155 lines
18 KiB
XML
155 lines
18 KiB
XML
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<mods:title id="EF1D6E72647176759ED044F3EEB94E30">Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)</mods:title>
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<mods:date id="E49F93245EA82A0F73BAFD909F5093D4">2011</mods:date>
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<treatment id="485787BFFFBAC32AFF0BF9B7FE71FDF6" ID-DOI="http://doi.org/10.5281/zenodo.5621728" ID-GBIF-Taxon="119417555" ID-Zenodo-Dep="5621728" LSID="urn:lsid:plazi:treatment:485787BFFFBAC32AFF0BF9B7FE71FDF6" httpUri="http://treatment.plazi.org/id/485787BFFFBAC32AFF0BF9B7FE71FDF6" lastPageId="8" lastPageNumber="44" pageId="7" pageNumber="44">
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<subSubSection id="88E46522FFBAC325FF0BF9B7FD31F93C" box="[151,694,1659,1686]" pageId="7" pageNumber="44" type="nomenclature">
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<paragraph id="C04136A9FFBAC325FF0BF9B7FD31F93C" blockId="7.[151,694,1659,1686]" box="[151,694,1659,1686]" pageId="7" pageNumber="44">
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<heading id="9B0981C5FFBAC325FF0BF9B7FD31F93C" bold="true" box="[151,694,1659,1686]" fontSize="11" level="1" pageId="7" pageNumber="44" reason="1">
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<emphasis id="F28AEABBFFBAC325FF0BF9B7FD31F93C" bold="true" box="[151,694,1659,1686]" pageId="7" pageNumber="44">
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<taxonomicName id="07FE4D2AFFBAC325FF0BF9B7FE7EF93C" authority="Tenkatei Lidth" authorityName="Tenkatei Lidth" box="[151,505,1659,1686]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
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<emphasis id="F28AEABBFFBAC325FF0BF9B7FE2CF93F" bold="true" box="[151,427,1660,1686]" italics="true" pageId="7" pageNumber="44">Hemidactylus Tenkatei</emphasis>
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Lidth
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</taxonomicName>
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de
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<bibRefCitation id="A46F4B58FFBAC325FDB8F9B0FD31F93C" author="Jeude" box="[548,694,1659,1685]" pageId="7" pageNumber="44" refString="Lidth de Jeude, T. W. V. (1895) Reptiles from Timor and the neighbouring islands. Notes from the Leyden Museum, 16, 119 - 127." type="journal article" year="1895">Jeude, 1895</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="88E46522FFBAC32AFF0BF908FE71FDF6" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="reference_group">
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<paragraph id="C04136A9FFBAC32AFF0BF908FE71FDF6" blockId="7.[151,1436,1730,2006]" lastBlockId="8.[151,1436,151,607]" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
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<taxonomicName id="07FE4D2AFFBAC325FF0BF908FE17F972" box="[151,400,1731,1755]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFBAC325FF0BF908FE17F972" box="[151,400,1731,1755]" italics="true" pageId="7" pageNumber="44">Hemidactylus tenkatei</emphasis>
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</taxonomicName>
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was described based on two unsexed adults and one juvenile specimen presented to the Leyden Museum,
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<collectingCountry id="B8E97639FFBAC325FEA7F92DFE70F957" box="[315,503,1766,1790]" name="Netherlands" pageId="7" pageNumber="44">The Netherlands</collectingCountry>
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(currently RMNH) by Dr. H. ten Kate, from “Rotti” (Lidth de
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<bibRefCitation id="A46F4B58FFBAC325FB49F92DFADCF957" author="Jeude" box="[1237,1371,1766,1790]" pageId="7" pageNumber="44" refString="Lidth de Jeude, T. W. V. (1895) Reptiles from Timor and the neighbouring islands. Notes from the Leyden Museum, 16, 119 - 127." type="journal article" year="1895">Jeude 1895</bibRefCitation>
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). The description, though detailed, is inadequate to identify the species affinities within
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<emphasis id="F28AEABBFFBAC325FBD8F8C7FB5CF88A" box="[1092,1243,1803,1827]" italics="true" pageId="7" pageNumber="44">
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<taxonomicName id="07FE4D2AFFBAC325FBD8F8C7FB3FF88A" box="[1092,1208,1803,1827]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="44" phylum="Chordata" rank="species" sensu="lato" species="brookii">H. brookii</taxonomicName>
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<taxonomicNameLabel id="E9B957C0FFBAC325FB5DF8C0FB5CF88A" box="[1217,1243,1803,1827]" pageId="7" pageNumber="44" sensu="lato">s.l</taxonomicNameLabel>
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</emphasis>
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. The
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<typeStatus id="1F45880BFFBAC325FABCF8C0FAD7F88A" box="[1312,1360,1803,1827]" pageId="7" pageNumber="44">type</typeStatus>
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specimens are still present in the collection of RMNH (RMNH 4353––three specimens; Ronald de Ruiter
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<emphasis id="F28AEABBFFBAC325FA95F8E4FACBF8EF" box="[1289,1356,1838,1862]" italics="true" pageId="7" pageNumber="44">in litt.</emphasis>
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2010).
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<bibRefCitation id="A46F4B58FFBAC325FF0BF899FE2FF8C3" author="Rosler" box="[151,424,1874,1898]" pageId="7" pageNumber="44" refString="Rosler, H. & Glaw, F. (2010) Morphologische Variation und Taxonomie von Hemidactylus brookii Gray, 1845 und Hemidactylus angulatus Hallowell, 1854 sowie phanotypisch ahnlicher Taxa. Spixiana, 33 (1), 139 - 160." type="journal article" year="2010">Rösler and Glaw (2010)</bibRefCitation>
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provide a description of two topotypes, which they used to elevate this nomen as a valid species. Photographs provided to me of the two adult
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<typeStatus id="1F45880BFFBAC325FC9CF8BCFCE5F826" box="[768,866,1911,1935]" pageId="7" pageNumber="44" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="07FE4D2AFFBAC325FC17F8B3FB8FF827" box="[907,1032,1910,1935]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFBAC325FC17F8B3FB8FF827" box="[907,1032,1910,1935]" italics="true" pageId="7" pageNumber="44">H. tenkatei</emphasis>
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</taxonomicName>
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reveal several important characters that diagnose this species from all but one other
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<taxonomicName id="07FE4D2AFFBAC325FD50F857FCC6F81A" box="[716,833,1947,1971]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFBAC325FD50F857FCC6F81A" box="[716,833,1947,1971]" italics="true" pageId="7" pageNumber="44">H. brookii</emphasis>
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</taxonomicName>
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group taxa redefined herein. Subcaudal scales completely transverse the tail width at approx. 50% of the original tail length and enlarged femoral scales of one of the female specimens indicate seven enlarged scales on each thigh separated medially by six smaller scales. These enlarged scales likely represent the enlarged pore-bearing scales of males, separated by a diastema of non-porebearing scales. These diagnosing characters are thus far only confirmed on
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<taxonomicName id="07FE4D2AFFB5C32AFC63FF2AFB42FF51" box="[1023,1221,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFC63FF2AFB42FF51" box="[1023,1221,224,248]" italics="true" pageId="8" pageNumber="45">H. subtriedroides</emphasis>
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</taxonomicName>
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. One
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<emphasis id="F28AEABBFFB5C32AFA92FF2AFAADFF51" box="[1294,1322,225,248]" italics="true" pageId="8" pageNumber="45">H.</emphasis>
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“
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<taxonomicName id="07FE4D2AFFB5C32AFADCFF2BFA17FF51" box="[1344,1424,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFB5C32AFADCFF2BFA17FF51" box="[1344,1424,224,248]" italics="true" pageId="8" pageNumber="45">brookii</emphasis>
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</taxonomicName>
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” specimen (BMNH 1926.10.30.46) from West Timor, nearby
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<collectingCountry id="B8E97639FFB5C32AFCA1FECFFCFCFEB5" box="[829,891,260,284]" name="Palau" pageId="8" pageNumber="45">Palau</collectingCountry>
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Roti, was examined in this study and is indistinguishable from
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<taxonomicName id="07FE4D2AFFB5C32AFEDBFEE2FD8CFEE9" box="[327,523,296,320]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFEDBFEE2FEDCFEE9" box="[327,347,297,320]" italics="true" pageId="8" pageNumber="45">H</emphasis>
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.
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<emphasis id="F28AEABBFFB5C32AFEF6FEE3FD8CFEE9" box="[362,523,296,320]" italics="true" pageId="8" pageNumber="45">subtriedroides</emphasis>
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</taxonomicName>
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. Additionally, I can find no characters from the
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<bibRefCitation id="A46F4B58FFB5C32AFBB7FEECFABDFE96" author="Rosler" box="[1067,1338,295,319]" pageId="8" pageNumber="45" refString="Rosler, H. & Glaw, F. (2010) Morphologische Variation und Taxonomie von Hemidactylus brookii Gray, 1845 und Hemidactylus angulatus Hallowell, 1854 sowie phanotypisch ahnlicher Taxa. Spixiana, 33 (1), 139 - 160." type="journal article" year="2010">Rösler and Glaw (2010)</bibRefCitation>
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description of
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<taxonomicName id="07FE4D2AFFB5C32AFF69FE86FEF0FECD" box="[245,375,332,356]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFB5C32AFF69FE86FEF0FECD" box="[245,375,332,356]" italics="true" pageId="8" pageNumber="45">H. tenkatei</emphasis>
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</taxonomicName>
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topotypes that distinguish this species from
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<taxonomicName id="07FE4D2AFFB5C32AFC05FE87FB66FECD" box="[921,1249,332,356]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFC05FE87FB66FECD" box="[921,1249,332,356]" italics="true" pageId="8" pageNumber="45">Hemidactylus subtriedroides</emphasis>
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</taxonomicName>
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, thus based on morphological evidence I here consider the two as one and the same species. The revalidation of
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<taxonomicName id="07FE4D2AFFB5C32AFB66FEBAFAF0FE21" box="[1274,1399,368,392]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFB5C32AFB66FEBAFAF0FE21" box="[1274,1399,368,392]" italics="true" pageId="8" pageNumber="45">H. tenkatei</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="A46F4B58FFB5C32AFF0BFE5FFE12FE05" author="Rosler" box="[151,405,404,428]" pageId="8" pageNumber="45" refString="Rosler, H. & Glaw, F. (2010) Morphologische Variation und Taxonomie von Hemidactylus brookii Gray, 1845 und Hemidactylus angulatus Hallowell, 1854 sowie phanotypisch ahnlicher Taxa. Spixiana, 33 (1), 139 - 160." type="journal article" year="2010">Rösler & Glaw (2010)</bibRefCitation>
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, disqualifies the possible retention of the more commonly used name
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<taxonomicName id="07FE4D2AFFB5C32AFB3AFE5EFAEDFE05" box="[1190,1386,404,428]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFB3AFE5EFAEDFE05" box="[1190,1386,404,428]" italics="true" pageId="8" pageNumber="45">H. subtriedroides</emphasis>
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</taxonomicName>
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as a
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<emphasis id="F28AEABBFFB5C32AFF0BFE72FEDCFE79" box="[151,347,441,464]" italics="true" pageId="8" pageNumber="45">nomen protectum</emphasis>
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(Article 23.9.1.2, 23.9.2). Chronologically
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<taxonomicName id="07FE4D2AFFB5C32AFCCCFE72FC4AFE79" box="[848,973,440,464]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFB5C32AFCCCFE72FC4AFE79" box="[848,973,440,464]" italics="true" pageId="8" pageNumber="45">H. tenkatei</emphasis>
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</taxonomicName>
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has nomenclatural priority over
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<taxonomicName id="07FE4D2AFFB5C32AFADBFE72FE96FE5D" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFADBFE72FE96FE5D" italics="true" pageId="8" pageNumber="45">H. subtriedroides</emphasis>
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</taxonomicName>
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, and thus
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<taxonomicName id="07FE4D2AFFB5C32AFE1FFE17FC0EFE5D" authority="Annandale, 1905" authorityName="Annandale" authorityYear="1905" box="[387,905,476,500]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFB5C32AFE1FFE17FD43FE5D" box="[387,708,476,500]" italics="true" pageId="8" pageNumber="45">Hemidactylus subtriedroides</emphasis>
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Annandale, 1905
|
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</taxonomicName>
|
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is here considered a junior subjective synonym of
|
||
<taxonomicName id="07FE4D2AFFB5C32AFF2AFDCBFE7FFDBE" authority="Tenkatei Lidth" authorityName="Tenkatei Lidth" box="[182,504,511,536]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="F28AEABBFFB5C32AFF2AFDCBFE32FDB1" box="[182,437,512,536]" italics="true" pageId="8" pageNumber="45">Hemidactylus Tenkatei</emphasis>
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Lidth
|
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</taxonomicName>
|
||
de
|
||
<bibRefCitation id="A46F4B58FFB5C32AFDBFFE34FD2BFDBE" author="Jeude" box="[547,684,511,535]" pageId="8" pageNumber="45" refString="Lidth de Jeude, T. W. V. (1895) Reptiles from Timor and the neighbouring islands. Notes from the Leyden Museum, 16, 119 - 127." type="journal article" year="1895">Jeude, 1895</bibRefCitation>
|
||
. The subsequent taxonomic section for this species is based on the morphological examination of
|
||
<taxonomicName id="07FE4D2AFFB5C32AFE6AFDEEFD3CFD95" box="[502,699,548,572]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="subtriedroides">
|
||
<emphasis id="F28AEABBFFB5C32AFE6AFDEEFD3CFD95" box="[502,699,548,572]" italics="true" pageId="8" pageNumber="45">H. subtriedroides</emphasis>
|
||
</taxonomicName>
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<typeStatus id="1F45880BFFB5C32AFD5FFDEEFD79FD94" box="[707,766,549,573]" pageId="8" pageNumber="45">types</typeStatus>
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and other material as the
|
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<typeStatus id="1F45880BFFB5C32AFBB5FDEEFB0CFD94" box="[1065,1163,549,573]" pageId="8" pageNumber="45" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="07FE4D2AFFB5C32AFB28FDEEFAB6FD95" box="[1204,1329,548,572]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="tenkatei">
|
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<emphasis id="F28AEABBFFB5C32AFB28FDEEFAB6FD95" box="[1204,1329,548,572]" italics="true" pageId="8" pageNumber="45">H. tenkatei</emphasis>
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</taxonomicName>
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were not directly examined in this study.
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |