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<document id="B9EF5A42BF31577EEBFD66A48BE09C98" ID-CLB-Dataset="120097" ID-DOI="10.11646/zootaxa.5174.3.4" ID-GBIF-Dataset="9ea0c18f-02e4-40b3-b1ff-27ed1b8eab15" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986947" ID-ZooBank="C75DE635-358E-40F0-A34B-AAA6BF1071AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660325937645" checkinUser="plazi" docAuthor="Nefediev, Pavel S." docDate="2022" docId="4663867A3E349C5174F2BECFFA24FF7B" docLanguage="en" docName="zootaxa.5174.3.4.pdf" docOrigin="Zootaxa 5174 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizoturanius krugovae Nefediev 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="274" masterDocId="BA5AFE023E309C5D7465BF00FFB8FF9C" masterDocTitle="Review of the genus Schizoturanius Verhoeff, 1931, with the description of a new species from the Altais, Asian Russia (Diplopoda: Polydesmida: Polydesmidae)" masterLastPageNumber="276" masterPageNumber="262" pageNumber="266" updateTime="1699366709504" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4D3DD5383BDA97C289C8FDF412D167D6">Review of the genus Schizoturanius Verhoeff, 1931, with the description of a new species from the Altais, Asian Russia (Diplopoda: Polydesmida: Polydesmidae)</mods:title>
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<mods:namePart id="E5762AC4A51E820D96364C0B1F9F7629">Nefediev, Pavel S.</mods:namePart>
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<mods:date id="4A514CAD05A2CB49E4227B7E437549EE">2022</mods:date>
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<mods:number id="32E73E4B6059AF93A550EC86DF749A28">2022-08-10</mods:number>
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<treatment id="4663867A3E349C5174F2BECFFA24FF7B" ID-DOI="http://doi.org/10.5281/zenodo.6989295" ID-GBIF-Taxon="199014938" ID-Zenodo-Dep="6989295" LSID="urn:lsid:plazi:treatment:4663867A3E349C5174F2BECFFA24FF7B" httpUri="http://treatment.plazi.org/id/4663867A3E349C5174F2BECFFA24FF7B" lastPageId="12" lastPageNumber="274" pageId="4" pageNumber="266">
<subSubSection id="86D064E73E349C5974F2BECFFD92FE76" box="[151,554,463,490]" pageId="4" pageNumber="266" type="nomenclature">
<paragraph id="CE75376C3E349C5974F2BECFFD92FE76" blockId="4.[151,554,463,526]" box="[151,554,463,490]" pageId="4" pageNumber="266">
<heading id="953D80003E349C5974F2BECFFD92FE76" bold="true" box="[151,554,463,490]" fontSize="11" level="1" pageId="4" pageNumber="266" reason="1">
<emphasis id="FCBEEB7E3E349C5974F2BECFFD92FE76" bold="true" box="[151,554,463,490]" pageId="4" pageNumber="266">
<taxonomicName id="09CA4CEF3E349C5974F2BECFFE7DFE76" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[151,453,463,490]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="266" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E349C5974F2BECFFE7DFE76" bold="true" box="[151,453,463,490]" italics="true" pageId="4" pageNumber="266">Schizoturanius krugovae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E78D56053E349C5975A8BED0FD92FE76" box="[461,554,464,490]" pageId="4" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E349C5974F2BEF4FE9AFD92" box="[151,290,500,526]" pageId="4" pageNumber="266" type="description">
<paragraph id="CE75376C3E349C5974F2BEF4FE9AFD92" blockId="4.[151,554,463,526]" box="[151,290,500,526]" pageId="4" pageNumber="266">
<figureCitation id="56F12BE93E349C5974F2BEF4FF64FD92" box="[151,220,500,526]" captionStart="FIGURES 12. 1" captionStartId="5.[151,264,670,694]" captionTargetBox="[152,1436,181,647]" captionTargetId="figure-18@5.[152,1436,181,647]" captionTargetPageId="5" captionText="FIGURES 12. 1. Schizoturanius krugovae sp. nov., male paratype, habitus (ZMUM Rd 4672). 2. Habitat of a male paratype ZMUM Rd 4672 (Betula pendula and Larix sibirica forest with bushes). Scale: 1 cm (Fig. 2 courtesy of T.M. Krugova)." figureDoi="http://doi.org/10.5281/zenodo.6986949" httpUri="https://zenodo.org/record/6986949/files/figure.png" pageId="4" pageNumber="266">Figs 1</figureCitation>
,
<figureCitation id="56F12BE93E349C59748FBEF4FE9AFD92" box="[234,290,500,526]" captionStart-0="FIGURES 39" captionStart-1="FIGURES 1021" captionStart-2="FIGURES 2232" captionStart-3="FIGURES 3344" captionStart-4="FIGURES 4546" captionStartId-0="6.[151,264,1824,1848]" captionStartId-1="7.[151,264,1434,1458]" captionStartId-2="8.[151,264,1824,1848]" captionStartId-3="9.[151,264,1674,1698]" captionStartId-4="10.[151,264,882,906]" captionTargetBox-0="[275,1313,181,1800]" captionTargetBox-1="[151,1435,181,1410]" captionTargetBox-2="[242,1345,181,1800]" captionTargetBox-3="[239,1349,181,1650]" captionTargetBox-4="[152,1436,181,858]" captionTargetId-0="figure-15@6.[275,1313,181,1800]" captionTargetId-1="figure-18@7.[151,1435,181,1410]" captionTargetId-2="figure-15@8.[242,1345,181,1800]" captionTargetId-3="figure-18@9.[239,1349,181,1650]" captionTargetId-4="figure-15@10.[152,1436,181,858]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." captionText-1="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." captionText-2="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." captionText-3="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." captionText-4="FIGURES 4546. Schizoturanius krugovae sp. nov., male paratype (ASU). 45 &amp; 46. Left gonopod, mesal and lateral views, respectively. Abbreviations: en, gonopod endomere; ep, external endomere process; ex, gonopod exomere; oip, oval inner plate; pu, pulvillus; sch, seminal chamber; sip, subtriangular inner plate. Scales: 500 μm." figureDoi-0="http://doi.org/10.5281/zenodo.6986953" figureDoi-1="http://doi.org/10.5281/zenodo.6986956" figureDoi-2="http://doi.org/10.5281/zenodo.6986958" figureDoi-3="http://doi.org/10.5281/zenodo.6986960" figureDoi-4="http://doi.org/10.5281/zenodo.6986962" httpUri-0="https://zenodo.org/record/6986953/files/figure.png" httpUri-1="https://zenodo.org/record/6986956/files/figure.png" httpUri-2="https://zenodo.org/record/6986958/files/figure.png" httpUri-3="https://zenodo.org/record/6986960/files/figure.png" httpUri-4="https://zenodo.org/record/6986962/files/figure.png" pageId="4" pageNumber="266">346</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E349C5974F2BD38FD3AFDCD" box="[151,642,568,593]" pageId="4" pageNumber="266" type="reference_group">
<paragraph id="CE75376C3E349C5974F2BD38FD3AFDCD" blockId="4.[151,642,568,593]" box="[151,642,568,593]" pageId="4" pageNumber="266">
<treatmentCitationGroup id="EEDA10423E349C5974F2BD38FD3AFDCD" box="[151,642,568,593]" pageId="4" pageNumber="266">
<taxonomicName id="09CA4CEF3E349C5974F2BD38FEDCFDCD" box="[151,356,568,593]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="266" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="FCBEEB7E3E349C5974F2BD38FE96FDCC" box="[151,302,568,592]" italics="true" pageId="4" pageNumber="266">Schizoturanius</emphasis>
sp. 2
</taxonomicName>
<treatmentCitation id="4F6B117D3E349C597518BD39FDC6FDCD" author="Krugova, T. M. &amp; Kuzmenkin, D. V. &amp; Nefediev, P. S." box="[381,638,568,593]" page="341" pageId="4" pageNumber="266" year="2018">
<bibRefCitation id="AA5B4A9D3E349C597518BD39FDC6FDCD" author="Krugova, T. M. &amp; Kuzmenkin, D. V. &amp; Nefediev, P. S." box="[381,638,568,593]" pageId="4" pageNumber="266" pagination="327 - 346" refId="ref8674" refString="Krugova, T. M., Kuzmenkin, D. V. &amp; Nefediev, P. S. (2018) Invertebrates of the Tigirek State Nature Reserve in 2017. Nature Chronicle of the Tigirek State Nature Reserve, 15 (1), 327 - 346. [in Russian]" type="journal article" year="2018">
Krugova
<emphasis id="FCBEEB7E3E349C5975B9BD39FDA8FDCC" box="[476,528,568,592]" italics="true" pageId="4" pageNumber="266">et al.</emphasis>
2018: 341
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E349C5974F2BD7EFE2BFBD5" pageId="4" pageNumber="266" type="materials_examined">
<paragraph id="CE75376C3E349C5974F2BD7EFC64FC99" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
<emphasis id="FCBEEB7E3E349C5974F2BD7EFEC7FD04" bold="true" box="[151,383,638,664]" pageId="4" pageNumber="266">Material examined.</emphasis>
<emphasis id="FCBEEB7E3E349C5975E0BD7FFE54FD05" box="[389,492,639,665]" italics="true" pageId="4" pageNumber="266">
<typeStatus id="117189CE3E349C5975E0BD7FFE54FD05" box="[389,492,639,665]" pageId="4" pageNumber="266" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="A8DBAFA93E349C597650BD7FFD36FD05" box="[565,654,639,665]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="266">ZMUM</collectionCode>
Rd 4671),
<collectingCountry id="B6DD77FC3E349C59776CBD7FFCEFFD05" box="[777,855,639,665]" name="Russia" pageId="4" pageNumber="266">Russia</collectingCountry>
, southwestern Siberia,
<collectingRegion id="0C0EF98E3E349C597031BD7FFB7FFD05" box="[1108,1223,639,665]" country="Russia" name="Altay" pageId="4" pageNumber="266">Altai Krai</collectingRegion>
, Zmeinogorsk District, left bank of Malaya Amelikha River,
<geoCoordinate id="ABFE51AB3E349C597614BDA2FD56FD21" box="[625,750,674,701]" degrees="51.0655" direction="north" orientation="latitude" pageId="4" pageNumber="266" precision="5" value="51.0655">51.0655°N</geoCoordinate>
,
<geoCoordinate id="ABFE51AB3E349C59769CBDA2FCCBFD21" box="[761,883,674,701]" degrees="82.7754" direction="east" orientation="longitude" pageId="4" pageNumber="266" precision="5" value="82.7754">82.7754°E</geoCoordinate>
,
<quantity id="09329A893E349C59771BBDA2FC7CFD21" box="[894,964,674,701]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="266" unit="m" value="570.0">570 m</quantity>
a.s.l., middle and low part of very steep N slope, chern taiga (
<taxonomicName id="09CA4CEF3E349C59750FBDC7FDBCFD7D" box="[362,516,711,737]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="FCBEEB7E3E349C59750FBDC7FDBCFD7D" box="[362,516,711,737]" italics="true" pageId="4" pageNumber="266">Abies sibirica</emphasis>
</taxonomicName>
forest with single
<taxonomicName id="09CA4CEF3E349C5976B5BDC7FCC2FD7D" box="[720,890,711,737]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="FCBEEB7E3E349C5976B5BDC7FCC2FD7D" box="[720,890,711,737]" italics="true" pageId="4" pageNumber="266">Betula pendula</emphasis>
</taxonomicName>
,
<taxonomicName id="09CA4CEF3E349C5977E2BDC7FB3FFD7D" box="[903,1159,711,737]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="FCBEEB7E3E349C5977E2BDC7FB3FFD7D" box="[903,1159,711,737]" italics="true" pageId="4" pageNumber="266">Caragana arborescens</emphasis>
</taxonomicName>
and
<taxonomicName id="09CA4CEF3E349C5970DEBDC7FAF3FD7C" box="[1211,1355,711,737]" class="Magnoliopsida" family="Rosaceae" genus="Padus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="avium">
<emphasis id="FCBEEB7E3E349C5970DEBDC7FAF3FD7C" box="[1211,1355,711,737]" italics="true" pageId="4" pageNumber="266">Padus avium</emphasis>
</taxonomicName>
,
<taxonomicName id="09CA4CEF3E349C59713CBDC7FA24FD7D" box="[1369,1436,711,737]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="genus">
<emphasis id="FCBEEB7E3E349C59713CBDC7FA24FD7D" box="[1369,1436,711,737]" italics="true" pageId="4" pageNumber="266">Carex</emphasis>
</taxonomicName>
plentifully), pitfall traps,
<date id="BA7411AC3E349C5975D7BDEBFD24FC99" box="[434,668,747,773]" pageId="4" pageNumber="266" value="2016-06-08" valueMax="2016-07-02" valueMin="2016-06-08">8 June 2 July 2016</date>
, collected by
<collectionCode id="A8DBAFA93E349C597750BDEBFCF0FC99" box="[821,840,747,773]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="266">T</collectionCode>
.M. Krugova.
</paragraph>
<paragraph id="CE75376C3E349C5974A2BC0FFE2BFBD5" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
<emphasis id="FCBEEB7E3E349C5974A2BC0FFE81FCB4" box="[199,313,783,808]" italics="true" pageId="4" pageNumber="266">
<typeStatus id="117189CE3E349C5974A2BC0FFE81FCB4" box="[199,313,783,808]" pageId="4" pageNumber="266" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="D8CCFCE53E349C59752FBC0FFE1DFCB5" box="[330,421,783,809]" pageId="4" pageNumber="266" type="male">2 males</specimenCount>
,
<specimenCount id="D8CCFCE53E349C5975D6BC0FFDA1FCB5" box="[435,537,783,809]" pageId="4" pageNumber="266" type="female">1 female</specimenCount>
,
<specimenCount id="D8CCFCE53E349C597642BC0EFD1BFCB5" box="[551,675,782,809]" pageId="4" pageNumber="266" type="juvenile">5 juveniles</specimenCount>
(
<collectionCode id="A8DBAFA93E349C5976D0BC0FFD4AFCB5" box="[693,754,783,809]" country="USA" lsid="urn:lsid:biocol.org:col:15600" name="Arizona State University" pageId="4" pageNumber="266" type="Herbarium">ASU</collectionCode>
), same data as
<typeStatus id="117189CE3E349C5977C6BC0FFBB1FCB5" box="[931,1033,783,809]" pageId="4" pageNumber="266" type="holotype">holotype</typeStatus>
;
<specimenCount id="D8CCFCE53E349C597073BC0FFBDFFCB5" box="[1046,1127,783,809]" pageId="4" pageNumber="266" type="male">1 male</specimenCount>
,
<specimenCount id="D8CCFCE53E349C597010BC0FFB5EFCB5" box="[1141,1254,783,809]" pageId="4" pageNumber="266" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="A8DBAFA93E349C59709DBC0FFA8DFCB5" box="[1272,1333,783,809]" country="USA" lsid="urn:lsid:biocol.org:col:15600" name="Arizona State University" pageId="4" pageNumber="266" type="Herbarium">ASU</collectionCode>
),
<collectingCountry id="B6DD77FC3E349C59712EBC0FFA21FCB5" box="[1355,1433,783,809]" name="Russia" pageId="4" pageNumber="266">Russia</collectingCountry>
, southwestern Siberia,
<collectingRegion id="0C0EF98E3E349C5975E9BC33FDB8FCD1" box="[396,512,819,845]" country="Russia" name="Altay" pageId="4" pageNumber="266">Altai Krai</collectingRegion>
, Charyshskoye District, left bank of Inya River, environs of abandon village of Kamyshenka, ca. 6 air-km SE of Tigirek,
<geoCoordinate id="ABFE51AB3E349C597623BC56FD7AFCED" box="[582,706,854,881]" degrees="51.1167" direction="north" orientation="latitude" pageId="4" pageNumber="266" precision="5" value="51.1167">51.1167°N</geoCoordinate>
,
<geoCoordinate id="ABFE51AB3E349C5976A8BC56FCFBFCED" box="[717,835,854,881]" degrees="83.1155" direction="east" orientation="longitude" pageId="4" pageNumber="266" precision="5" value="83.1155">83.1155°E</geoCoordinate>
(now included in
<collectionCode id="A8DBAFA93E349C597068BC57FBE1FCED" box="[1037,1113,855,881]" pageId="4" pageNumber="266">TSNR</collectionCode>
),
<quantity id="09329A893E349C597009BC56FB0AFCED" box="[1132,1202,854,881]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="4" pageNumber="266" unit="m" value="585.0">585 m</quantity>
a.s.l., N slope, chern taiga (
<taxonomicName id="09CA4CEF3E349C5974BBBC7BFEC1FC09" box="[222,377,891,917]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="FCBEEB7E3E349C5974BBBC7BFEC1FC09" box="[222,377,891,917]" italics="true" pageId="4" pageNumber="266">Abies sibirica</emphasis>
</taxonomicName>
forest with
<taxonomicName id="09CA4CEF3E349C59759ABC7BFD11FC09" box="[511,681,891,917]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="FCBEEB7E3E349C59759ABC7BFD11FC09" box="[511,681,891,917]" italics="true" pageId="4" pageNumber="266">Betula pendula</emphasis>
</taxonomicName>
), pitfall traps,
<date id="BA7411AC3E349C59772ABC7BFBFFFC09" box="[847,1095,891,917]" pageId="4" pageNumber="266" value="2016-06-18" valueMax="2016-07-03" valueMin="2016-06-18">18 June 3 July 2016</date>
, collected by
<collectionCode id="A8DBAFA93E349C597084BC7BFB4CFC09" box="[1249,1268,891,917]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="266">T</collectionCode>
.M. Krugova;
<specimenCount id="D8CCFCE53E349C5971EBBC7BFF75FC25" pageId="4" pageNumber="266" type="male">1 male</specimenCount>
(
<collectionCode id="A8DBAFA93E349C5974B9BC9FFE8DFC25" box="[220,309,927,953]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="266">ZMUM</collectionCode>
Rd 4672, voucher D113, GenBank
<accessionNumber id="D199AA8F3E349C5976A0BC9FFCF2FC25" box="[709,842,927,953]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM892667" pageId="4" pageNumber="266">
<collectionCode id="A8DBAFA93E349C5976A0BC9FFD4CFC25" box="[709,756,927,953]" country="New Zealand" name="Otago Museum" pageId="4" pageNumber="266">OM</collectionCode>
892667
</accessionNumber>
),
<collectingCountry id="B6DD77FC3E349C597738BC9FFC13FC25" box="[861,939,927,953]" name="Russia" pageId="4" pageNumber="266">Russia</collectingCountry>
, southwestern Siberia,
<collectingRegion id="0C0EF98E3E349C5970CEBC9FFAA7FC25" box="[1195,1311,927,953]" country="Russia" name="Altay" pageId="4" pageNumber="266">Altai Krai</collectingRegion>
, Krasnoshchiokovo District, ca. 6 air-km NW of Tigirek, right bank of Khankhara River,
<geoCoordinate id="ABFE51AB3E349C597062BCC2FB3CFC41" box="[1031,1156,962,989]" degrees="51.1931" direction="north" orientation="latitude" pageId="4" pageNumber="266" precision="5" value="51.1931">51.1931°N</geoCoordinate>
,
<geoCoordinate id="ABFE51AB3E349C5970EABCC3FAB2FC41" box="[1167,1290,963,989]" degrees="82.9800" direction="east" orientation="longitude" pageId="4" pageNumber="266" precision="5" value="82.98">82.9800°E</geoCoordinate>
,
<quantity id="09329A893E349C597171BCC3FAE3FC41" box="[1300,1371,963,989]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="4" pageNumber="266" unit="m" value="920.0">920 m</quantity>
a.s.l., upper part of N slope,
<taxonomicName id="09CA4CEF3E349C5975F4BCE7FD83FB9D" box="[401,571,999,1025]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="FCBEEB7E3E349C5975F4BCE7FD83FB9D" box="[401,571,999,1025]" italics="true" pageId="4" pageNumber="266">Betula pendula</emphasis>
</taxonomicName>
and
<taxonomicName id="09CA4CEF3E349C597614BCE7FCB2FB9D" box="[625,778,999,1025]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="266" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="FCBEEB7E3E349C597614BCE7FCB2FB9D" box="[625,778,999,1025]" italics="true" pageId="4" pageNumber="266">Larix sibirica</emphasis>
</taxonomicName>
forest with bushes, soil sample (
<quantity id="09329A893E349C597013BCE7FB60FB9D" box="[1142,1240,999,1025]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="4" pageNumber="266" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">010 cm</quantity>
deep),
<date id="BA7411AC3E349C59714FBCE7FF6AFBB9" pageId="4" pageNumber="266" value="2016-08-19">19 August 2016</date>
, collected by
<collectionCode id="A8DBAFA93E349C59750FBB0BFEC5FBB9" box="[362,381,1035,1061]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="266">T</collectionCode>
.M. Krugova, L.Yu. Gruntova,
<collectionCode id="A8DBAFA93E349C5976B4BB0BFD5DFBB9" box="[721,741,1035,1061]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="266" type="Museum">V</collectionCode>
.
<collectionCode id="A8DBAFA93E349C59768CBB0BFD45FBB9" box="[745,765,1035,1061]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="266" type="Museum">V</collectionCode>
. Zelenskii, K.
<collectionCode id="A8DBAFA93E349C5977F3BB0BFC14FBB9" box="[918,940,1035,1061]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="266" type="Museum">V</collectionCode>
. Smirnova, A.E. Pupkova, M.N. Terioshkina &amp;
<collectionCode id="A8DBAFA93E349C5974D1BB2FFF70FBD5" box="[180,200,1071,1097]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="266">R</collectionCode>
.
<collectionCode id="A8DBAFA93E349C5974A9BB2FFF5AFBD5" box="[204,226,1071,1097]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="266" type="Museum">V</collectionCode>
. Shcherbakova.
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E349C5974A2BB52FE88FAD9" pageId="4" pageNumber="266" type="diagnosis">
<paragraph id="CE75376C3E349C5974A2BB52FEEEFB61" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
<emphasis id="FCBEEB7E3E349C5974A2BB52FEFAFBF0" bold="true" box="[199,322,1106,1132]" pageId="4" pageNumber="266">Diagnosis.</emphasis>
Differs from all congeners mainly by the strongly elongated, slim and curved distal part of the gonopod endomere and its bifid apex (
<emphasis id="FCBEEB7E3E349C59766CBB77FD98FB0C" box="[521,544,1143,1168]" italics="true" pageId="4" pageNumber="266">vs</emphasis>
. shorter, subequal in size or less elongated, its apex never bifid in other species), and the postfemoral region of the endomere is with a thin, straight, subulate external process, which is almost nonserrate (
<emphasis id="FCBEEB7E3E349C597497BBBFFEB1FB44" box="[242,265,1215,1240]" italics="true" pageId="4" pageNumber="266">vs</emphasis>
. curved, hook-shaped, serrate or long, flat, not hook-shaped, serrate or even without an external process in other species).
</paragraph>
<paragraph id="CE75376C3E349C5974A2BA07FE88FAD9" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
<emphasis id="FCBEEB7E3E349C5974A2BA07FEACFABD" bold="true" box="[199,276,1287,1313]" pageId="4" pageNumber="266">Name.</emphasis>
Honours Mrs. Tatyana M. Krugova, the collector and a well-known Russian myrmecologist. Noun in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E349C5674A2BA4EFAE8FE81" lastPageId="11" lastPageNumber="273" pageId="4" pageNumber="266" type="description">
<paragraph id="CE75376C3E349C5974A2BA4EFD62FA2D" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
<emphasis id="FCBEEB7E3E349C5974A2BA4EFEE1FAF4" bold="true" box="[199,345,1358,1384]" pageId="4" pageNumber="266">Description.</emphasis>
Length
<quantity id="09329A893E349C5975D1BA4FFDEDFAF5" box="[436,597,1359,1385]" metricMagnitude="-2" metricUnit="m" metricValue="1.545" metricValueMax="1.63" metricValueMin="1.46" pageId="4" pageNumber="266" unit="mm" value="15.45" valueMax="16.3" valueMin="14.6">14.616.3 mm</quantity>
(males),
<quantity id="09329A893E349C5976D9BA4FFCA6FAF5" box="[700,798,1359,1385]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="4" pageNumber="266" unit="mm" value="17.3">17.3 mm</quantity>
(
<typeStatus id="117189CE3E349C59774EBA4FFC35FAF5" box="[811,909,1359,1385]" pageId="4" pageNumber="266" type="paratype">paratype</typeStatus>
female), width of midbody pro- and metazonae 1.0 and
<quantity id="09329A893E349C597494BA73FEF0FA11" box="[241,328,1395,1421]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="266" unit="mm" value="1.3">1.3 mm</quantity>
(males), 1.1 and
<quantity id="09329A893E349C59766EBA73FDD9FA11" box="[523,609,1395,1421]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="266" unit="mm" value="1.4">1.4 mm</quantity>
(
<typeStatus id="117189CE3E349C597615BA73FD6AFA11" box="[624,722,1395,1421]" pageId="4" pageNumber="266" type="paratype">paratype</typeStatus>
female), respectively.
<typeStatus id="117189CE3E349C5977B4BA73FB81FA11" box="[977,1081,1395,1421]" pageId="4" pageNumber="266" type="holotype">Holotype</typeStatus>
<quantity id="09329A893E349C597024BA73FB1DFA11" box="[1089,1189,1395,1421]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="4" pageNumber="266" unit="mm" value="14.6">14.6 mm</quantity>
long, 1.0 and
<quantity id="09329A893E349C597123BA73FA25FA11" box="[1350,1437,1395,1421]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="266" unit="mm" value="1.3">1.3 mm</quantity>
wide on midbody pro- and metazonae, respectively.
</paragraph>
<paragraph id="CE75376C3E349C5974A2BABBFF4CF9F9" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
Body moniliform, with 20 segments (including telson) in both sexes (
<figureCitation id="56F12BE93E349C5977AFBABBFBB3FA49" box="[970,1035,1467,1493]" captionStart="FIGURES 12. 1" captionStartId="5.[151,264,670,694]" captionTargetBox="[152,1436,181,647]" captionTargetId="figure-18@5.[152,1436,181,647]" captionTargetPageId="5" captionText="FIGURES 12. 1. Schizoturanius krugovae sp. nov., male paratype, habitus (ZMUM Rd 4672). 2. Habitat of a male paratype ZMUM Rd 4672 (Betula pendula and Larix sibirica forest with bushes). Scale: 1 cm (Fig. 2 courtesy of T.M. Krugova)." figureDoi="http://doi.org/10.5281/zenodo.6986949" httpUri="https://zenodo.org/record/6986949/files/figure.png" pageId="4" pageNumber="266">Fig. 1</figureCitation>
). Coloration in alcohol pale pinkish beige with darker anterior and posterior body parts, legs lighter, especially prefemora. Tegument moderately shining throughout; texture very delicately shagreened, alveolate, except convex metatergal surfaces. Prozonites (
<emphasis id="FCBEEB7E3E349C597158B903FAE1F981" bold="true" box="[1341,1369,1539,1565]" pageId="4" pageNumber="266">pz</emphasis>
) with
<specimenCount id="D8CCFCE53E349C5974F2B927FEBBF9DD" box="[151,259,1575,1601]" pageId="4" pageNumber="266" type="generic" typeStatus="types">two types</specimenCount>
of cuticular ornamentation: with or without serration along rear margin of alveolae (
<figureCitation id="56F12BE93E349C5970D7B927FABCF9DD" box="[1202,1284,1575,1601]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="4" pageNumber="266">Fig. 19</figureCitation>
, marked with arrows).
</paragraph>
<paragraph id="CE75376C3E349C5974A2B96FFEB0F86D" blockId="4.[151,1437,638,2033]" pageId="4" pageNumber="266">
Head mostly densely pubescent, with convex genae, clypeolabral region densely setose, but occiput bare (
<figureCitation id="56F12BE93E349C59710AB96FFF1FF931" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 9</figureCitation>
). Antennae moderately long, clavate, reaching past segment 3 dorsally. Length ratios of antennomeres 17 as 2.0:2.4:5.0:4.0:3.4:5.2:1, width ratios as 1.3:1.3:1.3:1.4:1.4:1.7:1, respectively (
<figureCitation id="56F12BE93E349C59707CB9B7FBE6F94D" box="[1049,1118,1719,1745]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 3</figureCitation>
); antennomeres 5 and 6 (
<emphasis id="FCBEEB7E3E349C5971E5B9B7FA24F94D" bold="true" box="[1408,1436,1719,1745]" pageId="4" pageNumber="266">a5</emphasis>
&amp;
<emphasis id="FCBEEB7E3E349C5974D0B9DAFF69F968" bold="true" box="[181,209,1754,1780]" pageId="4" pageNumber="266">a6</emphasis>
) each with a small, compact, distodorsal group of sensilla basiconica (
<emphasis id="FCBEEB7E3E349C597790B9DBFBB7F969" bold="true" box="[1013,1039,1755,1781]" pageId="4" pageNumber="266">sb</emphasis>
) (
<figureCitation id="56F12BE93E349C59704CB9DBFBD5F969" box="[1065,1133,1755,1781]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 3</figureCitation>
); antennomere 7 (
<emphasis id="FCBEEB7E3E349C597158B9DBFAE1F969" bold="true" box="[1341,1369,1755,1781]" pageId="4" pageNumber="266">a7</emphasis>
) with a terminal disc (
<emphasis id="FCBEEB7E3E349C597535B9FFFED1F885" bold="true" box="[336,361,1791,1817]" pageId="4" pageNumber="266">td</emphasis>
) bearing four sensory cones (
<emphasis id="FCBEEB7E3E349C5976DEB9FFFD6AF885" bold="true" box="[699,722,1791,1817]" pageId="4" pageNumber="266">sc</emphasis>
), each cone with 34 pores at its base (
<figureCitation id="56F12BE93E349C5970F0B9FFFB63F885" box="[1173,1243,1791,1817]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 4</figureCitation>
, marked with arrow), and a distodorsal group of sensilla basiconica spiniformia (
<emphasis id="FCBEEB7E3E349C597715B823FC2DF8A1" bold="true" box="[880,917,1827,1853]" pageId="4" pageNumber="266">sbs</emphasis>
) (
<figureCitation id="56F12BE93E349C5977CBB823FC4AF8A1" box="[942,1010,1826,1853]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 5</figureCitation>
). Mandibular gnathal lobe as follows: external tooth (
<emphasis id="FCBEEB7E3E349C597527B847FEE0F8FD" bold="true" box="[322,344,1863,1889]" pageId="4" pageNumber="266">et</emphasis>
) with large apical cusp and one smaller subapical cusp; internal tooth (
<emphasis id="FCBEEB7E3E349C597011B846FB3DF8FC" bold="true" box="[1140,1157,1862,1888]" pageId="4" pageNumber="266">it</emphasis>
) comb-shaped, with five apically rounded cusps; 6 rows of pectinate lamellae (
<emphasis id="FCBEEB7E3E349C597697B86BFCB1F819" bold="true" box="[754,777,1899,1925]" pageId="4" pageNumber="266">pl</emphasis>
) with simple, bacilliform teeth; intermediate area (
<emphasis id="FCBEEB7E3E349C597121B86AFAE2F818" bold="true" box="[1348,1370,1898,1924]" pageId="4" pageNumber="266">ia</emphasis>
) with serrate papillae; molar plate (
<emphasis id="FCBEEB7E3E349C5975BBB88FFDBDF835" bold="true" box="[478,517,1935,1961]" pageId="4" pageNumber="266">mp</emphasis>
) with transverse ridges, its anterior fringe (
<emphasis id="FCBEEB7E3E349C59778FB88EFBB9F834" bold="true" box="[1002,1025,1934,1960]" pageId="4" pageNumber="266">af</emphasis>
) with serrate spines apically (
<figureCitation id="56F12BE93E349C59712AB88FFF1FF851" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Figs 6, 7</figureCitation>
). Gnathochilarium (
<figureCitation id="56F12BE93E349C5975EEB8B3FE76F851" box="[395,462,1971,1997]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="4" pageNumber="266">Fig. 8</figureCitation>
) typical to
<taxonomicName id="09CA4CEF3E349C597635B8B3FD5CF851" authorityName="Pocock" authorityYear="1887" box="[592,740,1971,1997]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="266" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
, each stipes (
<emphasis id="FCBEEB7E3E349C59771EB8B3FC37F851" bold="true" box="[891,911,1971,1997]" pageId="4" pageNumber="266">st</emphasis>
) with 15 setae, each lamella lingualis (
<emphasis id="FCBEEB7E3E349C59712FB8B3FAE2F851" bold="true" box="[1354,1370,1971,1997]" pageId="4" pageNumber="266">ll</emphasis>
) with 68 setae.
</paragraph>
<caption id="9AB567E43E359C5874F2BD9EFAE2FD47" ID-DOI="http://doi.org/10.5281/zenodo.6986949" ID-Zenodo-Dep="6986949" httpUri="https://zenodo.org/record/6986949/files/figure.png" pageId="5" pageNumber="267" startId="5.[151,264,670,694]" targetBox="[152,1436,181,647]" targetPageId="5">
<paragraph id="CE75376C3E359C5874F2BD9EFAE2FD47" blockId="5.[151,1436,670,731]" pageId="5" pageNumber="267">
<emphasis id="FCBEEB7E3E359C5874F2BD9EFEF7FD2A" bold="true" box="[151,335,670,694]" pageId="5" pageNumber="267">FIGURES 12. 1</emphasis>
.
<taxonomicName id="09CA4CEF3E359C587539BD9FFDEFFD2B" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[348,599,671,695]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="267" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E359C587539BD9FFDEFFD2B" box="[348,599,671,695]" italics="true" pageId="5" pageNumber="267">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E359C58763BBD9FFD17FD2B" bold="true" box="[606,687,671,695]" pageId="5" pageNumber="267">
<taxonomicNameLabel id="E78D56053E359C58763BBD9FFD17FD2B" box="[606,687,671,695]" pageId="5" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype, habitus (ZMUM Rd 4672).
<emphasis id="FCBEEB7E3E359C587011BD9EFB39FD2A" bold="true" box="[1140,1153,670,694]" pageId="5" pageNumber="267">2</emphasis>
. Habitat of a male paratype ZMUM Rd 4672 (
<taxonomicName id="09CA4CEF3E359C587537BDC3FE54FD47" box="[338,492,707,731]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="267" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="FCBEEB7E3E359C587537BDC3FE54FD47" box="[338,492,707,731]" italics="true" pageId="5" pageNumber="267">Betula pendula</emphasis>
</taxonomicName>
and
<taxonomicName id="09CA4CEF3E359C587678BDC3FD1FFD47" box="[541,679,707,731]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="267" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="FCBEEB7E3E359C587678BDC3FD1FFD47" box="[541,679,707,731]" italics="true" pageId="5" pageNumber="267">Larix sibirica</emphasis>
</taxonomicName>
forest with bushes).
<emphasis id="FCBEEB7E3E359C58771EBDC2FC0BFD46" bold="true" box="[891,947,706,730]" pageId="5" pageNumber="267">Scale</emphasis>
: 1 cm (Fig. 2 courtesy of T.M. Krugova).
</paragraph>
</caption>
<paragraph id="CE75376C3E359C5874A2BC09FDEDFA1B" blockId="5.[151,1437,777,1991]" pageId="5" pageNumber="267">
In males, width of head (
<quantity id="09329A893E359C5875BBBC09FDDFFCBF" box="[478,615,777,803]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="5" pageNumber="267" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
broad)&gt; collum &lt;ring 2 = 3 &lt;4 &lt;5 &lt;6 = 12&gt; 13 = 16; thereafter body gradually, but significantly tapering towards telson (
<quantity id="09329A893E359C5876BABC2DFC8EFCDB" box="[735,822,813,839]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="267" unit="mm" value="0.6">0.6 mm</quantity>
broad). In the
<typeStatus id="117189CE3E359C5877BEBC2DFB83FCDB" box="[987,1083,813,839]" pageId="5" pageNumber="267" type="paratype">paratype</typeStatus>
female, width of head (
<quantity id="09329A893E359C587123BC2DFA25FCDB" box="[1350,1437,813,839]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="267" unit="mm" value="1.1">1.1 mm</quantity>
broad)&gt; collum &lt;ring 2 = 3 &lt;4 &lt;5 &lt;6 = 12&gt; 13 = 16; thereafter body rather rapidly tapering towards telson (
<quantity id="09329A893E359C58711DBC51FF7BFC13" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="267" unit="mm" value="0.7">0.7 mm</quantity>
broad). Collum (
<emphasis id="FCBEEB7E3E359C5875E0BC75FE1FFC13" bold="true" box="[389,423,885,911]" pageId="5" pageNumber="267">col</emphasis>
) transversely ovoid, ellipsoid (
<figureCitation id="56F12BE93E359C587761BC75FCF1FC13" box="[772,841,885,911]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="5" pageNumber="267">Fig. 9</figureCitation>
). Convex metatergal surfaces of rings 24 somewhat shorter than subsequent ones. Metatergal polygonal sculpture poorly-developed in adults, with three transverse rows of subsquarish bosses growing obliterate towards lateral portions of metaterga; juveniles with well-developed bosses on metaterga; each boss with a simple, very short tergal seta at its rear margin, more distinct on collum, as well as body rings 24 and 19, almost obliterate on midbody ones (
<figureCitation id="56F12BE93E359C5877C0BB05FB93FBBC" box="[933,1067,1029,1056]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Figs 1015</figureCitation>
). Collum with 1516 setae only in first row, and eight setae in each of two following rows (
<figureCitation id="56F12BE93E359C587758BB29FC3BFBDF" box="[829,899,1065,1091]" captionStart="FIGURES 39" captionStartId="6.[151,264,1824,1848]" captionTargetBox="[275,1313,181,1800]" captionTargetId="figure-15@6.[275,1313,181,1800]" captionTargetPageId="6" captionText="FIGURES 39. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3, 59) &amp; female paratype (ASU) (Fig. 4). 3. Antenna; 4. Terminal disc of antenna. 5. Distal part of antenna. 6 &amp; 7. Mandibular gnathal lobe, front and lateral views, respectively. 8. Gnathochilarium. 9. Head and collum, dorsal view. Abbreviations: a1a7, antennomeres 17; af, anterior fringe; col, collum; et, external tooth; ia, intermediate area; it, internal tooth; ll, lamella lingualis; mp, molar plate; pl, pectinate lamellae; sb, sensilla basiconica; sbs, sensilla basiconica spiniformia; sc, sensory cone; st, stipes; td, terminal disc; arrow shows pores on terminal disc. Scales: 100 μm (Figs 57), 200 μm (Fig. 4), 250 μm (Fig. 3), 400 μm (Fig. 8), 500 μm (Fig. 9)." figureDoi="http://doi.org/10.5281/zenodo.6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="5" pageNumber="267">Fig. 9</figureCitation>
), while following metaterga with three rows of eight setae each (
<figureCitation id="56F12BE93E359C58753CBB4DFE65FBF4" box="[345,477,1101,1128]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Figs 1015</figureCitation>
). Frontolateral corners of paratergite 2 slightly elongated anteriorly and pointed (
<figureCitation id="56F12BE93E359C58710ABB4DFF0DFB17" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Fig. 16</figureCitation>
, marked with arrow); lateral edges of subsequent swollen paraterga weakly rounded, smooth, with three setae on each side. Limbus, or caudal margin of metazona (
<emphasis id="FCBEEB7E3E359C58769DBB95FCA4FB33" bold="true" box="[760,796,1173,1199]" pageId="5" pageNumber="267">mz</emphasis>
) with tiny serrate spikes (
<emphasis id="FCBEEB7E3E359C587021BB95FBE2FB33" bold="true" box="[1092,1114,1173,1199]" pageId="5" pageNumber="267">ss</emphasis>
) (
<figureCitation id="56F12BE93E359C587016BB95FB7EFB33" box="[1139,1222,1173,1199]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Fig. 19</figureCitation>
), excluding telson. Metazona 1618 with slightly pointed tips of a couple of subsquarish bosses in row 3, closer to lateral sides of trunk, with an apical seta each (marked with arrow in
<figureCitation id="56F12BE93E359C5876D9BBDDFCA9FB64" box="[700,785,1245,1272]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Fig. 15</figureCitation>
). Epiproct process rather long, rounded at tip, carrying a couple of dorsal and lateral setae, and a group of four setiform spinnerets (
<emphasis id="FCBEEB7E3E359C5877BDBA01FC4AFA87" bold="true" box="[984,1010,1281,1307]" pageId="5" pageNumber="267">sp</emphasis>
) (some spinnerets may be broken off) (
<figureCitation id="56F12BE93E359C5874FABA25FE99FAA3" box="[159,289,1317,1343]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Figs 20, 21</figureCitation>
); it seems very likely that caudal projection of epiproct carrying four setiform spinnerets in all species of the genus (
<emphasis id="FCBEEB7E3E359C587553BA4AFEF5FAFF" box="[310,333,1354,1379]" italics="true" pageId="5" pageNumber="267">vs</emphasis>
. “two apical setae” in
<taxonomicName id="09CA4CEF3E359C587631BA49FD1DFAFF" authorityName="Mikhaljova" authorityYear="2013" box="[596,677,1353,1379]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="267" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="FCBEEB7E3E359C587631BA49FDDAFAFF" box="[596,610,1353,1379]" italics="true" pageId="5" pageNumber="267">S</emphasis>
.
<emphasis id="FCBEEB7E3E359C587617BA49FD1DFAFF" box="[626,677,1353,1379]" italics="true" pageId="5" pageNumber="267">levis</emphasis>
</taxonomicName>
and
<taxonomicName id="09CA4CEF3E359C5876BABA49FC2CFAFF" authorityName="Golovatch" authorityYear="1979" box="[735,916,1353,1379]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="267" phylum="Arthropoda" rank="species" species="dshungaricus">
<emphasis id="FCBEEB7E3E359C5876BABA49FD55FAFF" box="[735,749,1353,1379]" italics="true" pageId="5" pageNumber="267">S</emphasis>
.
<emphasis id="FCBEEB7E3E359C587698BA49FC2CFAFF" box="[765,916,1353,1379]" italics="true" pageId="5" pageNumber="267">dshungaricus</emphasis>
</taxonomicName>
). Paraprocts with a couple of setae each, and hypoproct with a pair of setae (
<figureCitation id="56F12BE93E359C587591BA6DFDFFFA1B" box="[500,583,1389,1415]" captionStart="FIGURES 1021" captionStartId="7.[151,264,1434,1458]" captionTargetBox="[151,1435,181,1410]" captionTargetId="figure-18@7.[151,1435,181,1410]" captionTargetPageId="7" captionText="FIGURES 1021. Schizoturanius krugovae sp. nov., male paratypes (ASU). 10 &amp; 13. Front body part, dorsal and lateral views, respectively. 11, 14 &amp; 17. Mid-body part, dorsal, lateral and ventral views, respectively. 12, 15 &amp; 18. Rear body part, dorsal, lateral and ventral views, respectively. 16. Segment 2, dorsal view. 19. Margin of prozonite and metazonite between rings 3 and 4, lateral view. 20 &amp; 21. Distal part of epiproct, lateral and ventral views, respectively. Abbreviations: mz, metazona; pz, prozona; sp, spinneret; ss, serrate spikes; arrows show details indicated in text. Scales: 50 μm (Fig. 19), 100 μm (Figs 20, 21), 400 μm (Fig. 16), 500 μm (Figs 12, 14, 15, 17, 18), 1 mm (Figs 10, 11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="5" pageNumber="267">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="CE75376C3E359C5874A2BA91FC75F8C7" blockId="5.[151,1437,777,1991]" pageId="5" pageNumber="267">
Legs generally rather long and slender, incrassate and slightly longer in males compared to females, podomere setose. Male podomeres ventrally with minute papillae (? sphaerotrichomes) on prefemora, femora, postfemora, tibiae and tarsi; prefemora papillate dorsally as well; in female, all podomeres with simple setae, these located mostly ventrally; prefemora clearly bulged dorsad only in males. Male leg-pair 1 (
<figureCitation id="56F12BE93E359C587020BAFDFB20F98B" box="[1093,1176,1533,1559]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 22</figureCitation>
) somewhat reduced as compared to following walking legs, coxae elongate ventrally, each coxa, prefemur, femur and postfemur with a single long seta caudoventrally; tibiae and most of tarsi densely pilose on ventral surface with relatively robust setae; each claw (
<emphasis id="FCBEEB7E3E359C58753BB969FED2F91F" bold="true" box="[350,362,1641,1667]" pageId="5" pageNumber="267">c</emphasis>
) with an accessory claw (
<emphasis id="FCBEEB7E3E359C5876EAB969FD11F91F" bold="true" box="[655,681,1641,1667]" pageId="5" pageNumber="267">ac</emphasis>
) ventrally (
<figureCitation id="56F12BE93E359C587748B969FCC6F91F" box="[813,894,1641,1667]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 23</figureCitation>
) (an additional minute claw has also been noted only in the original description of
<taxonomicName id="09CA4CEF3E359C587670B98DFD24F93B" authorityName="Timotheew" authorityYear="1897" box="[533,668,1677,1703]" class="Diplopoda" family="Polydesmidae" genus="Trachynotus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="267" phylum="Arthropoda" rank="species" species="dmitriewi">
<emphasis id="FCBEEB7E3E359C587670B98DFD9BF93B" box="[533,547,1677,1703]" italics="true" pageId="5" pageNumber="267">S</emphasis>
.
<emphasis id="FCBEEB7E3E359C587654B98DFD24F93B" box="[561,668,1677,1703]" italics="true" pageId="5" pageNumber="267">dmitriewi</emphasis>
</taxonomicName>
). Female leg-pair 1 subequal in size compared to following ones, all other characters as in male leg 1 (
<figureCitation id="56F12BE93E359C587677B9B1FDDFF957" box="[530,615,1713,1739]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 24</figureCitation>
). Leg-pair 2 (
<figureCitation id="56F12BE93E359C587765B9B1FC38F950" box="[768,896,1713,1740]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Figs 2527</figureCitation>
) in both sexes with an accessory claw (
<emphasis id="FCBEEB7E3E359C58715AB9B1FAE1F957" bold="true" box="[1343,1369,1713,1739]" pageId="5" pageNumber="267">ac</emphasis>
) ventrally (
<figureCitation id="56F12BE93E359C587484B9D5FE8DF973" box="[225,309,1749,1775]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 26</figureCitation>
). Gonapophysis (
<emphasis id="FCBEEB7E3E359C587592B9D5FDADF973" bold="true" box="[503,533,1749,1775]" pageId="5" pageNumber="267">gp</emphasis>
) on male coxa 2 (
<emphasis id="FCBEEB7E3E359C5876B9B9D5FCBDF973" bold="true" box="[732,773,1749,1775]" pageId="5" pageNumber="267">cx2</emphasis>
) relatively short, fimbriate apically (
<figureCitation id="56F12BE93E359C5870F9B9D5FB48F973" box="[1180,1264,1749,1775]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 27</figureCitation>
). Female coxae 2 (
<emphasis id="FCBEEB7E3E359C5874D3B9F9FF67F88F" bold="true" box="[182,223,1785,1811]" pageId="5" pageNumber="267">cx2</emphasis>
) flattened, with a lateral prominence (marked with arrow in
<figureCitation id="56F12BE93E359C5877EFB9F9FC66F88F" box="[906,990,1785,1811]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="5" pageNumber="267">Fig. 42</figureCitation>
). Other walking legs in both sexes with normal claws devoid of accessory claws. Pregonopodal male sternum 7 enlarged, flattened, double shield-shaped, obviously protecting telopodites of gonopods mechanically (
<figureCitation id="56F12BE93E359C587758B841FC79F8C7" box="[829,961,1857,1883]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Figs 2831</figureCitation>
).
</paragraph>
<paragraph id="CE75376C3E359C5874A2B865FDD5F85B" blockId="5.[151,1437,777,1991]" pageId="5" pageNumber="267">
Gonopod aperture of segment 7 enlarged, bean-shaped, with a transverse internal shelf anteriorly and a modest median projection on sternum posteriorly (
<figureCitation id="56F12BE93E359C587613B889FD72F83F" box="[630,714,1929,1955]" captionStart="FIGURES 2232" captionStartId="8.[151,264,1824,1848]" captionTargetBox="[242,1345,181,1800]" captionTargetId="figure-15@8.[242,1345,181,1800]" captionTargetPageId="8" captionText="FIGURES 2232. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 22 &amp; 23, 2532) &amp; female paratype (ASU) (Fig. 24). 22. Leg-pair 1, caudal view. 23. Terminal part of right tarsus 1, caudal view. 24. Leg-pair 1, caudal view. 25. Ring 3, caudal view. 26. Terminal part of right tarsus 2, front view. 27. Coxa of left leg 2, ventral view. 28 &amp; 30. Segment 6, caudal and ventral views, respectively. 29 &amp; 31. Sternum 7, caudal and ventral views, respectively. 32. Segment 7, ventral view (gonopods dissected, legs 9 removed). Abbreviations: ac, accessory claw; c, claw; cx2, coxa 2; gp, gonapophysis. Scales: 50 μm (Figs 23, 26), 100 μm (Fig. 27), 200 μm (Fig. 29), 300 μm (Fig. 31), 400 μm (Figs 22, 24), 500 μm (Figs 25, 30, 32), 1 mm (Fig. 28)." figureDoi="http://doi.org/10.5281/zenodo.6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="5" pageNumber="267">Fig. 32</figureCitation>
). Sternum 9 at base of coxa 9 (
<emphasis id="FCBEEB7E3E359C587040B889FBF5F83F" bold="true" box="[1061,1101,1929,1955]" pageId="5" pageNumber="267">cx9</emphasis>
) with a setose conical projection (
<emphasis id="FCBEEB7E3E359C5874B6B8ADFF57F85B" bold="true" box="[211,239,1965,1991]" pageId="5" pageNumber="267">cp</emphasis>
) extended ventrally (
<figureCitation id="56F12BE93E359C5875BBB8ADFDD9F85B" box="[478,609,1965,1991]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="5" pageNumber="267">Figs 33, 34</figureCitation>
).
</paragraph>
<caption id="9AB567E43E369C5B74F2B820FB54F870" ID-DOI="http://doi.org/10.5281/zenodo.6986953" ID-Zenodo-Dep="6986953" httpUri="https://zenodo.org/record/6986953/files/figure.png" pageId="6" pageNumber="268" startId="6.[151,264,1824,1848]" targetBox="[275,1313,181,1800]" targetPageId="6">
<paragraph id="CE75376C3E369C5B74F2B820FB54F870" blockId="6.[151,1436,1824,2028]" pageId="6" pageNumber="268">
<emphasis id="FCBEEB7E3E369C5B74F2B820FE84F8A4" bold="true" box="[151,316,1824,1848]" pageId="6" pageNumber="268">FIGURES 39.</emphasis>
<taxonomicName id="09CA4CEF3E369C5B7526B820FD87F8A4" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[323,575,1824,1848]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="268" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E369C5B7526B820FD87F8A4" box="[323,575,1824,1848]" italics="true" pageId="6" pageNumber="268">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E369C5B7623B820FD20F8A4" bold="true" box="[582,664,1824,1848]" pageId="6" pageNumber="268">
<taxonomicNameLabel id="E78D56053E369C5B7623B820FD20F8A4" box="[582,664,1824,1848]" pageId="6" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratypes (ASU) (Figs
<emphasis id="FCBEEB7E3E369C5B77ABB820FC63F8A4" bold="true" box="[974,987,1824,1848]" pageId="6" pageNumber="268">3</emphasis>
,
<emphasis id="FCBEEB7E3E369C5B778DB820FBB6F8A4" bold="true" box="[1000,1038,1824,1848]" pageId="6" pageNumber="268">59</emphasis>
) &amp; female paratype (ASU) (Fig.
<emphasis id="FCBEEB7E3E369C5B7103B820FACBF8A4" bold="true" box="[1382,1395,1824,1848]" pageId="6" pageNumber="268">4</emphasis>
).
<emphasis id="FCBEEB7E3E369C5B71ECB820FA2EF8A4" bold="true" box="[1417,1430,1824,1848]" pageId="6" pageNumber="268">3</emphasis>
. Antenna;
<emphasis id="FCBEEB7E3E369C5B749FB844FEBFF8C0" bold="true" box="[250,263,1860,1884]" pageId="6" pageNumber="268">4</emphasis>
. Terminal disc of antenna.
<emphasis id="FCBEEB7E3E369C5B767CB844FD9EF8C0" bold="true" box="[537,550,1860,1884]" pageId="6" pageNumber="268">5</emphasis>
. Distal part of antenna.
<emphasis id="FCBEEB7E3E369C5B777CB844FC9EF8C0" bold="true" box="[793,806,1860,1884]" pageId="6" pageNumber="268">6</emphasis>
&amp;
<emphasis id="FCBEEB7E3E369C5B7723B844FCEBF8C0" bold="true" box="[838,851,1860,1884]" pageId="6" pageNumber="268">7</emphasis>
. Mandibular gnathal lobe, front and lateral views, respectively.
<emphasis id="FCBEEB7E3E369C5B74B9B868FF51F81C" bold="true" box="[220,233,1896,1920]" pageId="6" pageNumber="268">8</emphasis>
. Gnathochilarium.
<emphasis id="FCBEEB7E3E369C5B75C8B868FE02F81C" bold="true" box="[429,442,1896,1920]" pageId="6" pageNumber="268">9</emphasis>
. Head and collum, dorsal view.
<emphasis id="FCBEEB7E3E369C5B769AB868FC20F81C" bold="true" box="[767,920,1896,1920]" pageId="6" pageNumber="268">Abbreviations</emphasis>
:
<emphasis id="FCBEEB7E3E369C5B77C0B868FC5DF81C" bold="true" box="[933,997,1896,1920]" pageId="6" pageNumber="268">a1a7</emphasis>
, antennomeres 17;
<emphasis id="FCBEEB7E3E369C5B70D2B868FB74F81C" bold="true" box="[1207,1228,1896,1920]" pageId="6" pageNumber="268">af</emphasis>
, anterior fringe;
<emphasis id="FCBEEB7E3E369C5B7112B868FA2EF81C" bold="true" box="[1399,1430,1896,1920]" pageId="6" pageNumber="268">col</emphasis>
, collum;
<emphasis id="FCBEEB7E3E369C5B748EB88CFF47F838" bold="true" box="[235,255,1932,1956]" pageId="6" pageNumber="268">et</emphasis>
, external tooth;
<emphasis id="FCBEEB7E3E369C5B75C2B88CFE03F838" bold="true" box="[423,443,1932,1956]" pageId="6" pageNumber="268">ia</emphasis>
, intermediate area;
<emphasis id="FCBEEB7E3E369C5B76E0B88CFD2CF838" bold="true" box="[645,660,1932,1956]" pageId="6" pageNumber="268">it</emphasis>
, internal tooth;
<emphasis id="FCBEEB7E3E369C5B775DB88CFCFEF838" bold="true" box="[824,838,1932,1956]" pageId="6" pageNumber="268">ll</emphasis>
, lamella lingualis;
<emphasis id="FCBEEB7E3E369C5B706CB88CFB94F838" bold="true" box="[1033,1068,1932,1956]" pageId="6" pageNumber="268">mp</emphasis>
, molar plate;
<emphasis id="FCBEEB7E3E369C5B70DEB88CFB68F838" bold="true" box="[1211,1232,1932,1956]" pageId="6" pageNumber="268">pl</emphasis>
, pectinate lamellae;
<emphasis id="FCBEEB7E3E369C5B74F2B8B0FF17F854" bold="true" box="[151,175,1968,1992]" pageId="6" pageNumber="268">sb</emphasis>
, sensilla basiconica;
<emphasis id="FCBEEB7E3E369C5B75E0B8B0FE1FF854" bold="true" box="[389,423,1968,1992]" pageId="6" pageNumber="268">sbs</emphasis>
, sensilla basiconica spiniformia;
<emphasis id="FCBEEB7E3E369C5B7692B8B0FCB4F854" bold="true" box="[759,780,1968,1992]" pageId="6" pageNumber="268">sc</emphasis>
, sensory cone;
<emphasis id="FCBEEB7E3E369C5B77CDB8B0FC02F854" bold="true" box="[936,954,1968,1992]" pageId="6" pageNumber="268">st</emphasis>
, stipes;
<emphasis id="FCBEEB7E3E369C5B7068B8B0FB9CF854" bold="true" box="[1037,1060,1968,1992]" pageId="6" pageNumber="268">td</emphasis>
, terminal disc; arrow shows pores on terminal disc.
<emphasis id="FCBEEB7E3E369C5B7542B8D4FED1F870" bold="true" box="[295,361,2004,2028]" pageId="6" pageNumber="268">Scales</emphasis>
: 100 μm (Figs
<emphasis id="FCBEEB7E3E369C5B7660B8D4FD93F870" bold="true" box="[517,555,2004,2028]" pageId="6" pageNumber="268">57</emphasis>
), 200 μm (Fig.
<emphasis id="FCBEEB7E3E369C5B76AEB8D4FD60F870" bold="true" box="[715,728,2004,2028]" pageId="6" pageNumber="268">4</emphasis>
), 250 μm (Fig.
<emphasis id="FCBEEB7E3E369C5B771DB8D4FC3DF870" bold="true" box="[888,901,2004,2028]" pageId="6" pageNumber="268">3</emphasis>
), 400 μm (Fig.
<emphasis id="FCBEEB7E3E369C5B7041B8D4FB89F870" bold="true" box="[1060,1073,2004,2028]" pageId="6" pageNumber="268">8</emphasis>
), 500 μm (Fig.
<emphasis id="FCBEEB7E3E369C5B70B4B8D4FB66F870" bold="true" box="[1233,1246,2004,2028]" pageId="6" pageNumber="268">9</emphasis>
).
</paragraph>
</caption>
<caption id="9AB567E43E379C5A74F2BA9AFC2BF9FA" ID-DOI="http://doi.org/10.5281/zenodo.6986956" ID-Zenodo-Dep="6986956" httpUri="https://zenodo.org/record/6986956/files/figure.png" pageId="7" pageNumber="269" startId="7.[151,264,1434,1458]" targetBox="[151,1435,181,1410]" targetPageId="7">
<paragraph id="CE75376C3E379C5A74F2BA9AFC2BF9FA" blockId="7.[151,1437,1434,1639]" pageId="7" pageNumber="269">
<emphasis id="FCBEEB7E3E379C5A74F2BA9AFEEBFA2E" bold="true" box="[151,339,1434,1458]" pageId="7" pageNumber="269">FIGURES 1021.</emphasis>
<taxonomicName id="09CA4CEF3E379C5A753CBA9AFDEBFA2E" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[345,595,1434,1458]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="269" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E379C5A753CBA9AFDEBFA2E" box="[345,595,1434,1458]" italics="true" pageId="7" pageNumber="269">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E379C5A763CBA9AFD11FA2E" bold="true" box="[601,681,1434,1458]" pageId="7" pageNumber="269">
<taxonomicNameLabel id="E78D56053E379C5A763CBA9AFD11FA2E" box="[601,681,1434,1458]" pageId="7" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratypes (ASU).
<emphasis id="FCBEEB7E3E379C5A77C7BA9AFC03FA2E" bold="true" box="[930,955,1434,1458]" pageId="7" pageNumber="269">10</emphasis>
&amp;
<emphasis id="FCBEEB7E3E379C5A77BEBA9AFC4CFA2E" bold="true" box="[987,1012,1434,1458]" pageId="7" pageNumber="269">13</emphasis>
. Front body part, dorsal and lateral views, respectively.
<emphasis id="FCBEEB7E3E379C5A757BBABEFE8EFA4A" bold="true" box="[286,310,1470,1494]" pageId="7" pageNumber="269">11</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7521BABEFEE5FA4A" bold="true" box="[324,349,1470,1494]" pageId="7" pageNumber="269">14</emphasis>
&amp;
<emphasis id="FCBEEB7E3E379C5A75E5BABEFE21FA4A" bold="true" box="[384,409,1470,1494]" pageId="7" pageNumber="269">17</emphasis>
. Mid-body part, dorsal, lateral and ventral views, respectively.
<emphasis id="FCBEEB7E3E379C5A7043BABEFB87FA4A" bold="true" box="[1062,1087,1470,1494]" pageId="7" pageNumber="269">12</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7028BABEFBDEFA4A" bold="true" box="[1101,1126,1470,1494]" pageId="7" pageNumber="269">15</emphasis>
&amp;
<emphasis id="FCBEEB7E3E379C5A70EFBABEFB1BFA4A" bold="true" box="[1162,1187,1470,1494]" pageId="7" pageNumber="269">18</emphasis>
. Rear body part, dorsal, lateral and ventral views, respectively.
<emphasis id="FCBEEB7E3E379C5A7649BAE2FDFDFA66" bold="true" box="[556,581,1506,1530]" pageId="7" pageNumber="269">16</emphasis>
. Segment 2, dorsal view.
<emphasis id="FCBEEB7E3E379C5A7736BAE2FCD4FA66" bold="true" box="[851,876,1506,1530]" pageId="7" pageNumber="269">19</emphasis>
. Margin of prozonite and metazonite between rings 3 and 4, lateral view.
<emphasis id="FCBEEB7E3E379C5A753AB906FEC0F982" bold="true" box="[351,376,1542,1566]" pageId="7" pageNumber="269">20</emphasis>
&amp;
<emphasis id="FCBEEB7E3E379C5A75FCB906FE0AF982" bold="true" box="[409,434,1542,1566]" pageId="7" pageNumber="269">21</emphasis>
. Distal part of epiproct, lateral and ventral views, respectively.
<emphasis id="FCBEEB7E3E379C5A705FB906FB6BF982" bold="true" box="[1082,1235,1542,1566]" pageId="7" pageNumber="269">Abbreviations</emphasis>
:
<emphasis id="FCBEEB7E3E379C5A7085B906FAB8F982" bold="true" box="[1248,1280,1542,1566]" pageId="7" pageNumber="269">mz</emphasis>
, metazona;
<emphasis id="FCBEEB7E3E379C5A7118B906FA2EF982" bold="true" box="[1405,1430,1542,1566]" pageId="7" pageNumber="269">pz</emphasis>
, prozona;
<emphasis id="FCBEEB7E3E379C5A7492B92AFEB7F9DE" bold="true" box="[247,271,1578,1602]" pageId="7" pageNumber="269">sp</emphasis>
, spinneret;
<emphasis id="FCBEEB7E3E379C5A75E3B92AFE22F9DE" bold="true" box="[390,410,1578,1602]" pageId="7" pageNumber="269">ss</emphasis>
, serrate spikes; arrows show details indicated in text.
<emphasis id="FCBEEB7E3E379C5A77A4B92AFBBBF9DE" bold="true" box="[961,1027,1578,1602]" pageId="7" pageNumber="269">Scales</emphasis>
: 50 μm (Fig.
<emphasis id="FCBEEB7E3E379C5A70F5B92AFB11F9DE" bold="true" box="[1168,1193,1578,1602]" pageId="7" pageNumber="269">19</emphasis>
), 100 μm (Figs
<emphasis id="FCBEEB7E3E379C5A712BB92AFADFF9DE" bold="true" box="[1358,1383,1578,1602]" pageId="7" pageNumber="269">20</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7111B92AFA35F9DE" bold="true" box="[1396,1421,1578,1602]" pageId="7" pageNumber="269">21</emphasis>
), 400 μm (Fig.
<emphasis id="FCBEEB7E3E379C5A7547B94EFE83F9FA" bold="true" box="[290,315,1614,1638]" pageId="7" pageNumber="269">16</emphasis>
), 500 μm (Figs
<emphasis id="FCBEEB7E3E379C5A75BAB94EFE40F9FA" bold="true" box="[479,504,1614,1638]" pageId="7" pageNumber="269">12</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7660B94EFDA6F9FA" bold="true" box="[517,542,1614,1638]" pageId="7" pageNumber="269">14</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A764EB94EFDFCF9FA" bold="true" box="[555,580,1614,1638]" pageId="7" pageNumber="269">15</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7634B94EFDD2F9FA" bold="true" box="[593,618,1614,1638]" pageId="7" pageNumber="269">17</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7612B94EFD28F9FA" bold="true" box="[631,656,1614,1638]" pageId="7" pageNumber="269">18</emphasis>
), 1 mm (Figs
<emphasis id="FCBEEB7E3E379C5A7745B94EFC81F9FA" bold="true" box="[800,825,1614,1638]" pageId="7" pageNumber="269">10</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A7723B94EFCE6F9FA" bold="true" box="[838,862,1614,1638]" pageId="7" pageNumber="269">11</emphasis>
,
<emphasis id="FCBEEB7E3E379C5A770EB94EFC3CF9FA" bold="true" box="[875,900,1614,1638]" pageId="7" pageNumber="269">13</emphasis>
).
</paragraph>
</caption>
<caption id="9AB567E43E389C5574F2B820FAC4F871" ID-DOI="http://doi.org/10.5281/zenodo.6986958" ID-Zenodo-Dep="6986958" httpUri="https://zenodo.org/record/6986958/files/figure.png" pageId="8" pageNumber="270" startId="8.[151,264,1824,1848]" targetBox="[242,1345,181,1800]" targetPageId="8">
<paragraph id="CE75376C3E389C5574F2B820FAC4F871" blockId="8.[151,1437,1824,2029]" pageId="8" pageNumber="270">
<emphasis id="FCBEEB7E3E389C5574F2B820FEEDF8A4" bold="true" box="[151,341,1824,1848]" pageId="8" pageNumber="270">FIGURES 2232.</emphasis>
<taxonomicName id="09CA4CEF3E389C557539B820FDEFF8A4" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[348,599,1824,1848]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="270" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E389C557539B820FDEFF8A4" box="[348,599,1824,1848]" italics="true" pageId="8" pageNumber="270">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E389C55763AB821FD08F8A5" bold="true" box="[607,688,1825,1849]" pageId="8" pageNumber="270">
<taxonomicNameLabel id="E78D56053E389C55763AB821FD08F8A5" box="[607,688,1825,1849]" pageId="8" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratypes (ASU) (Figs
<emphasis id="FCBEEB7E3E389C557786B820FC44F8A4" bold="true" box="[995,1020,1824,1848]" pageId="8" pageNumber="270">22</emphasis>
&amp;
<emphasis id="FCBEEB7E3E389C55707BB820FB8FF8A4" bold="true" box="[1054,1079,1824,1848]" pageId="8" pageNumber="270">23</emphasis>
,
<emphasis id="FCBEEB7E3E389C557020B820FB3CF8A4" bold="true" box="[1093,1156,1824,1848]" pageId="8" pageNumber="270">2532</emphasis>
) &amp; female paratype (ASU) (Fig.
<emphasis id="FCBEEB7E3E389C5574ABB844FF5FF8C0" bold="true" box="[206,231,1860,1884]" pageId="8" pageNumber="270">24</emphasis>
).
<emphasis id="FCBEEB7E3E389C557498B844FEAEF8C0" bold="true" box="[253,278,1860,1884]" pageId="8" pageNumber="270">22</emphasis>
. Leg-pair 1, caudal view.
<emphasis id="FCBEEB7E3E389C557645B844FD81F8C0" bold="true" box="[544,569,1860,1884]" pageId="8" pageNumber="270">23</emphasis>
. Terminal part of right tarsus 1, caudal view.
<emphasis id="FCBEEB7E3E389C55706DB844FB99F8C0" bold="true" box="[1032,1057,1860,1884]" pageId="8" pageNumber="270">24</emphasis>
. Leg-pair 1, caudal view.
<emphasis id="FCBEEB7E3E389C55714EB844FAFCF8C0" bold="true" box="[1323,1348,1860,1884]" pageId="8" pageNumber="270">25</emphasis>
. Ring 3, caudal view.
<emphasis id="FCBEEB7E3E389C55757EB868FE8CF81C" bold="true" box="[283,308,1896,1920]" pageId="8" pageNumber="270">26</emphasis>
. Terminal part of right tarsus 2, front view.
<emphasis id="FCBEEB7E3E389C55768BB868FCBFF81C" bold="true" box="[750,775,1896,1920]" pageId="8" pageNumber="270">27</emphasis>
. Coxa of left leg 2, ventral view.
<emphasis id="FCBEEB7E3E389C55703CB868FBCAF81C" bold="true" box="[1113,1138,1896,1920]" pageId="8" pageNumber="270">28</emphasis>
&amp;
<emphasis id="FCBEEB7E3E389C5570F7B868FB13F81C" bold="true" box="[1170,1195,1896,1920]" pageId="8" pageNumber="270">30</emphasis>
. Segment 6, caudal and ventral views, respectively.
<emphasis id="FCBEEB7E3E389C5575D5B88CFE71F838" bold="true" box="[432,457,1932,1956]" pageId="8" pageNumber="270">29</emphasis>
&amp;
<emphasis id="FCBEEB7E3E389C55758CB88CFDBAF838" bold="true" box="[489,514,1932,1956]" pageId="8" pageNumber="270">31</emphasis>
. Sternum 7, caudal and ventral views, respectively.
<emphasis id="FCBEEB7E3E389C557075B88CFB91F838" bold="true" box="[1040,1065,1932,1956]" pageId="8" pageNumber="270">32</emphasis>
. Segment 7, ventral view (gonopods dissected, legs 9 removed).
<emphasis id="FCBEEB7E3E389C5575CBB8B0FDFFF854" bold="true" box="[430,583,1968,1992]" pageId="8" pageNumber="270">Abbreviations</emphasis>
:
<emphasis id="FCBEEB7E3E389C557631B8B1FDD4F855" bold="true" box="[596,620,1969,1993]" pageId="8" pageNumber="270">ac</emphasis>
, accessory claw;
<emphasis id="FCBEEB7E3E389C557779B8B1FC9FF855" bold="true" box="[796,807,1969,1993]" pageId="8" pageNumber="270">c</emphasis>
, claw;
<emphasis id="FCBEEB7E3E389C55770AB8B0FC2CF854" bold="true" box="[879,916,1968,1992]" pageId="8" pageNumber="270">cx2</emphasis>
, coxa 2;
<emphasis id="FCBEEB7E3E389C55778AB8B1FBB2F855" bold="true" box="[1007,1034,1969,1993]" pageId="8" pageNumber="270">gp</emphasis>
, gonapophysis.
<emphasis id="FCBEEB7E3E389C5570C8B8B0FB57F854" bold="true" box="[1197,1263,1968,1992]" pageId="8" pageNumber="270">Scales</emphasis>
: 50 μm (Figs
<emphasis id="FCBEEB7E3E389C557118B8B0FA2EF854" bold="true" box="[1405,1430,1968,1992]" pageId="8" pageNumber="270">23</emphasis>
,
<emphasis id="FCBEEB7E3E389C5574F2B8D4FF08F870" bold="true" box="[151,176,2004,2028]" pageId="8" pageNumber="270">26</emphasis>
), 100 μm (Fig.
<emphasis id="FCBEEB7E3E389C557535B8D4FED1F870" bold="true" box="[336,361,2004,2028]" pageId="8" pageNumber="270">27</emphasis>
), 200 μm (Fig.
<emphasis id="FCBEEB7E3E389C55766FB8D4FD9BF870" bold="true" box="[522,547,2004,2028]" pageId="8" pageNumber="270">29</emphasis>
), 300 μm (Fig.
<emphasis id="FCBEEB7E3E389C5576A6B8D4FD64F870" bold="true" box="[707,732,2004,2028]" pageId="8" pageNumber="270">31</emphasis>
), 400 μm (Figs
<emphasis id="FCBEEB7E3E389C5577E5B8D4FC21F870" bold="true" box="[896,921,2004,2028]" pageId="8" pageNumber="270">22</emphasis>
,
<emphasis id="FCBEEB7E3E389C5577C3B8D4FC07F870" bold="true" box="[934,959,2004,2028]" pageId="8" pageNumber="270">24</emphasis>
), 500 μm (Figs
<emphasis id="FCBEEB7E3E389C557006B8D4FBC4F870" bold="true" box="[1123,1148,2004,2028]" pageId="8" pageNumber="270">25</emphasis>
,
<emphasis id="FCBEEB7E3E389C5570ECB8D4FB1AF870" bold="true" box="[1161,1186,2004,2028]" pageId="8" pageNumber="270">30</emphasis>
,
<emphasis id="FCBEEB7E3E389C5570CAB8D4FB70F870" bold="true" box="[1199,1224,2004,2028]" pageId="8" pageNumber="270">32</emphasis>
), 1 mm (Fig.
<emphasis id="FCBEEB7E3E389C557130B8D4FAD6F870" bold="true" box="[1365,1390,2004,2028]" pageId="8" pageNumber="270">28</emphasis>
).
</paragraph>
</caption>
<caption id="9AB567E43E399C5474F2B98AFBA5F87A" ID-DOI="http://doi.org/10.5281/zenodo.6986960" ID-Zenodo-Dep="6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="9" pageNumber="271" startId="9.[151,264,1674,1698]" targetBox="[239,1349,181,1650]" targetPageId="9">
<paragraph id="CE75376C3E399C5474F2B98AFBA5F87A" blockId="9.[151,1437,1674,2022]" pageId="9" pageNumber="271">
<emphasis id="FCBEEB7E3E399C5474F2B98AFEECF93E" bold="true" box="[151,340,1674,1698]" pageId="9" pageNumber="271">FIGURES 3344.</emphasis>
<taxonomicName id="09CA4CEF3E399C54753FB98AFDEDF93E" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[346,597,1674,1698]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="271" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E399C54753FB98AFDEDF93E" box="[346,597,1674,1698]" italics="true" pageId="9" pageNumber="271">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E399C547639B98AFD14F93E" bold="true" box="[604,684,1674,1698]" pageId="9" pageNumber="271">
<taxonomicNameLabel id="E78D56053E399C547639B98AFD14F93E" box="[604,684,1674,1698]" pageId="9" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratypes (ASU) (Figs
<emphasis id="FCBEEB7E3E399C5477B8B98AFBA4F93E" bold="true" box="[989,1052,1674,1698]" pageId="9" pageNumber="271">3341</emphasis>
) &amp; female paratype (Figs
<emphasis id="FCBEEB7E3E399C54714DB98AFADFF93E" bold="true" box="[1320,1383,1674,1698]" pageId="9" pageNumber="271">4244</emphasis>
).
<emphasis id="FCBEEB7E3E399C547118B98AFA2EF93E" bold="true" box="[1405,1430,1674,1698]" pageId="9" pageNumber="271">33</emphasis>
. Segment 7, caudal view (legs 9 omitted).
<emphasis id="FCBEEB7E3E399C54765AB9AEFDE0F95A" bold="true" box="[575,600,1710,1734]" pageId="9" pageNumber="271">34</emphasis>
. Sternum 9, caudal view (legs 9 omitted).
<emphasis id="FCBEEB7E3E399C54706FB9AEFB9BF95A" bold="true" box="[1034,1059,1710,1734]" pageId="9" pageNumber="271">35</emphasis>
. Distal part of right gonopod, caudal view.
<emphasis id="FCBEEB7E3E399C5474B1B9D2FF55F976" bold="true" box="[212,237,1746,1770]" pageId="9" pageNumber="271">36</emphasis>
. Segment 7 with gonopods, ventral view.
<emphasis id="FCBEEB7E3E399C5476FFB9D2FD0BF976" bold="true" box="[666,691,1746,1770]" pageId="9" pageNumber="271">37</emphasis>
&amp;
<emphasis id="FCBEEB7E3E399C5476B1B9D2FD55F976" bold="true" box="[724,749,1746,1770]" pageId="9" pageNumber="271">38</emphasis>
. Right gonopod, mesal and lateral views, respectively.
<emphasis id="FCBEEB7E3E399C547145B9D2FA81F976" bold="true" box="[1312,1337,1746,1770]" pageId="9" pageNumber="271">39</emphasis>
. External process of right gonopod endomere, caudal view.
<emphasis id="FCBEEB7E3E399C5476E2B9F6FD18F892" bold="true" box="[647,672,1782,1806]" pageId="9" pageNumber="271">40</emphasis>
. Tip of right gonopod endomere, mesal view.
<emphasis id="FCBEEB7E3E399C54700FB9F6FB3BF892" bold="true" box="[1130,1155,1782,1806]" pageId="9" pageNumber="271">41</emphasis>
. Pulvillus of right gonopod, lateral view.
<emphasis id="FCBEEB7E3E399C54757FB81AFE8BF8AE" bold="true" box="[282,307,1818,1842]" pageId="9" pageNumber="271">42</emphasis>
. Segment 3, ventral view (leg 2 almost omitted).
<emphasis id="FCBEEB7E3E399C547748B81AFCFEF8AE" bold="true" box="[813,838,1818,1842]" pageId="9" pageNumber="271">43</emphasis>
&amp;
<emphasis id="FCBEEB7E3E399C54770DB81AFC39F8AE" bold="true" box="[872,897,1818,1842]" pageId="9" pageNumber="271">44</emphasis>
. Vulva, front and lateral views, respectively.
<emphasis id="FCBEEB7E3E399C54712AB81AFF53F8CA" bold="true" pageId="9" pageNumber="271">Abbreviations</emphasis>
:
<emphasis id="FCBEEB7E3E399C54749CB83EFEADF8CA" bold="true" box="[249,277,1854,1878]" pageId="9" pageNumber="271">bu</emphasis>
, bursa of vulva;
<emphasis id="FCBEEB7E3E399C5475A5B83EFE60F8CA" bold="true" box="[448,472,1854,1878]" pageId="9" pageNumber="271">co</emphasis>
, mesal conical coxal outgrowth;
<emphasis id="FCBEEB7E3E399C547742B83EFCF8F8CA" bold="true" box="[807,832,1854,1878]" pageId="9" pageNumber="271">cp</emphasis>
, conical projection;
<emphasis id="FCBEEB7E3E399C547075B83EFB9EF8CA" bold="true" box="[1040,1062,1854,1878]" pageId="9" pageNumber="271">cr</emphasis>
, small crest;
<emphasis id="FCBEEB7E3E399C5470CBB83EFB6AF8CA" bold="true" box="[1198,1234,1854,1878]" pageId="9" pageNumber="271">cx2</emphasis>
&amp;
<emphasis id="FCBEEB7E3E399C547097B83EFAAFF8CA" bold="true" box="[1266,1303,1854,1878]" pageId="9" pageNumber="271">cx9</emphasis>
, coxae 2 and coxae 9;
<emphasis id="FCBEEB7E3E399C547496B862FEB6F8E6" bold="true" box="[243,270,1890,1914]" pageId="9" pageNumber="271">do</emphasis>
, digitiform coxal outgrowth;
<emphasis id="FCBEEB7E3E399C547658B862FDEEF8E6" bold="true" box="[573,598,1890,1914]" pageId="9" pageNumber="271">en</emphasis>
, gonopod endomere;
<emphasis id="FCBEEB7E3E399C547751B862FCF5F8E6" bold="true" box="[820,845,1890,1914]" pageId="9" pageNumber="271">ep</emphasis>
, external endomere process;
<emphasis id="FCBEEB7E3E399C54701DB862FB36F8E6" bold="true" box="[1144,1166,1890,1914]" pageId="9" pageNumber="271">er</emphasis>
, epigynal ridge;
<emphasis id="FCBEEB7E3E399C547159B862FAECF8E6" bold="true" box="[1340,1364,1890,1914]" pageId="9" pageNumber="271">ex</emphasis>
, gonopod exomere;
<emphasis id="FCBEEB7E3E399C54754CB886FEEEF802" bold="true" box="[297,342,1926,1950]" pageId="9" pageNumber="271">msp</emphasis>
, minute spinous process;
<emphasis id="FCBEEB7E3E399C54763AB886FDC2F802" bold="true" box="[607,634,1926,1950]" pageId="9" pageNumber="271">op</emphasis>
, operculum of vulva;
<emphasis id="FCBEEB7E3E399C547739B886FCC0F802" bold="true" box="[860,888,1926,1950]" pageId="9" pageNumber="271">pu</emphasis>
, pulvillus;
<emphasis id="FCBEEB7E3E399C547789B886FBBBF802" bold="true" box="[1004,1027,1926,1950]" pageId="9" pageNumber="271">ro</emphasis>
, rounded external coxal outgrowth;
<emphasis id="FCBEEB7E3E399C547112B886FA2EF802" bold="true" box="[1399,1430,1926,1950]" pageId="9" pageNumber="271">sip</emphasis>
, subtriangular inner plate; arrows show details indicated in text.
<emphasis id="FCBEEB7E3E399C547779B8AAFCE6F85E" bold="true" box="[796,862,1962,1986]" pageId="9" pageNumber="271">Scales</emphasis>
: 30 μm (Fig.
<emphasis id="FCBEEB7E3E399C54778FB8AAFBBBF85E" bold="true" box="[1002,1027,1962,1986]" pageId="9" pageNumber="271">40</emphasis>
), 50 μm (Figs
<emphasis id="FCBEEB7E3E399C5470FFB8AAFB0BF85E" bold="true" box="[1178,1203,1962,1986]" pageId="9" pageNumber="271">39</emphasis>
,
<emphasis id="FCBEEB7E3E399C5470A5B8AAFB61F85E" bold="true" box="[1216,1241,1962,1986]" pageId="9" pageNumber="271">41</emphasis>
), 100 μm (Figs
<emphasis id="FCBEEB7E3E399C547118B8AAFA2EF85E" bold="true" box="[1405,1430,1962,1986]" pageId="9" pageNumber="271">34</emphasis>
,
<emphasis id="FCBEEB7E3E399C5474F2B8CEFF08F87A" bold="true" box="[151,176,1998,2022]" pageId="9" pageNumber="271">35</emphasis>
), 200 μm (Fig.
<emphasis id="FCBEEB7E3E399C547535B8CEFED1F87A" bold="true" box="[336,361,1998,2022]" pageId="9" pageNumber="271">43</emphasis>
), 300 μm (Figs
<emphasis id="FCBEEB7E3E399C547668B8CEFD9EF87A" bold="true" box="[525,550,1998,2022]" pageId="9" pageNumber="271">36</emphasis>
,
<emphasis id="FCBEEB7E3E399C547656B8CEFDF4F87A" bold="true" box="[563,588,1998,2022]" pageId="9" pageNumber="271">44</emphasis>
), 400 μm (Figs
<emphasis id="FCBEEB7E3E399C547695B8CEFCB1F87A" bold="true" box="[752,777,1998,2022]" pageId="9" pageNumber="271">37</emphasis>
,
<emphasis id="FCBEEB7E3E399C547773B8CEFC97F87A" bold="true" box="[790,815,1998,2022]" pageId="9" pageNumber="271">38</emphasis>
,
<emphasis id="FCBEEB7E3E399C547759B8CEFCEDF87A" bold="true" box="[828,853,1998,2022]" pageId="9" pageNumber="271">42</emphasis>
), 500 μm (Fig.
<emphasis id="FCBEEB7E3E399C547790B8CEFBB6F87A" bold="true" box="[1013,1038,1998,2022]" pageId="9" pageNumber="271">33</emphasis>
).
</paragraph>
</caption>
<caption id="9AB567E43E3A9C5774F2BC72FB9AFC4F" ID-DOI="http://doi.org/10.5281/zenodo.6986962" ID-Zenodo-Dep="6986962" httpUri="https://zenodo.org/record/6986962/files/figure.png" pageId="10" pageNumber="272" startId="10.[151,264,882,906]" targetBox="[152,1436,181,858]" targetPageId="10">
<paragraph id="CE75376C3E3A9C5774F2BC72FB9AFC4F" blockId="10.[151,1436,882,979]" pageId="10" pageNumber="272">
<emphasis id="FCBEEB7E3E3A9C5774F2BC72FEEDFC16" bold="true" box="[151,341,882,906]" pageId="10" pageNumber="272">FIGURES 4546.</emphasis>
<taxonomicName id="09CA4CEF3E3A9C577538BC72FDE1FC16" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[349,601,882,906]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="272" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E3A9C577538BC72FDE1FC16" box="[349,601,882,906]" italics="true" pageId="10" pageNumber="272">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E3A9C577604BC72FD0AFC16" bold="true" box="[609,690,882,906]" pageId="10" pageNumber="272">
<taxonomicNameLabel id="E78D56053E3A9C577604BC72FD0AFC16" box="[609,690,882,906]" pageId="10" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (ASU).
<emphasis id="FCBEEB7E3E3A9C5777CFBC72FC7BFC16" bold="true" box="[938,963,882,906]" pageId="10" pageNumber="272">45</emphasis>
&amp;
<emphasis id="FCBEEB7E3E3A9C577783BC72FC47FC16" bold="true" box="[998,1023,882,906]" pageId="10" pageNumber="272">46</emphasis>
. Left gonopod, mesal and lateral views, respectively.
<emphasis id="FCBEEB7E3E3A9C57757ABC96FE00FC32" bold="true" box="[287,440,918,942]" pageId="10" pageNumber="272">Abbreviations</emphasis>
:
<emphasis id="FCBEEB7E3E3A9C5775ADBC96FE59FC32" bold="true" box="[456,481,918,942]" pageId="10" pageNumber="272">en</emphasis>
, gonopod endomere;
<emphasis id="FCBEEB7E3E3A9C5776A3BC96FD67FC32" bold="true" box="[710,735,918,942]" pageId="10" pageNumber="272">ep</emphasis>
, external endomere process;
<emphasis id="FCBEEB7E3E3A9C577077BC96FB92FC32" bold="true" box="[1042,1066,918,942]" pageId="10" pageNumber="272">ex</emphasis>
, gonopod exomere;
<emphasis id="FCBEEB7E3E3A9C577167BC96FA9CFC32" bold="true" box="[1282,1316,918,942]" pageId="10" pageNumber="272">oip</emphasis>
, oval inner plate;
<emphasis id="FCBEEB7E3E3A9C5774B3BCBAFF4AFC4E" bold="true" box="[214,242,954,978]" pageId="10" pageNumber="272">pu</emphasis>
, pulvillus;
<emphasis id="FCBEEB7E3E3A9C577500BCBAFE30FC4E" bold="true" box="[357,392,954,978]" pageId="10" pageNumber="272">sch</emphasis>
, seminal chamber;
<emphasis id="FCBEEB7E3E3A9C57762BBCBAFDD5FC4E" bold="true" box="[590,621,954,978]" pageId="10" pageNumber="272">sip</emphasis>
, subtriangular inner plate.
<emphasis id="FCBEEB7E3E3A9C57771BBCBAFC78FC4E" bold="true" box="[894,960,954,978]" pageId="10" pageNumber="272">Scales</emphasis>
: 500 μm.
</paragraph>
</caption>
<caption id="9AB567E43E3A9C5774F2BAB2FCC7FA72" ID-DOI="http://doi.org/10.5281/zenodo.6986966" ID-Zenodo-Dep="6986966" httpUri="https://zenodo.org/record/6986966/files/figure.png" pageId="10" pageNumber="272" startId="10.[151,250,1458,1482]" targetBox="[331,1226,1073,1385]" targetPageId="10">
<paragraph id="CE75376C3E3A9C5774F2BAB2FCC7FA72" blockId="10.[151,1437,1458,1518]" pageId="10" pageNumber="272">
<emphasis id="FCBEEB7E3E3A9C5774F2BAB2FE98FA56" bold="true" box="[151,288,1458,1482]" pageId="10" pageNumber="272">FIGURE 47.</emphasis>
Neighbor-joining tree of COI mtDNA gene sequence divergences (Tamura-Nei model) of studied
<taxonomicName id="09CA4CEF3E3A9C577163BAB2FF5BFA72" pageId="10" pageNumber="272">
<emphasis id="FCBEEB7E3E3A9C577163BAB2FA25FA56" box="[1286,1437,1458,1482]" italics="true" pageId="10" pageNumber="272">Schizoturanius</emphasis>
species
</taxonomicName>
; bootstrap support for branch nodes shown for 1,000 replications.
</paragraph>
</caption>
<paragraph id="CE75376C3E3A9C5774A2B91CFB7FF87A" blockId="10.[151,1437,1564,2023]" pageId="10" pageNumber="272">
Gonopods falcate, telopodites amber-yellowish,
<emphasis id="FCBEEB7E3E3A9C57768FB91DFC8BF9AA" box="[746,819,1565,1590]" italics="true" pageId="10" pageNumber="272">in situ</emphasis>
crossing each other, directed caudodorsally (
<figureCitation id="56F12BE93E3A9C57715CB91CFA28F9AA" box="[1337,1424,1564,1590]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 36</figureCitation>
). Gonopod coxa prominently enlarged, alveolate, with a rounded external outgrowth (
<emphasis id="FCBEEB7E3E3A9C57704BB941FBFFF9C7" bold="true" box="[1070,1095,1601,1627]" pageId="10" pageNumber="272">ro</emphasis>
) bearing an apical seta, as well as a digitiform outgrowth (
<emphasis id="FCBEEB7E3E3A9C5775DFB965FE6FF9E3" bold="true" box="[442,471,1637,1663]" pageId="10" pageNumber="272">do</emphasis>
) and a small crest (
<emphasis id="FCBEEB7E3E3A9C5776CCB965FD79F9E3" bold="true" box="[681,705,1637,1663]" pageId="10" pageNumber="272">cr</emphasis>
) located a little more caudally; cannula region with a mesal conical outgrowth (
<emphasis id="FCBEEB7E3E3A9C57757CB989FE8BF93F" bold="true" box="[281,307,1673,1699]" pageId="10" pageNumber="272">co</emphasis>
) (
<figureCitation id="56F12BE93E3A9C577529B988FE75F93E" box="[332,461,1672,1698]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Figs 3638</figureCitation>
). Basal (= prefemoral) part of telopodite densely setose. Distal part of telopodite deeply branched into an endomere (
<emphasis id="FCBEEB7E3E3A9C5775AAB9ADFE52F95B" bold="true" box="[463,490,1709,1735]" pageId="10" pageNumber="272">en</emphasis>
) and an exomere (
<emphasis id="FCBEEB7E3E3A9C5776D3B9ADFD68F95B" bold="true" box="[694,720,1709,1735]" pageId="10" pageNumber="272">ex</emphasis>
) (
<figureCitation id="56F12BE93E3A9C577683B9ACFCDFF95B" box="[742,871,1708,1735]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Figs 3538</figureCitation>
). Endomere significantly longer than exomere, with strongly elongated, slim and curved distal part anteriad; endomere tip bifid (
<figureCitation id="56F12BE93E3A9C5777B3B9D0FB90F977" box="[982,1064,1744,1771]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 40</figureCitation>
). Postfemoral region of endomere with a thin, straight, subulate external process (
<emphasis id="FCBEEB7E3E3A9C5776FCB9F5FD0CF893" bold="true" box="[665,692,1781,1807]" pageId="10" pageNumber="272">ep</emphasis>
), directed caudolaterad, almost devoid of serration (
<figureCitation id="56F12BE93E3A9C577088B9F4FA87F893" box="[1261,1343,1780,1807]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 39</figureCitation>
). Pulvillus (
<emphasis id="FCBEEB7E3E3A9C5774ADB819FF5FF8AF" bold="true" box="[200,231,1817,1843]" pageId="10" pageNumber="272">pu</emphasis>
) densely micropilose, supplied with a minute spinous process (
<emphasis id="FCBEEB7E3E3A9C5777C3B819FC6FF8AF" bold="true" box="[934,983,1817,1843]" pageId="10" pageNumber="272">msp</emphasis>
) (= thin process in
<bibRefCitation id="AA5B4A9D3E3A9C5770D4B818FAEDF8AF" author="Nefediev, P. S." box="[1201,1365,1816,1843]" pageId="10" pageNumber="272" pagination="331 - 339" refId="ref8973" refString="Nefediev, P. S. (2019) New records of the millipede genus Schizoturanius Verhoeff, 1931 from Asian Russia (Diplopoda: Polydesmida: Polydesmidae). Russian Entomological Journal, 28 (3), 331 - 339. https: // doi. org / 10.15298 / rusentj. 28.3.14" type="journal article" year="2019">Nefediev 2019</bibRefCitation>
) in its mid portion (
<figureCitation id="56F12BE93E3A9C57754DB83CFEC3F8CB" box="[296,379,1852,1879]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 41</figureCitation>
). Exomere thin, flattened on sides, with a blunt apex, a small, subtriangular, inner plate (
<emphasis id="FCBEEB7E3E3A9C577134B83CFACAF8CA" bold="true" box="[1361,1394,1852,1878]" pageId="10" pageNumber="272">sip</emphasis>
) (= small triangular blade in
<bibRefCitation id="AA5B4A9D3E3A9C5775CCB860FDF4F8E7" author="Nefediev, P. S." box="[425,588,1888,1915]" pageId="10" pageNumber="272" pagination="331 - 339" refId="ref8973" refString="Nefediev, P. S. (2019) New records of the millipede genus Schizoturanius Verhoeff, 1931 from Asian Russia (Diplopoda: Polydesmida: Polydesmidae). Russian Entomological Journal, 28 (3), 331 - 339. https: // doi. org / 10.15298 / rusentj. 28.3.14" type="journal article" year="2019">Nefediev 2019</bibRefCitation>
) at mesal edge of its middle part (
<figureCitation id="56F12BE93E3A9C5777A6B860FBAAF8E7" box="[963,1042,1888,1915]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 37</figureCitation>
) and a small, oval, inner plate close to exomeres base (
<emphasis id="FCBEEB7E3E3A9C57750FB884FE37F802" bold="true" box="[362,399,1924,1950]" pageId="10" pageNumber="272">oip</emphasis>
) (
<figureCitation id="56F12BE93E3A9C5775C3B884FE4EF803" box="[422,502,1924,1951]" captionStart="FIGURES 4546" captionStartId="10.[151,264,882,906]" captionTargetBox="[152,1436,181,858]" captionTargetId="figure-15@10.[152,1436,181,858]" captionTargetPageId="10" captionText="FIGURES 4546. Schizoturanius krugovae sp. nov., male paratype (ASU). 45 &amp; 46. Left gonopod, mesal and lateral views, respectively. Abbreviations: en, gonopod endomere; ep, external endomere process; ex, gonopod exomere; oip, oval inner plate; pu, pulvillus; sch, seminal chamber; sip, subtriangular inner plate. Scales: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.6986962" httpUri="https://zenodo.org/record/6986962/files/figure.png" pageId="10" pageNumber="272">Fig. 45</figureCitation>
) (the latter structure
<emphasis id="FCBEEB7E3E3A9C577688B884FCAAF802" bold="true" box="[749,786,1924,1950]" pageId="10" pageNumber="272">oip</emphasis>
is not visible in
<figureCitation id="56F12BE93E3A9C5777BCB884FB90F803" box="[985,1064,1924,1951]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="10" pageNumber="272">Fig. 37</figureCitation>
, being hidden by the inner wall of gonopod coxa). Seminal groove forming a loop level to telopodite division into an endo- and an exomere, ending up on a micropilose pulvillus supplied with a subterminal accessory seminal chamber (
<emphasis id="FCBEEB7E3E3A9C57704EB8CDFBE9F87B" bold="true" box="[1067,1105,1997,2023]" pageId="10" pageNumber="272">sch</emphasis>
) (
<figureCitation id="56F12BE93E3A9C57700CB8CCFB03F87A" box="[1129,1211,1996,2022]" captionStart="FIGURES 4546" captionStartId="10.[151,264,882,906]" captionTargetBox="[152,1436,181,858]" captionTargetId="figure-15@10.[152,1436,181,858]" captionTargetPageId="10" captionText="FIGURES 4546. Schizoturanius krugovae sp. nov., male paratype (ASU). 45 &amp; 46. Left gonopod, mesal and lateral views, respectively. Abbreviations: en, gonopod endomere; ep, external endomere process; ex, gonopod exomere; oip, oval inner plate; pu, pulvillus; sch, seminal chamber; sip, subtriangular inner plate. Scales: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.6986962" httpUri="https://zenodo.org/record/6986962/files/figure.png" pageId="10" pageNumber="272">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="CE75376C3E3B9C5674A2BF97FAE8FE81" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="273">
Vulvar aperture rounded; transverse epigynal ridge (
<emphasis id="FCBEEB7E3E3B9C56776BBF97FC9FFF2D" bold="true" box="[782,807,151,177]" pageId="11" pageNumber="273">er</emphasis>
) located behind leg-pair 2 on ventral surface of segment 3, well developed, slightly concave ventrally (
<figureCitation id="56F12BE93E3B9C5676CCBFBBFD47FF49" box="[681,767,187,213]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="11" pageNumber="273">Fig. 42</figureCitation>
). Vulvae bean-shaped, front surface of bursa (
<emphasis id="FCBEEB7E3E3B9C567177BFBBFA89FF49" bold="true" box="[1298,1329,187,213]" pageId="11" pageNumber="273">bu</emphasis>
) covered with two rows of relatively large transverse folds with smooth edges (marked with arrow in
<figureCitation id="56F12BE93E3B9C5670F2BFDFFB5EFF65" box="[1175,1254,223,249]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="11" pageNumber="273">Fig. 43</figureCitation>
) and long setae; operculum (
<emphasis id="FCBEEB7E3E3B9C56757ABE03FE85FE81" bold="true" box="[287,317,259,285]" pageId="11" pageNumber="273">op</emphasis>
) also with long setae (
<figureCitation id="56F12BE93E3B9C56765FBE03FD36FE81" box="[570,654,259,285]" captionStart="FIGURES 3344" captionStartId="9.[151,264,1674,1698]" captionTargetBox="[239,1349,181,1650]" captionTargetId="figure-18@9.[239,1349,181,1650]" captionTargetPageId="9" captionText="FIGURES 3344. Schizoturanius krugovae sp. nov., male paratypes (ASU) (Figs 3341) &amp; female paratype (Figs 4244). 33. Segment 7, caudal view (legs 9 omitted). 34. Sternum 9, caudal view (legs 9 omitted). 35. Distal part of right gonopod, caudal view. 36. Segment 7 with gonopods, ventral view. 37 &amp; 38. Right gonopod, mesal and lateral views, respectively. 39. External process of right gonopod endomere, caudal view. 40. Tip of right gonopod endomere, mesal view. 41. Pulvillus of right gonopod, lateral view. 42. Segment 3, ventral view (leg 2 almost omitted). 43 &amp; 44. Vulva, front and lateral views, respectively. Abbreviations: bu, bursa of vulva; co, mesal conical coxal outgrowth; cp, conical projection; cr, small crest; cx2 &amp; cx9, coxae 2 and coxae 9; do, digitiform coxal outgrowth; en, gonopod endomere; ep, external endomere process; er, epigynal ridge; ex, gonopod exomere; msp, minute spinous process; op, operculum of vulva; pu, pulvillus; ro, rounded external coxal outgrowth; sip, subtriangular inner plate; arrows show details indicated in text. Scales: 30 μm (Fig. 40), 50 μm (Figs 39, 41), 100 μm (Figs 34, 35), 200 μm (Fig. 43), 300 μm (Figs 36, 44), 400 μm (Figs 37, 38, 42), 500 μm (Fig. 33)." figureDoi="http://doi.org/10.5281/zenodo.6986960" httpUri="https://zenodo.org/record/6986960/files/figure.png" pageId="11" pageNumber="273">Fig. 44</figureCitation>
). Vulvae devoid of spiral structures at bottom of bursas gutter.
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E3B9C5674A2BE26FD9CFEF9" pageId="11" pageNumber="273" type="distribution">
<paragraph id="CE75376C3E3B9C5674A2BE26FD9CFEF9" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="273">
<emphasis id="FCBEEB7E3E3B9C5674A2BE26FED9FEDC" bold="true" box="[199,353,294,320]" pageId="11" pageNumber="273">Distribution.</emphasis>
Southern parts of the
<collectingRegion id="0C0EF98E3E3B9C56763BBE27FD69FEDD" box="[606,721,295,321]" country="Russia" name="Altay" pageId="11" pageNumber="273">Altai Krai</collectingRegion>
in the territory of the Tigirek State Nature Reserve and its immediate adjacent areas (
<figureCitation id="56F12BE93E3B9C5675C6BE4BFE40FEF9" box="[419,504,331,357]" captionStart="MAP 1" captionStartId="3.[151,207,1380,1404]" captionTargetBox="[151,1436,669,1357]" captionTargetId="figure-230@3.[151,1436,669,1357]" captionTargetPageId="3" captionText="MAP 1. Distributions of Schizoturanius species (circles): clavatipes (dark blue), dmitriewi (green), dshungaricus (light blue), kitabensis (violet), krugovae sp. nov. (red), levis (brown), montivagus (pink), strongylosomoides (orange), tabescens (yellow). Abbreviations: KG, Kyrgyzstan; UZ, Uzbekistan." figureDoi="http://doi.org/10.5281/zenodo.6989293" httpUri="https://zenodo.org/record/6989293/files/figure.png" pageId="11" pageNumber="273">Maps 1</figureCitation>
,
<figureCitation id="56F12BE93E3B9C567663BE4BFDAFFEF9" box="[518,535,331,357]" captionStart="MAP 2" captionStartId="11.[151,207,1385,1409]" captionTargetBox="[151,1436,453,1362]" captionTargetId="figure-143@11.[151,1436,453,1362]" captionTargetPageId="11" captionText="MAP 2. Distribution of Schizoturanius krugovae sp. nov. (red circle) within a strictly protected area and a buffer zone of the Tigirek State Nature Reserve (marked in green and yellow, respectively) and its immediate adjacent areas. Abbreviations: CN, China; KZ, Kazakhstan; MN, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.6989297" httpUri="https://zenodo.org/record/6989297/files/figure.png" pageId="11" pageNumber="273">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86D064E73E3B9C5174A2BE6FFA24FF7B" lastPageId="12" lastPageNumber="274" pageId="11" pageNumber="273" type="discussion">
<paragraph id="CE75376C3E3B9C5674A2BE6FFE4AFE31" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="273">
<emphasis id="FCBEEB7E3E3B9C5674A2BE6FFE83FE15" bold="true" box="[199,315,367,393]" pageId="11" pageNumber="273">Remarks.</emphasis>
A low-mountain species that inhabits the chern taiga and mixed forests at
<quantity id="09329A893E3B9C56701ABE6EFB46FE15" box="[1151,1278,366,393]" metricMagnitude="2" metricUnit="m" metricValue="7.45" metricValueMax="9.2" metricValueMin="5.7" pageId="11" pageNumber="273" unit="m" value="745.0" valueMax="920.0" valueMin="570.0">570920 m</quantity>
a.s.l. (
<figureCitation id="56F12BE93E3B9C56712CBE6FFA37FE15" box="[1353,1423,367,393]" captionStart="FIGURES 12. 1" captionStartId="5.[151,264,670,694]" captionTargetBox="[152,1436,181,647]" captionTargetId="figure-18@5.[152,1436,181,647]" captionTargetPageId="5" captionText="FIGURES 12. 1. Schizoturanius krugovae sp. nov., male paratype, habitus (ZMUM Rd 4672). 2. Habitat of a male paratype ZMUM Rd 4672 (Betula pendula and Larix sibirica forest with bushes). Scale: 1 cm (Fig. 2 courtesy of T.M. Krugova)." figureDoi="http://doi.org/10.5281/zenodo.6986949" httpUri="https://zenodo.org/record/6986949/files/figure.png" pageId="11" pageNumber="273">Fig. 2</figureCitation>
). Endemic of the Russian Altais.
</paragraph>
<caption id="9AB567E43E3B9C5674F2BA69FD8FFA55" ID-DOI="http://doi.org/10.5281/zenodo.6989297" ID-Zenodo-Dep="6989297" httpUri="https://zenodo.org/record/6989297/files/figure.png" pageId="11" pageNumber="273" startId="11.[151,207,1385,1409]" subCaptionStartIDs="11.[1236,1384,1421,1445]" subCaptionStarts="Abbr" targetBox="[151,1436,453,1362]" targetPageId="11">
<paragraph id="CE75376C3E3B9C5674F2BA69FD8FFA55" blockId="11.[151,1436,1385,1481]" pageId="11" pageNumber="273">
<emphasis id="FCBEEB7E3E3B9C5674F2BA69FF52FA1D" bold="true" box="[151,234,1385,1409]" pageId="11" pageNumber="273">MAP 2.</emphasis>
Distribution of
<taxonomicName id="09CA4CEF3E3B9C5675F5BA69FD34FA1D" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[400,652,1385,1409]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E3B9C5675F5BA69FD34FA1D" box="[400,652,1385,1409]" italics="true" pageId="11" pageNumber="273">Schizoturanius krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E3B9C5676F1BA69FD5DFA1D" bold="true" box="[660,741,1385,1409]" pageId="11" pageNumber="273">
<taxonomicNameLabel id="E78D56053E3B9C5676F1BA69FD5DFA1D" box="[660,741,1385,1409]" pageId="11" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red circle) within a strictly protected area and a buffer zone of the Tigirek State Nature Reserve (marked in green and yellow, respectively) and its immediate adjacent areas. Abbreviations:
<emphasis id="FCBEEB7E3E3B9C567114BA8DFA2EFA39" bold="true" box="[1393,1430,1421,1445]" pageId="11" pageNumber="273">CN</emphasis>
,
<collectingCountry id="B6DD77FC3E3B9C5674F2BAB1FF6FFA55" box="[151,215,1457,1481]" name="China" pageId="11" pageNumber="273">China</collectingCountry>
;
<emphasis id="FCBEEB7E3E3B9C567484BAB1FEBEFA55" bold="true" box="[225,262,1457,1481]" pageId="11" pageNumber="273">KZ</emphasis>
,
<collectingCountry id="B6DD77FC3E3B9C567577BAB1FE34FA55" box="[274,396,1457,1481]" name="Kazakhstan" pageId="11" pageNumber="273">Kazakhstan</collectingCountry>
;
<emphasis id="FCBEEB7E3E3B9C5675F3BAB1FE78FA55" bold="true" box="[406,448,1457,1481]" pageId="11" pageNumber="273">MN</emphasis>
,
<collectingCountry id="B6DD77FC3E3B9C5675A8BAB1FD8BFA55" box="[461,563,1457,1481]" name="Mongolia" pageId="11" pageNumber="273">Mongolia</collectingCountry>
.
</paragraph>
</caption>
<paragraph id="CE75376C3E3B9C5674F2BAF7FD79F9A9" blockId="11.[151,1437,1527,1590]" pageId="11" pageNumber="273">
Despite the obvious need for a full-scale revision of
<taxonomicName id="09CA4CEF3E3B9C567684BAF7FC3FF98D" baseAuthorityName="Mikhaljova" baseAuthorityYear="1993" box="[737,903,1527,1553]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="FCBEEB7E3E3B9C567684BAF7FC3FF98D" box="[737,903,1527,1553]" italics="true" pageId="11" pageNumber="273">Schizoturanius</emphasis>
</taxonomicName>
, the following provisional key can be proposed for all nine species of the genus currently known.
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B960FD2EF9EA" blockId="11.[151,1437,1632,2036]" box="[151,662,1632,1654]" pageId="11" pageNumber="273">1(16) Gonopod endomere with an external process.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B97EFD1FF90F" blockId="11.[151,1437,1632,2036]" box="[151,679,1662,1684]" pageId="11" pageNumber="273">2(11) External process of gonopod endomere serrate.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B99BFA25F92D" blockId="11.[151,1437,1632,2036]" box="[151,1437,1691,1713]" pageId="11" pageNumber="273">
3(4) Gonopod endomere longer than gonopod exomere…...............................................
<taxonomicName id="09CA4CEF3E3B9C567173B99BFA25F92D" authorityName="Lohmander" authorityYear="1933" box="[1302,1437,1691,1713]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="FCBEEB7E3E3B9C567173B99BFA9AF92C" box="[1302,1314,1691,1712]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C56714BB99CFA25F92D" box="[1326,1437,1692,1713]" italics="true" pageId="11" pageNumber="273">montivagus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B9B8FD6AF952" blockId="11.[151,1437,1632,2036]" box="[151,722,1720,1742]" pageId="11" pageNumber="273">4(3) Gonopod endomere shorter than gonopod exomere.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B9D5FC07F970" blockId="11.[151,1437,1632,2036]" box="[151,959,1749,1772]" pageId="11" pageNumber="273">5(8) External process of gonopod endomere curved, hook-shaped. Pulvillus bare.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B9F3FA24F894" blockId="11.[151,1437,1632,2036]" box="[151,1436,1779,1801]" pageId="11" pageNumber="273">
6(7) Lateral edge of gonopod endomere smooth. A small inner plate on gonopod exomere present…..............
<taxonomicName id="09CA4CEF3E3B9C567146B9F3FA24F894" authorityName="Stuxberg" authorityYear="1876" box="[1315,1436,1779,1800]" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="clavatipes">
<emphasis id="FCBEEB7E3E3B9C567146B9F3FA97F894" box="[1315,1327,1779,1800]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C56715EB9F3FA24F894" box="[1339,1436,1779,1800]" italics="true" pageId="11" pageNumber="273">clavatipes</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B810FA24F8B9" blockId="11.[151,1437,1632,2036]" box="[151,1436,1808,1830]" pageId="11" pageNumber="273">
7(6) Lateral edge of gonopod endomere serrate. A small inner plate on gonopod exomere absent.................
<taxonomicName id="09CA4CEF3E3B9C56714DB811FA24F8B9" authorityName="Timotheew" authorityYear="1897" box="[1320,1436,1808,1830]" class="Diplopoda" family="Polydesmidae" genus="Trachynotus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="dmitriewi">
<emphasis id="FCBEEB7E3E3B9C56714DB811FA8CF8BA" box="[1320,1332,1809,1830]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C567125B810FA24F8B9" box="[1344,1436,1808,1829]" italics="true" pageId="11" pageNumber="273">dmitriewi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B82DFC44F8D8" blockId="11.[151,1437,1632,2036]" box="[151,1020,1837,1860]" pageId="11" pageNumber="273">8(5) External process of gonopod endomere and pulvillus different or poorly described.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B84BFA24F8FC" blockId="11.[151,1437,1632,2036]" box="[151,1436,1867,1889]" pageId="11" pageNumber="273">
9(10) External process of gonopod endomere long, flat, not hook-shaped. Pulvillus micropilose.......................
<taxonomicName id="09CA4CEF3E3B9C56713DB84BFA24F8FC" authorityName="Mikhaljova" authorityYear="2013" box="[1368,1436,1867,1888]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="FCBEEB7E3E3B9C56713DB84BFADCF8FC" box="[1368,1380,1867,1888]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C567115B84BFA24F8FC" box="[1392,1436,1867,1888]" italics="true" pageId="11" pageNumber="273">levis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B868FA24F807" blockId="11.[151,1437,1632,2036]" pageId="11" pageNumber="273">
10(9) Shape of external process of gonopod endomere and pubescence of pulvillus uncertain (in the poor original description)..............................................................................................
<taxonomicName id="09CA4CEF3E3B9C567141B886FA24F807" baseAuthorityName="Gulicka" baseAuthorityYear="1963" box="[1316,1436,1926,1947]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="kitabensis">
<emphasis id="FCBEEB7E3E3B9C567141B886FA88F807" box="[1316,1328,1926,1947]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C567159B886FA24F807" box="[1340,1436,1926,1947]" italics="true" pageId="11" pageNumber="273">kitabensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B8A3FBA0F825" blockId="11.[151,1437,1632,2036]" box="[151,1048,1955,1977]" pageId="11" pageNumber="273">11(2) External process of gonopod endomere devoid or almost devoid of serration, straight.</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B8C0FA24F849" blockId="11.[151,1437,1632,2036]" box="[151,1436,1984,2006]" pageId="11" pageNumber="273">
12(13) Gonopod endomere subequal in size to gonopod exomere.....................................
<taxonomicName id="09CA4CEF3E3B9C5670B4B8C1FA24F849" baseAuthorityName="Attems" baseAuthorityYear="1904" box="[1233,1436,1984,2006]" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="273" phylum="Arthropoda" rank="species" species="strongylosomoides">
<emphasis id="FCBEEB7E3E3B9C5670B4B8C1FB65F84A" box="[1233,1245,1985,2006]" italics="true" pageId="11" pageNumber="273">S</emphasis>
.
<emphasis id="FCBEEB7E3E3B9C56708CB8C0FA24F849" box="[1257,1436,1984,2005]" italics="true" pageId="11" pageNumber="273">strongylosomoides</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3B9C5674F2B8DEFD74F86F" blockId="11.[151,1437,1632,2036]" box="[151,716,2014,2036]" pageId="11" pageNumber="273">13(12) Gonopod endomere longer than gonopod exomere.</paragraph>
<paragraph id="CE75376C3E3C9C5174F2BF97FA24FF30" blockId="12.[151,1436,151,232]" box="[151,1436,151,173]" pageId="12" pageNumber="274">
14(15) Gonopod endomere slightly longer than gonopod exomere.........................................
<taxonomicName id="09CA4CEF3E3C9C517166BF98FA24FF30" authorityName="Golovatch" authorityYear="1979" box="[1283,1436,151,173]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="274" phylum="Arthropoda" rank="species" species="dshungaricus">
<emphasis id="FCBEEB7E3E3C9C517166BF98FAB7FF31" box="[1283,1295,152,173]" italics="true" pageId="12" pageNumber="274">S</emphasis>
.
<emphasis id="FCBEEB7E3E3C9C51717EBF97FA24FF30" box="[1307,1436,151,172]" italics="true" pageId="12" pageNumber="274">dshungaricus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="CE75376C3E3C9C5174F2BFB5FA24FF56" blockId="12.[151,1436,151,232]" box="[151,1436,181,203]" pageId="12" pageNumber="274">
15(14) Gonopod endomere significantly longer than gonopod exomere. Endomere tip bifid.................
<taxonomicName id="09CA4CEF3E3C9C5170BDBFB5FAF1FF56" authority="Nefediev, 2022" authorityName="Nefediev" authorityYear="2022" box="[1240,1353,181,202]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="274" phylum="Arthropoda" rank="species" species="krugovae" status="sp. nov.">
<emphasis id="FCBEEB7E3E3C9C5170BDBFB5FB5CFF56" box="[1240,1252,181,202]" italics="true" pageId="12" pageNumber="274">S</emphasis>
.
<emphasis id="FCBEEB7E3E3C9C517095BFB5FAF1FF56" box="[1264,1353,181,202]" italics="true" pageId="12" pageNumber="274">krugovae</emphasis>
</taxonomicName>
<emphasis id="FCBEEB7E3E3C9C517135BFB5FA24FF56" bold="true" box="[1360,1436,181,202]" pageId="12" pageNumber="274">
<taxonomicNameLabel id="E78D56053E3C9C517135BFB5FA24FF56" box="[1360,1436,181,202]" pageId="12" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="CE75376C3E3C9C5174F2BFD2FA24FF7B" blockId="12.[151,1436,151,232]" box="[151,1436,210,232]" pageId="12" pageNumber="274">
16(1) Gonopod endomere without an external process.....................................................
<taxonomicName id="09CA4CEF3E3C9C517142BFD2FA24FF7B" baseAuthorityName="Stuxberg" baseAuthorityYear="1876" box="[1319,1436,210,232]" class="Diplopoda" family="Polydesmidae" genus="Schizoturanius" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="274" phylum="Arthropoda" rank="species" species="tabescens">
<emphasis id="FCBEEB7E3E3C9C517142BFD2FA8BFF7B" box="[1319,1331,210,231]" italics="true" pageId="12" pageNumber="274">S</emphasis>
.
<emphasis id="FCBEEB7E3E3C9C51715ABFD2FA24FF7B" box="[1343,1436,210,231]" italics="true" pageId="12" pageNumber="274">tabescens</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>