BFFED1127FF59BF7CFEE6FEC7" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="EAF05655FFED1127FF59BF7CFEE6FEC7" bold="true" box="[189,303,267,293]" pageId="3" pageNumber="4">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D83B8A47FFED1127FF59BF4CFB0DFEB4" blockId="3.[189,1220,315,342]" box="[189,1220,315,342]" pageId="3" pageNumber="4">
This species was referred to as “FI4
<taxonomicName id="1F84F1C4FFED1127FD8DBF4CFD31FEB7" authorityName="Johanson" authorityYear="2011" box="[617,760,315,341]" class="Insecta" family="Ecnomidae" genus="Caledomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EAF05655FFED1127FD8DBF4CFD31FEB7" box="[617,760,315,341]" italics="true" pageId="3" pageNumber="4">Caledomina</emphasis>
</taxonomicName>
sp. 2” in Espeland &amp;
<bibRefCitation id="BC15F7B6FFED1127FC1EBF4CFB76FEB4" author="Johanson K. A. &amp; Espeland M." box="[1018,1215,315,342]" pageId="3" pageNumber="4" pagination="36 - 48" refId="ref3441" refString="Johanson K. A. &amp; Espeland M. 2010. Phylogeny of the Ecnomidae (Insecta: Trichoptera). Cladistics 26: 36 - 48. https: // doi. org / 10.1111 / j. 1096 - 0031.2009.00276. x" type="journal article" year="2010">Johanson (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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