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<mods:title id="E84549C0BD45D432D57643FF56AE6A21">Review of the Stomodes tolutarius species group (Coleoptera: Curculionidae: Entiminae: Otiorhynchini) from the Balkan Peninsula with description of two new species from Greece</mods:title>
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<treatment id="3E6F2970F5098A74FF35FD0806D9FA59" ID-DOI="http://doi.org/10.5281/zenodo.5964270" ID-GBIF-Taxon="148651500" ID-Zenodo-Dep="5964270" LSID="urn:lsid:plazi:treatment:3E6F2970F5098A74FF35FD0806D9FA59" httpUri="http://treatment.plazi.org/id/3E6F2970F5098A74FF35FD0806D9FA59" lastPageNumber="123" pageId="12" pageNumber="123">
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<paragraph id="B6799866F5098A74FF35FD0801A5FFF8" blockId="12.[151,736,727,754]" box="[151,736,727,754]" pageId="12" pageNumber="123">
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Key to the species of
<taxonomicName id="71C6E3E5F5098A74FE3EFD070175FFFB" ID-CoL="6ZWQB" authorityName=": Wagner" authorityYear="1912" box="[412,560,727,753]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="tolutarius">
<emphasis id="84B24474F5098A74FE3EFD070175FFFB" bold="true" box="[412,560,727,753]" italics="true" pageId="12" pageNumber="123">S. tolutarius</emphasis>
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species group
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<keyLead id="0D373B53F5098A74FF35FCCD06D9FE5B" pageId="12" pageNumber="123">1. Elytra (Figs 23, 31) with apex not visible in dorsal view, hidden by slope overhanging apex; pronotum with punctures of two different sizes, with unevenly scattered coarse and conspicuously smaller and fine punctures.......................... 2</keyLead>
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<paragraph id="B6799866F5098A74FF35FC8906DEFE81" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35FC8906DEFE81" pageId="12" pageNumber="123">- Elytra (Fig. 9) with apex visible in dorsal view; pronotum with punctures of equal size, in some species weakly variable in size but never distinctly small............................................................................... 3</keyLead>
</paragraph>
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<keyStep id="0D3280C3F5098A74FF35FC4306D9F90B" pageId="12" pageNumber="123">
<paragraph id="B6799866F5098A74FF35FC4306DEFECC" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35FC4306DEFECC" pageId="12" pageNumber="123">
2. Rostrum 1.611.67× as wide as long; pronotum 1.031.06× as wide as long; elytra (Fig. 31) laterally flat, slope overhanging apex, evenly descending downwards; tarsi with segment 2 1.11.2× as wide as long..............
<taxonomicName id="71C6E3E5F5098A74FB11FC6006DEFECC" ID-CoL="6ZWQB" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1826" box="[1203,1435,944,966]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="tolutarius">
<emphasis id="84B24474F5098A74FB11FC60066DFECC" box="[1203,1320,944,966]" italics="true" pageId="12" pageNumber="123">S. tolutarius</emphasis>
Schoenherr
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<paragraph id="B6799866F5098A74FF35FC1E06D9F90B" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
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- Rostrum 1.85× as wide as long; pronotum 1.11× as wide as long; elytra (Fig. 23) with slope overhanging apex, rounded and then perpendicularly falling downwards; tarsi with segment 2 1.4× as wide as long...............
<taxonomicName id="71C6E3E5F5098A74FB10FC3B06D9F90B" ID-CoL="6ZWS8" authority="Hoffmann" authorityName="Hoffmann" authorityYear="1956" box="[1202,1436,1003,1025]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="marocanus">
<emphasis id="84B24474F5098A74FB10FC3B0671F90B" box="[1202,1332,1003,1025]" italics="true" pageId="12" pageNumber="123">S. marocanus</emphasis>
Hoffmann
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<paragraph id="B6799866F5098A74FF35FBD806D9F97C" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35FBD806D9F97C" pageId="12" pageNumber="123">
3. Pronotum shiny, with very fine and sparse punctures; distance between two nearest pronotal punctures greater than diameter of a puncture; elytra at most 1.68× as long as wide, with semierect setae on disc, and with finely punctate striae; distance between elytral punctures greater than a puncture diameter; interstriae distinctly wider than striae; funicle segments in females wider than long................................................................................
<taxonomicName id="71C6E3E5F5098A74FB5BFBB006D9F97C" ID-CoL="6ZX3W" authority="Lona" authorityName="Lona" authorityYear="1922" box="[1273,1436,1120,1142]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="muelleri">
<emphasis id="84B24474F5098A74FB5BFBB00621F97C" box="[1273,1380,1120,1142]" italics="true" pageId="12" pageNumber="123">S. muelleri</emphasis>
Lona
</taxonomicName>
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<keyLead id="0D373B53F5098A74FF35FBAE06DEF9E1" pageId="12" pageNumber="123">
- Pronotum matt, densely punctate; distance between two nearest pronotal punctures smaller than diameter of a puncture; elytra at least 1.81× as long as wide, with semi-appressed setae on disc, and with coarsely punctate striae; distance between elytral punctures shorter than their diameter; interstriae weakly wider or equally wide as striae; funicle segments in females longer than wide or as long as wide (except for
<taxonomicName id="71C6E3E5F5098A74FDC4FB05019FF9E1" ID-CoL="6ZX49" authorityName="Wagner" authorityYear="1912" box="[614,730,1237,1259]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="leonhardi">
<emphasis id="84B24474F5098A74FDC4FB05019FF9E1" box="[614,730,1237,1259]" italics="true" pageId="12" pageNumber="123">S. leonhardi</emphasis>
</taxonomicName>
)......................................................... 4
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<paragraph id="B6799866F5098A74FF35FB2306DFF849" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35FB2306DFF849" pageId="12" pageNumber="123">
4. Epifrons (Fig. 16) wide, with distinctly concave sides, posteriorly significantly enlarged, at base only slightly narrower than space between anterior margins of eyes; funicle segments
<quantity id="713E3583F5098A74FCAFFAC0000FF82C" box="[781,842,1296,1318]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="12" pageNumber="123" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">37 in</quantity>
females wider than long; pronotum in female wider than long....................................................................................
<taxonomicName id="71C6E3E5F5098A74FB75FAFD06DFF849" ID-CoL="6ZX49" authority="Wagner" authorityName="Wagner" authorityYear="1912" box="[1239,1434,1325,1347]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="leonhardi">
<emphasis id="84B24474F5098A74FB75FAFD060EF849" box="[1239,1355,1325,1347]" italics="true" pageId="12" pageNumber="123">S. leonhardi</emphasis>
Wagner
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- Epifrons (Fig. 19) slender, at basal half subparallel-sided, at base significantly narrower than space between anterior margins of eyes; funicle segments
<quantity id="713E3583F5098A74FE40FAB80164F874" box="[482,545,1384,1406]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="12" pageNumber="123" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">37 in</quantity>
females isodiametric or longer than wide; pronotum in female isodiametric or longer than wide................................................................................................ 5
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<paragraph id="B6799866F5098A74FF35FA7306D9FB1B" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35FA7306D9FB1B" pageId="12" pageNumber="123">
5. Femora ventraly with small but distinct tooth (
<figureCitation id="2EFD84E3F5098A74FD12FA7301A8F8B3" box="[688,749,1443,1465]" captionStart="FIGURES 12. 1" captionStartId="4.[151,264,1375,1398]" captionTargetBox="[155,1428,213,1354]" captionTargetId="figure@4.[151,1436,193,1354]" captionTargetPageId="4" captionText="FIGURES 12. 1—Stomodes benedikti, habitus; 2—Stomodes dodocunevi, habitus." httpUri="https://zenodo.org/record/1451247/files/figure.png" pageId="12" pageNumber="123">Fig. 2</figureCitation>
); rostrum with slightly concave sides and slightly laterally projecting pterygiae; striae and interstriae with equally large punctures; interstriae weakly wider than striae; female sternite VIII (Fig. 3) with isodiametric plate, narrowly sclerotised along anterior margin and with large fenestra; spermatheca (Fig. 4) with ramus and nodulus perpendicular.................................................................
<taxonomicName id="71C6E3E5F5098A74FB4CFA2B0625FB1B" ID-CoL="6ZWRZ" box="[1262,1376,1531,1553]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="benedikti" status="sp. nov.">
<emphasis id="84B24474F5098A74FB4CFA2B0625FB1B" box="[1262,1376,1531,1553]" italics="true" pageId="12" pageNumber="123">S. benedikti</emphasis>
</taxonomicName>
<emphasis id="84B24474F5098A74FAC4FA2B06D9FB1B" bold="true" box="[1382,1436,1531,1553]" pageId="12" pageNumber="123">
<taxonomicNameLabel id="9F81F90FF5098A74FAC4FA2B06D9FB1B" box="[1382,1436,1531,1553]" pageId="12" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="B6799866F5098A74FF35F9C906DEFB63" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35F9C906DEFB63" pageId="12" pageNumber="123">- Femora adentate (Fig. 8); rostrum with straight sides, without laterally projecting pterygiae; punctures in striae distinctly larger than those on interstriae; striae and interstriae equally wide; plate of female sternite VIII (Figs 10, 20) wider than long, with apical half uniformly sclerotized and lacking fenestra; spermatheca (Fig. 12) with ramus and nodulus divergent, V-shaped.. 6</keyLead>
</paragraph>
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<keyStep id="0D3280C3F5098A74FF35F9A006D9FA59" pageId="12" pageNumber="123">
<paragraph id="B6799866F5098A74FF35F9A006D9FBD5" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35F9A006D9FBD5" pageId="12" pageNumber="123">
6. Epifrons with longitudinal median stria; pronotum widest slightly before the middle; pronotal disc with well isolated punctures; elytra longer, 1.891.98× as long as wide; rostrum longer, 1.551.67× as wide as long; metatarsal segment 2 1.1× as wide as long; penis (Fig. 13) in dorsal view at apex evenly regularly tapered, tip shortly subtruncate; plate of female sternite VIII (Fig. 10) with apical margin concave...................................................
<taxonomicName id="71C6E3E5F5098A74FB77F918061BFBD4" ID-CoL="6ZWRY" box="[1237,1374,1736,1758]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="dodocunevi" status="sp. nov.">
<emphasis id="84B24474F5098A74FB77F918061BFBD4" box="[1237,1374,1736,1758]" italics="true" pageId="12" pageNumber="123">S. dodocunevi</emphasis>
</taxonomicName>
<emphasis id="84B24474F5098A74FAC4F91906D9FBD5" bold="true" box="[1382,1436,1737,1759]" pageId="12" pageNumber="123">
<taxonomicNameLabel id="9F81F90FF5098A74FAC4F91906D9FBD5" box="[1382,1436,1737,1759]" pageId="12" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="B6799866F5098A74FF35F93606D9FA59" blockId="12.[151,1436,797,1875]" pageId="12" pageNumber="123">
<keyLead id="0D373B53F5098A74FF35F93606D9FA59" pageId="12" pageNumber="123">
- Epifrons convex, lacking stria; pronotum widest at middle; pronotal disc with space between punctures very narrow; elytra shorter, 1.811.91× as long as wide; rostrum shorter, 1.781.91× as wide as long; metatarsal segment 2 1.4× as wide as long; penis in dorsal view (Fig. 21) with apex weakly enlarged and then tapered and pointed apicad; plate of female sternite VIII (Fig. 20) with round apical margin...........................................................
<taxonomicName id="71C6E3E5F5098A74FB51F8ED06D9FA59" ID-CoL="6ZWRW" authority="Reitter" authorityName="Reitter" authorityYear="1889" box="[1267,1436,1853,1875]" class="Insecta" family="Curculionidae" genus="Stomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="letzneri">
<emphasis id="84B24474F5098A74FB51F8ED0611FA59" box="[1267,1364,1853,1875]" italics="true" pageId="12" pageNumber="123">S. letzneri</emphasis>
Reitter
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