513 lines
55 KiB
XML
513 lines
55 KiB
XML
<document id="AAC4A6028F59CB617CE949FBC3F1BE16" ID-DOI="10.5281/zenodo.293973" ID-GBIF-Dataset="e22a2a00-99d4-4f31-aa1d-50a15aee1ee9" ID-ISSN="1175-5326" ID-Zenodo-Dep="293973" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227972717" checkinUser="plazi" docAuthor="M, José, Costas, Salgado, Moreda, Lucía Labrada & Luque, Carlos G." docDate="2010" docId="2F1887F79927793F16890DDA38B8F874" docLanguage="en" docName="zt02484p034.pdf" docOrigin="Zootaxa 2484" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Quaestus (Quaesticulus) pasensis Salgado, Labrada & Luque, new species" docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="D321FF8F99247938161E0D4D3979FF94" masterDocTitle="A new cave-dwelling species of Quaestus Schaufuss, 1861 from the Montes de Pas, Northern Spain (Coleoptera: Leiodidae: Leptodirini)" masterLastPageNumber="34" masterPageNumber="25" pageNumber="28" updateTime="1698238730522" updateUser="plazi">
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<mods:mods id="11CA3DA320E9E73749DDA4AD3A7E2C57" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5925B22C29340E583C1BD6E560EF4F55">
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<mods:title id="0EAD7403A17CAA8BD8996393D3659741">A new cave-dwelling species of Quaestus Schaufuss, 1861 from the Montes de Pas, Northern Spain (Coleoptera: Leiodidae: Leptodirini)</mods:title>
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</mods:titleInfo>
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<mods:name id="070A5071B6B9A748B7B3957374492012" type="personal">
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<mods:role id="38A21B2DB2984882CD66DD1DE8F1125C">
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<mods:roleTerm id="F6E43A7ADFE5765D0D01187626712435">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E91CC40E78E645F8FB40D46EDA57A71F">M, José</mods:namePart>
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</mods:name>
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<mods:name id="4BEA74D2556BD4B75589223BD59EEC70" type="personal">
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<mods:role id="15A13DD15964159E5246CD825F56B808">
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<mods:roleTerm id="874658E41831C2B5C3933D1ABBD6EF3F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8CF4D7F3BC1C3E8192C20C6A4486B069">Costas, Salgado</mods:namePart>
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</mods:name>
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<mods:name id="11C469D60499D408A06077842376855F" type="personal">
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<mods:role id="1497A02A19446DF111C8087C256771D2">
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<mods:roleTerm id="DC5B59AE02BD8FAE7A6E90A11BFA346F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="239D14FDD5D315CF1C645C94F91B3019">Moreda, Lucía Labrada</mods:namePart>
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</mods:name>
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<mods:name id="748DD7AF00A304D6B0D2BE6C23F018AA" type="personal">
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<mods:role id="37CC0636B4937840C65C7D5286D4AD13">
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<mods:roleTerm id="2994010C1B60959AC8A776557D257ECE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02209C04E09841BF318BBE5C16FA04E7">Luque, Carlos G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1F698CCA9C3438AF398FE1682000FF25">text</mods:typeOfResource>
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<mods:titleInfo id="6C9E0BF7AE95B2ED9E47E4F38316AAAD">
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<mods:title id="9343527FAF7C98C3B9731BC39E09FFD1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A007D71EF43089BA7654E8B6ABF41B6F">
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<mods:date id="96E810CA98DF1B283E72302A9BF14ACE">2010</mods:date>
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<mods:detail id="4B3E81FD5BE1AF63E5BFC3953E1178EC" type="volume">
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<mods:number id="56A0DC9873BC97437FED9892857ADD9A">2484</mods:number>
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</mods:detail>
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<mods:start id="BD0AE669EB02D22C9E81164994D93D6D">25</mods:start>
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<mods:end id="E6285811153C38216A6AA7AD76E3EB38">34</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="74BF083B7F6EFE03B1135C8ABF15294A">journal article</mods:classification>
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<mods:identifier id="2D08EFFFA277ADAAE516EBD2CBD8A87E" type="DOI">10.5281/zenodo.293973</mods:identifier>
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<mods:identifier id="A413180D5036D4C5C4303AE35A1DB6D2" type="GBIF-Dataset">e22a2a00-99d4-4f31-aa1d-50a15aee1ee9</mods:identifier>
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<mods:identifier id="20DCE4611B7F94D7840073EEC7F0E950" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6833292BA528497FBA085E95A78587B6" type="Zenodo-Dep">293973</mods:identifier>
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</mods:mods>
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<treatment id="2F1887F79927793F16890DDA38B8F874" ID-DOI="http://doi.org/10.5281/zenodo.6212106" ID-GBIF-Taxon="119386877" ID-Zenodo-Dep="6212106" LSID="urn:lsid:plazi:treatment:2F1887F79927793F16890DDA38B8F874" httpUri="http://treatment.plazi.org/id/2F1887F79927793F16890DDA38B8F874" lastPageId="7" lastPageNumber="32" pageId="3" pageNumber="28">
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<subSubSection id="EFAB656A9927793B16890DDA386EFF40" pageId="3" pageNumber="28" type="nomenclature">
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<paragraph id="A70E36E19927793B16890DDA3D53FF25" blockId="3.[151,1066,151,212]" box="[151,1066,151,177]" pageId="3" pageNumber="28">
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<heading id="FC46818D9927793B16890DDA3D53FF25" bold="true" box="[151,1066,151,177]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
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<emphasis id="95C5EAF39927793B16890DDA3D53FF25" bold="true" box="[151,1066,151,177]" pageId="3" pageNumber="28">
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<taxonomicName id="60B14D629927793B16890DDA3AF4FF25" ID-CoL="4QZLX" authority="Salgado, Labrada & Luque" authorityName="Salgado, Labrada & Luque" box="[151,909,151,177]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov." subGenus="Quaesticulus">
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||
<emphasis id="95C5EAF39927793B16890DDA387EFF25" bold="true" box="[151,263,151,177]" italics="true" pageId="3" pageNumber="28">Quaestus</emphasis>
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||
(
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||
<emphasis id="95C5EAF39927793B17060DDA38CFFF25" bold="true" box="[280,438,151,177]" italics="true" pageId="3" pageNumber="28">Quaesticulus</emphasis>
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||
)
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||
<emphasis id="95C5EAF39927793B17D90DDA3B55FF25" bold="true" box="[455,556,151,177]" italics="true" pageId="3" pageNumber="28">pasensis</emphasis>
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Salgado, Labrada & Luque
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</taxonomicName>
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,
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<taxonomicNameLabel id="8EF657889927793B15840DDA3D53FF25" box="[922,1066,151,177]" pageId="3" pageNumber="28" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="A70E36E19927793B16890DF7386EFF40" blockId="3.[151,1066,151,212]" box="[151,279,186,212]" pageId="3" pageNumber="28">
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||
(
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<figureCitation id="3F8A2A649927793B16810DF73995FF40" box="[159,236,186,212]" captionStart="FIGURE 1" captionStartId="4.[151,255,1368,1392]" captionText="FIGURE 1. Habitus of Quaestus pasensis sp. nov., holotype." pageId="3" pageNumber="28">Figs. 1</figureCitation>
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–7)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EFAB656A9927793B16890C483A19FD00" pageId="3" pageNumber="28" type="materials_examined">
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||
<paragraph id="A70E36E19927793B16890C483A19FD00" blockId="3.[151,1437,260,1117]" pageId="3" pageNumber="28">
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||
<emphasis id="95C5EAF39927793B16890C4838BBFE8B" bold="true" box="[151,450,261,287]" pageId="3" pageNumber="28">
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||
<typeStatus id="780A88439927793B16890C4839AAFE8B" box="[151,211,261,287]" pageId="3" pageNumber="28">Type</typeStatus>
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material.
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<typeStatus id="780A88439927793B174E0C4838BBFE8B" box="[336,450,261,287]" pageId="3" pageNumber="28" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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(3) and
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<emphasis id="95C5EAF39927793B14330C483BE9FE8B" bold="true" box="[557,656,261,287]" pageId="3" pageNumber="28">allotype</emphasis>
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(Ƥ):
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<emphasis id="95C5EAF39927793B14CC0C483AC0FE8B" bold="true" box="[722,953,261,287]" pageId="3" pageNumber="28">
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<collectingCountry id="DFA676719927793B14CC0C483A54FE8B" box="[722,813,261,287]" name="Spain" pageId="3" pageNumber="28">SPAIN</collectingCountry>
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: Cantabria
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</emphasis>
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: Luena: El Churrón Cave (The Churrón- Millajo Cave karst system, 30TVN
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<quantity id="60499B049927793B144C0C673A28FED0" box="[594,849,298,324]" metricMagnitude="12" metricUnit="m" metricValue="2.988980475255" pageId="3" pageNumber="28" unit="m" value="2.988980475255E12">2988980475, 255 m</quantity>
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||
),
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<date id="D30F10219927793B15760C673A86FED0" box="[872,1023,298,324]" pageId="3" pageNumber="28" value="2007-06-27">27-VI-2007</date>
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, Ocejo, leg. C.G.Luque (
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<collectionCode id="C1A0AE249927793B13260C673CE9FED0" box="[1336,1424,298,324]" pageId="3" pageNumber="28">CZUL</collectionCode>
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||
).
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||
<typeStatus id="780A88439927793B16890C02386DFEFD" box="[151,276,335,361]" pageId="3" pageNumber="28" type="paratype">
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||
<emphasis id="95C5EAF39927793B16890C02386DFEFD" bold="true" box="[151,276,335,361]" pageId="3" pageNumber="28">Paratypes</emphasis>
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||
</typeStatus>
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||
:
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<emphasis id="95C5EAF39927793B173D0C023B72FEFD" bold="true" box="[291,523,335,361]" pageId="3" pageNumber="28">
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||
<collectingCountry id="DFA676719927793B173D0C023804FEFD" box="[291,381,335,361]" name="Spain" pageId="3" pageNumber="28">SPAIN</collectingCountry>
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||
: Cantabria
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||
</emphasis>
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||
: Luena: 253, 30
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||
<collectionCode id="C1A0AE249927793B14F90C023A6AFEFD" box="[743,787,335,361]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
, same data as the
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||
<typeStatus id="780A88439927793B15F60C1D3D28FEFE" box="[1000,1105,336,362]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
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||
, (33, 3
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||
<collectionCode id="C1A0AE249927793B12A00C023D93FEFD" box="[1214,1258,335,361]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
<collectionCode id="C1A0AE249927793B12EF0C1D3C37FEFE" LSID="urn:lsid:biocol.org:col:34988" box="[1265,1358,336,362]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="28">MNHN</collectionCode>
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||
; 33, 3
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||
<collectionCode id="C1A0AE249927793B16BB0C3839A8FE1B" box="[165,209,373,399]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
<collectionCode id="C1A0AE249927793B16C40C383857FE1B" box="[218,302,373,399]" httpUri="http://grbio.org/cool/z0c5-kfkv" name="Museo Zoologia" pageId="3" pageNumber="28">MZBS</collectionCode>
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||
; 43, 4
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||
<collectionCode id="C1A0AE249927793B178A0C383B77FE1B" box="[404,526,373,399]" pageId="3" pageNumber="28">ƤƤ JFCL</collectionCode>
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||
; 43, 4
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||
<collectionCode id="C1A0AE249927793B146A0C383B8CFE1B" box="[628,757,373,399]" pageId="3" pageNumber="28">ƤƤ JMSC</collectionCode>
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||
; the rest
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||
<collectionCode id="C1A0AE249927793B157C0C383ACDFE1B" box="[866,948,373,399]" pageId="3" pageNumber="28">CZUL</collectionCode>
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||
); 83, 8
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||
<collectionCode id="C1A0AE249927793B123D0C383D36FE1B" box="[1059,1103,373,399]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
, El Churrón Cave,
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||
<date id="D30F10219927793B13260C3839ACFE20" pageId="3" pageNumber="28" value="2008-08-23">23-VIII- 2008</date>
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||
, leg. C.G.Luque & L.Labrada (
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||
<collectionCode id="C1A0AE249927793B14580CD73BE3FE20" box="[582,666,410,436]" pageId="3" pageNumber="28">CZUL</collectionCode>
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||
); 403, 38
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||
<collectionCode id="C1A0AE249927793B153A0CD73A29FE20" box="[804,848,410,436]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
, Rellano Cave (30TVN2897080570, altitude
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<quantity id="60499B049927793B136E0CD739CBFE4E" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="28" unit="m" value="270.0">270 m</quantity>
|
||
),
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||
<date id="D30F10219927793B16D60C8D3814FE4E" box="[200,365,448,474]" pageId="3" pageNumber="28" value="2008-08-23">23-VIII-2008</date>
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||
, Ocejo, leg. C.G.Luque & L.Labrada (33, 3
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<collectionCode id="C1A0AE249927793B15870CF23ABCFE4D" box="[921,965,447,473]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
, in
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<collectionCode id="C1A0AE249927793B15EB0C8D3D2BFE4E" LSID="urn:lsid:biocol.org:col:35275" box="[1013,1106,448,474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="28">MNCN</collectionCode>
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||
; 33, 3
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<collectionCode id="C1A0AE249927793B12A40CF23D9FFE4D" box="[1210,1254,447,473]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
<collectionCode id="C1A0AE249927793B12F10C8D3C35FE4E" box="[1263,1356,448,474]" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="3" pageNumber="28">MCNA</collectionCode>
|
||
; 43, 4
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||
<collectionCode id="C1A0AE249927793B16BB0CA839A8FE6B" box="[165,209,485,511]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
,
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||
<collectionCode id="C1A0AE249927793B16FE0CA8384EFE6B" box="[224,311,485,511]" pageId="3" pageNumber="28">PMGC</collectionCode>
|
||
; the rest in
|
||
<collectionCode id="C1A0AE249927793B17A10CA83B68FE6B" box="[447,529,485,511]" pageId="3" pageNumber="28">CZUL</collectionCode>
|
||
); 23, 3
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||
<collectionCode id="C1A0AE249927793B14630CA83BD0FE6B" box="[637,681,485,511]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
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||
, Picón de Riolango Cave (30TVN2983880286, altitude
|
||
<quantity id="60499B049927793B135E0CA83CF7FE6B" box="[1344,1422,485,511]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="3" pageNumber="28" unit="m" value="265.0">265 m</quantity>
|
||
),
|
||
<date id="D30F10219927793B16890F473857FDB0" box="[151,302,522,548]" pageId="3" pageNumber="28" value="2008-10-25">25-X- 2008</date>
|
||
, Riolango, leg. C.G.Luque & L.Labrada (
|
||
<collectionCode id="C1A0AE249927793B15910F473A9DFDB0" box="[911,996,522,548]" pageId="3" pageNumber="28">CZUL</collectionCode>
|
||
); 23, 4
|
||
<collectionCode id="C1A0AE249927793B126D0F473DDBFDB0" box="[1139,1186,522,548]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
|
||
, La Resaca Cave (30TVN3000181278, altitude
|
||
<quantity id="60499B049927793B17E20F7D3B33FDDE" box="[508,586,560,586]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="28" unit="m" value="320.0">320 m</quantity>
|
||
),
|
||
<date id="D30F10219927793B147E0F7D3BAEFDDE" box="[608,727,560,586]" pageId="3" pageNumber="28" value="2009-01-10">10-I-2009</date>
|
||
, Ocejo, leg. C.G.Luque & L.Labrada (
|
||
<collectionCode id="C1A0AE249927793B12830F7D3D89FDDE" box="[1181,1264,560,586]" pageId="3" pageNumber="28">CZUL</collectionCode>
|
||
); Vega de Pas: 23, 4
|
||
<collectionCode id="C1A0AE249927793B16F10F183862FDFB" box="[239,283,597,623]" pageId="3" pageNumber="28">ƤƤ</collectionCode>
|
||
, La Millajo Cave (The Churrón-Millajo Cave karst system, 30TVN3031081276, altitude
|
||
<quantity id="60499B049927793B135E0F183CF7FDFB" box="[1344,1422,597,623]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="28" unit="m" value="320.0">320 m</quantity>
|
||
),
|
||
<date id="D30F10219927793B16890F373876FD00" box="[151,271,634,660]" pageId="3" pageNumber="28" value="2009-01-10">10-I-2009</date>
|
||
, Guzparras, leg. C.G.Luque & L.Labrada (
|
||
<collectionCode id="C1A0AE249927793B151F0F373A2DFD00" box="[769,852,634,660]" pageId="3" pageNumber="28">CZUL</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFAB656A9927793B16D80FEF3DE6FCB8" pageId="3" pageNumber="28" type="diagnosis">
|
||
<paragraph id="A70E36E19927793B16D80FEF3DE6FCB8" blockId="3.[151,1437,260,1117]" pageId="3" pageNumber="28">
|
||
<emphasis id="95C5EAF39927793B16D80FEF383AFD28" bold="true" box="[198,323,674,700]" pageId="3" pageNumber="28">Diagnosis.</emphasis>
|
||
10th antennomere almost as long as 11th; all antennomeres slender and much longer than wide; male protarsus somewhat wider than apex of protibia; elytra bearing sutural striae; median lobe of aedeagus pointed, internal sac of aedeagus exhibiting small scales and a short, slender stylet, without basal plate; spermathecal complex with short and fine spermathecal duct, somewhat dilated anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFAB656A9927793E16D80E753D69FF26" lastPageId="6" lastPageNumber="31" pageId="3" pageNumber="28" type="description">
|
||
<paragraph id="A70E36E19927793B16D80E7538B7FC08" blockId="3.[151,1437,260,1117]" pageId="3" pageNumber="28">
|
||
<emphasis id="95C5EAF39927793B16D80E753827FCC6" bold="true" box="[198,350,824,850]" pageId="3" pageNumber="28">Description.</emphasis>
|
||
<emphasis id="95C5EAF39927793B17760E753B5CFCC5" box="[360,549,824,849]" italics="true" pageId="3" pageNumber="28">
|
||
<typeStatus id="780A88439927793B17760E7538AFFCC5" box="[360,470,824,849]" pageId="3" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
, male.
|
||
</emphasis>
|
||
Body length
|
||
<quantity id="60499B049927793B14D50E753A40FCC6" box="[715,825,824,850]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="3" pageNumber="28" unit="mm" value="2.44">2.44 mm</quantity>
|
||
; maximum width
|
||
<quantity id="60499B049927793B120A0E753DFBFCC6" box="[1044,1154,824,850]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="28" unit="mm" value="1.3">1.30 mm</quantity>
|
||
. Uniformly ovoid body tapering posteriad; pronotum and elytra convex, completely covered with short, fine, sparse and recumbent yellowish pilosity (
|
||
<figureCitation id="3F8A2A649927793B17660ECF38C4FC08" box="[376,445,898,924]" captionStart="FIGURE 1" captionStartId="4.[151,255,1368,1392]" captionText="FIGURE 1. Habitus of Quaestus pasensis sp. nov., holotype." pageId="3" pageNumber="28">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19927793B16D80EE5387CFBC9" blockId="3.[151,1437,260,1117]" pageId="3" pageNumber="28">Punctures on head extremely fine and superficial, seem almost non-existent giving tegument very shiny reddish brown appearance. Antennae long, surpassing half of body length; antennomeres long and cylindrical, 1st, 2nd, 7th, 9th and 10th are slightly dilated apically, all at least twice as long as wide; 11th antennomere the longest, slightly longer than 10th, 10th is slightly shorter than 9th; 4th almost as long as 3rd; 3rd, 5th and 6th are equally long (Table 1).</paragraph>
|
||
<paragraph id="A70E36E19927793B168909C53BC8FB56" blockId="3.[151,1436,1160,1352]" pageId="3" pageNumber="28">
|
||
<emphasis id="95C5EAF39927793B168909C53870FB34" bold="true" box="[151,265,1160,1184]" pageId="3" pageNumber="28">TABLE 1.</emphasis>
|
||
Measurements of the
|
||
<typeStatus id="780A88439927793B17E409C53B2EFB0B" box="[506,599,1160,1183]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
(male) of
|
||
<taxonomicName id="60B14D629927793B14D709C43AF4FB34" box="[713,909,1161,1184]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39927793B14D709C43AF4FB34" box="[713,909,1161,1184]" italics="true" pageId="3" pageNumber="28">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C5EAF39927793B158B09C53A92FB34" bold="true" box="[917,1003,1160,1184]" pageId="3" pageNumber="28">
|
||
<taxonomicNameLabel id="8EF657889927793B158B09C53A92FB34" box="[917,1003,1160,1184]" pageId="3" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Length (L) and maximum width (W) of antennomeres 1st to 11th (all measurements in mm).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19927793B171609933C05FADC" blockId="3.[151,1436,1160,1352]" pageId="3" pageNumber="28">
|
||
1st 2nd 3rd 4th 5th 6th 7th 8th 9th 10th 11th L 0.145 0.162 0.107 0.100 0.107 0.107 0.133 0.091 0.140 0.136
|
||
<geoCoordinate id="C28550269927793B1325084539C7FADC" direction="west" orientation="longitude" pageId="3" pageNumber="28" precision="55" value="-0.167">
|
||
0.167
|
||
<geoCoordinate id="C28550269927793B16BB087C39C7FADC" box="[165,190,1329,1352]" direction="west" orientation="longitude" pageId="3" pageNumber="28" precision="55" value="-0.051">W</geoCoordinate>
|
||
</geoCoordinate>
|
||
0.051 0.047 0.036 0.036 0.036 0.036 0.047 0.036 0.047 0.047 0.056
|
||
</paragraph>
|
||
<paragraph id="A70E36E19927793B16D808333C25F9B9" blockId="3.[151,1437,1406,2032]" pageId="3" pageNumber="28">
|
||
Pronotum strongly transverse, widest at base, almost twice as wide as long, as wide as anterior part of elytra; lateral margins uniformly arcuate, posterior corners blunt and slightly prolonged posteriorly, angle slightly obtuse; tegument with very fine superficial punctation. Mesoventral carina as in group 2C (
|
||
<bibRefCitation id="C3204B109927793B13250884380AF99C" author="Salgado" pageId="3" pageNumber="33" refString="Salgado, J. M. (1996) Estudio sobre la importancia de la quilla mesosternal. Descripcion de Leonesiella n. gen. y " status " del gen. Speogeus (Col. Cholevidae, Leptodirinae). Memoires de Biospeologie, 23, 149 - 155." type="journal article" year="1996">Salgado 1996: see Figures</bibRefCitation>
|
||
), high, laminar and long, reaching metaventrite; anterior edge of carina weakly curved, slightly prominent, ventral edge straight and narrow; the strongly rounded vertex forms an obtuse angle.
|
||
</paragraph>
|
||
<paragraph id="A70E36E19927793B16D80B743D86F9EC" blockId="3.[151,1437,1406,2032]" pageId="3" pageNumber="28">Elytra elongated, 1.44 times as long as wide, widest near basal ¼, somewhat wider than pronotum (1.1 ratio); lateral margins uniformly curved to strongly rounded apex. Sutural striae well developed.</paragraph>
|
||
<paragraph id="A70E36E19927793B16D80BCB3827F97F" blockId="3.[151,1437,1406,2032]" pageId="3" pageNumber="28">Legs long and slender. Foretarsus with 1st–4th tarsomeres dilated, 1st tarsomere 1.13 times as wide as apex of tibia; protarsus elongate, one and a half times longer than wide; mesotibia somewhat arcuate and metatibia slightly undulate.</paragraph>
|
||
<paragraph id="A70E36E19927793B16D80BBB3BB9F864" blockId="3.[151,1437,1406,2032]" pageId="3" pageNumber="28">
|
||
Aedeagus (Figs. 2, 3)
|
||
<quantity id="60499B049927793B17D60BBB3B49F884" box="[456,560,1782,1808]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="28" unit="mm" value="0.49">0.49 mm</quantity>
|
||
long; in lateral view (Fig. 3) widely arcuate, forming obtuse angle near the middle; in dorsal view (Fig. 2) median lobe with uniformly arcuate lateral margins and weakly pointed apex. Ventral lamina of tegmen wide and rounded, well sclerotized, shorter than basal lamina. Parameres slender, almost reaching apex of median lobe, apical club weakly dilated and subrectangular, with three fairly long setae, two apically with insertion pores very close to each other and one basally, clearly distant from the other two (Fig. 4). Internal sac with short slender stylet lacking basal plate and surrounded by small scales; in apical half with narrow poorly visible reinforcing bands (Fig. 5).
|
||
</paragraph>
|
||
<caption id="F3CE66699920793C168908153A51FAE4" box="[151,808,1368,1392]" pageId="4" pageNumber="29">
|
||
<paragraph id="A70E36E19920793C168908153A51FAE4" blockId="4.[151,808,1368,1392]" box="[151,808,1368,1392]" pageId="4" pageNumber="29">
|
||
<emphasis id="95C5EAF39920793C168908153860FAE4" bold="true" box="[151,281,1368,1392]" pageId="4" pageNumber="29">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="60B14D629920793C178808143B23FAE4" box="[406,602,1369,1392]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39920793C178808143B23FAE4" box="[406,602,1369,1392]" italics="true" pageId="4" pageNumber="29">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8EF657889920793C147F08153BCFFAE4" box="[609,694,1368,1392]" pageId="4" pageNumber="29" rank="species">
|
||
<emphasis id="95C5EAF39920793C147F08153BCFFAE4" bold="true" box="[609,694,1368,1392]" pageId="4" pageNumber="29">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A70E36E19920793C16D808EC3A9EFA74" blockId="4.[151,1436,1438,1504]" pageId="4" pageNumber="29">
|
||
<emphasis id="95C5EAF39920793C16D808EC38F7FA2E" box="[198,398,1441,1466]" italics="true" pageId="4" pageNumber="29">Allotype, female.</emphasis>
|
||
Body length
|
||
<quantity id="60499B049920793C142508EC3BD5FA2F" box="[571,684,1441,1467]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" pageId="4" pageNumber="29" unit="mm" value="2.28">2.28 mm</quantity>
|
||
. Similar to male, except 1st–4th protarsomeres not dilated and antennae slightly shorter, with antennomeres wider than in male (Table 2).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19920793C16890B463BCCF9D1" blockId="4.[151,1436,1547,1741]" pageId="4" pageNumber="29">
|
||
<emphasis id="95C5EAF39920793C16890B463871F9B7" bold="true" box="[151,264,1547,1571]" pageId="4" pageNumber="29">TABLE 2.</emphasis>
|
||
Measurements of the allotype (female) of
|
||
<taxonomicName id="60B14D629920793C14CE0B413AECF9B7" box="[720,917,1548,1571]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39920793C14CE0B413AECF9B7" box="[720,917,1548,1571]" italics="true" pageId="4" pageNumber="29">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8EF657889920793C15850B463A89F9B7" box="[923,1008,1547,1571]" pageId="4" pageNumber="29" rank="species">
|
||
<emphasis id="95C5EAF39920793C15850B463A89F9B7" bold="true" box="[923,1008,1547,1571]" pageId="4" pageNumber="29">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Length (L) and maximum width (W) of antennomeres 1st to 11th (all measurements in mm).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19920793C17160B2C3C3EF8A5" blockId="4.[151,1436,1547,1741]" lastBlockId="4.[151,1437,1811,2027]" pageId="4" pageNumber="29">
|
||
1st 2nd 3rd 4th 5th 6th 7th 8th 9th 10th 11th L 0.138 0.147 0.100 0.096 0.100 0.100 0.113 0.076 0.113 0.109
|
||
<geoCoordinate id="C28550269920793C13220BC139C1F959" direction="west" orientation="longitude" pageId="4" pageNumber="29" precision="55" value="-0.151">
|
||
0.151
|
||
<geoCoordinate id="C28550269920793C16810BFB39C1F959" box="[159,184,1718,1741]" direction="west" orientation="longitude" pageId="4" pageNumber="29" precision="55" value="-0.047">W</geoCoordinate>
|
||
</geoCoordinate>
|
||
0.047 0.040 0.033 0.033 0.033 0.033 0.040 0.033 0.040 0.040 0.051 Spiculum ventrale of 8th urosternite short and fine, sides almost parallel and apex rounded (Fig. 6).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19920793C16D80A713DDFF87F" blockId="4.[151,1437,1811,2027]" pageId="4" pageNumber="29">
|
||
Spermathecal complex distinct,
|
||
<quantity id="60499B049920793C14210A713BD1F8C2" box="[575,680,1852,1878]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="29" unit="mm" value="0.14">0.14 mm</quantity>
|
||
long, bilobed, basal lobe ovoid, longer and wider than spherical apical lobe, both lobes well-sclerotized; the part joining lobes membranous, as long as apical lobe; spermathecal duct short and thin, 3-4 times as long as spermatheca, widened at beginning, posterior part directly joined to dorsal face of bursa copulatrix; spermathecal gland small, rounded and membranous usually not easily discernible, opens into widened anterior part of the spermathecal duct (Fig. 7).
|
||
</paragraph>
|
||
<paragraph id="A70E36E19921793D16890A1638F4F821" blockId="5.[151,1437,1883,1973]" pageId="5" pageNumber="30">
|
||
<emphasis id="95C5EAF39921793D16890A16383DF8E7" bold="true" box="[151,324,1883,1907]" pageId="5" pageNumber="30">FIGURES 2–7.</emphasis>
|
||
Details of
|
||
<taxonomicName id="60B14D629921793D17A50A113BF8F8E7" box="[443,641,1884,1907]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39921793D17A50A113BF8F8E7" box="[443,641,1884,1907]" italics="true" pageId="5" pageNumber="30">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8EF657889921793D14960A163BA7F8E7" box="[648,734,1883,1907]" pageId="5" pageNumber="30" rank="species">
|
||
<emphasis id="95C5EAF39921793D14960A163BA7F8E7" bold="true" box="[648,734,1883,1907]" pageId="5" pageNumber="30">
|
||
sp.
|
||
<date id="D30F10219921793D14B10A163BA7F8E7" box="[687,734,1883,1907]" pageId="5" pageNumber="30">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
2: aedeagus in dorsal view; 3: aedeagus in lateral view; 4: apical club of the right paramere; 5: sclerotized structures of internal sac; 6: ventral spine of 8th female urosternite; 7: spermathecal complex.
|
||
</paragraph>
|
||
<paragraph id="A70E36E19922793E16D80DDA3D69FF26" blockId="6.[151,1437,151,554]" box="[198,1040,151,178]" pageId="6" pageNumber="31">
|
||
<emphasis id="95C5EAF39922793E16D80DDA382BFF25" bold="true" box="[198,338,151,177]" pageId="6" pageNumber="31">Variability.</emphasis>
|
||
Body length of the
|
||
<typeStatus id="780A88439922793E14250DD53BD2FF26" box="[571,683,152,178]" pageId="6" pageNumber="31" type="paratype">paratypes</typeStatus>
|
||
ranges from
|
||
<quantity id="60499B049922793E15590DD53D72FF26" box="[839,1035,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.5" metricValueMin="2.25" pageId="6" pageNumber="31" unit="mm" value="2.375" valueMax="2.5" valueMin="2.25">2.25 to 2.50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFAB656A9922793E16D80DF038EDFED3" pageId="6" pageNumber="31" type="discussion">
|
||
<paragraph id="A70E36E19922793E16D80DF038EDFED3" blockId="6.[151,1437,151,554]" pageId="6" pageNumber="31">
|
||
<emphasis id="95C5EAF39922793E16D80DF0382BFF43" bold="true" box="[198,338,189,215]" pageId="6" pageNumber="31">Discussion.</emphasis>
|
||
<taxonomicName id="60B14D629922793E17400DF03B42FF42" box="[350,571,189,214]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39922793E17400DF03B42FF42" box="[350,571,189,214]" italics="true" pageId="6" pageNumber="31">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the subgenus
|
||
<taxonomicName id="60B14D629922793E15610DF03D94FF43" authority="Schaufuss, 1861" authorityName="Schaufuss" authorityYear="1861" box="[895,1261,189,215]" class="Insecta" family="Leiodidae" genus="Quaesticulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="95C5EAF39922793E15610DF03D61FF42" box="[895,1048,189,214]" italics="true" pageId="6" pageNumber="31">Quaesticulus</emphasis>
|
||
<bibRefCitation id="C3204B109922793E123A0DF03D94FF43" author="Schaufuss" box="[1060,1261,189,215]" pageId="6" pageNumber="33" refString="Schaufuss, L. W. (1861) Zwei neue Silphiden Gattungen. Stettiner entomologische Zeitung, 22, 423 - 428." type="journal article" year="1861">Schaufuss, 1861</bibRefCitation>
|
||
</taxonomicName>
|
||
, based on the following characters: elytra bearing sutural stria, internal sac without developed sclerotized formations and with stylet lacking basal plate, spermathecal duct short with spermathecal gland small and membranous (
|
||
<bibRefCitation id="C3204B109922793E16810C6038FEFED3" author="Salgado" box="[159,391,301,327]" pageId="6" pageNumber="33" refString="Salgado, J. M., Blas, M. & Fresneda, J. (2008) Fauna Iberica. Vol. 31. Coleoptera, Cholevidae. Museo Nacional de Ciencias Naturales, Consejo Superior de Investigaciones Cientificas, Madrid, 802 pp." type="book" year="2008">
|
||
Salgado
|
||
<emphasis id="95C5EAF39922793E17180C603838FED2" box="[262,321,301,326]" italics="true" pageId="6" pageNumber="31">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFAB656A9922793F16D80C1F3D0EFF68" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31" type="reference_group">
|
||
<paragraph id="A70E36E19922793E16D80C1F3C50FDBE" blockId="6.[151,1437,151,554]" pageId="6" pageNumber="31">
|
||
Within
|
||
<taxonomicName id="60B14D629922793E17380C1F38EAFEFF" ID-CoL="8X7SL" box="[294,403,338,363]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="95C5EAF39922793E17380C1F38EAFEFF" box="[294,403,338,363]" italics="true" pageId="6" pageNumber="31">Quaestus</emphasis>
|
||
</taxonomicName>
|
||
the new species belongs to section IV (originally established by
|
||
<bibRefCitation id="C3204B109922793E12FB0C1F3CE1FEF8" author="Jeannel" box="[1253,1432,338,364]" pageId="6" pageNumber="33" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel(1924))</bibRefCitation>
|
||
, characterized by two characters: 11th antennomere slightly longer than 10th, and all antennomeres clearly longer than wide. Section IV includes two additional species:
|
||
<taxonomicName id="60B14D629922793E158B0CED3DB3FE2E" ID-CoL="79285" authority="Jeannel, 1909" authorityName="Jeannel" authorityYear="1909" box="[917,1226,416,442]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="minos">
|
||
<emphasis id="95C5EAF39922793E158B0CED3D7AFE2D" box="[917,1027,416,441]" italics="true" pageId="6" pageNumber="31">Q. minos</emphasis>
|
||
(
|
||
<bibRefCitation id="C3204B109922793E120B0CED3DB8FE2E" author="Jeannel" box="[1045,1217,416,442]" pageId="6" pageNumber="33" refString="Jeannel, R. (1909) Contribution a l'etude des Silphides cavernicoles d'Espagne. Boletin de la Real Sociedad espanola de Historia natural, 9 (19), 462 - 474." type="journal article" year="1909">Jeannel, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60B14D629922793E13190CED382DFE4B" ID-CoL="4QZLT" authority="Uhagon, 1881" authorityName="Uhagon" authorityYear="1881" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="filicornis">
|
||
<emphasis id="95C5EAF39922793E13190CED3CE5FE2D" box="[1287,1436,416,441]" italics="true" pageId="6" pageNumber="31">Q. filicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="C3204B109922793E16810C883832FE4B" author="Uhagon" box="[159,331,453,479]" pageId="6" pageNumber="33" refString="Uhagon, S. (1881) Especies nuevas del genero Bathyscia encontradas en Vizcaya. Anales de la Sociedad espanola de Historia natural, 10, 113 - 126." type="journal article" year="1881">Uhagón, 1881</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The new species differs from them in the shape of antennae, protarsi, mesoventral carina and the structure of the aedeagus; in
|
||
<taxonomicName id="60B14D629922793E142D0CA63BDDFD90" ID-CoL="79285" box="[563,676,491,516]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="minos">
|
||
<emphasis id="95C5EAF39922793E142D0CA63BDDFD90" box="[563,676,491,516]" italics="true" pageId="6" pageNumber="31">Q. minos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60B14D629922793E14FC0CA63A02FD90" ID-CoL="4QZLT" box="[738,891,491,516]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="filicornis">
|
||
<emphasis id="95C5EAF39922793E14FC0CA63A02FD90" box="[738,891,491,516]" italics="true" pageId="6" pageNumber="31">Q. filicornis</emphasis>
|
||
</taxonomicName>
|
||
the antennae are longer, protarsi are wider, mesoventral carina is stronger, the parameres and stylet of the aedeagus are longer and more robust.
|
||
</paragraph>
|
||
<caption id="F3CE66699922793E16890BAC3C7DF8AC" httpUri="https://zenodo.org/record/293974/files/figure.png" pageId="6" pageNumber="31" targetBox="[393,1193,608,1740]" targetPageId="6">
|
||
<paragraph id="A70E36E19922793E16890BAC3C7DF8AC" blockId="6.[151,1436,1761,1848]" pageId="6" pageNumber="31">
|
||
<emphasis id="95C5EAF39922793E16890BAC3859F96D" bold="true" box="[151,288,1761,1785]" pageId="6" pageNumber="31">FIGURES 8</emphasis>
|
||
. Location of the caves inhabited by
|
||
<taxonomicName id="60B14D629922793E14BE0BAF3AF8F96D" ID-CoL="4QZLP" box="[672,897,1762,1785]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39922793E14BE0BAF3AF8F96D" box="[672,897,1762,1785]" italics="true" pageId="6" pageNumber="31">Quaestus autumnalis</emphasis>
|
||
</taxonomicName>
|
||
(1–29),
|
||
<taxonomicName id="60B14D629922793E15C40BAF3D2DF96D" ID-CoL="4QZLX" box="[986,1108,1762,1785]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39922793E15C40BAF3D2DF96D" box="[986,1108,1762,1785]" italics="true" pageId="6" pageNumber="31">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8EF657889922793E12450BAC3DD6F96D" box="[1115,1199,1761,1785]" pageId="6" pageNumber="31" rank="species">
|
||
<emphasis id="95C5EAF39922793E12450BAC3DD6F96D" bold="true" box="[1115,1199,1761,1785]" pageId="6" pageNumber="31">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(30–34) and
|
||
<taxonomicName id="60B14D629922793E13220BAF3CE5F96D" ID-CoL="79285" box="[1340,1436,1762,1785]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="minos">
|
||
<emphasis id="95C5EAF39922793E13220BAF3CE5F96D" box="[1340,1436,1762,1785]" italics="true" pageId="6" pageNumber="31">Q. minos</emphasis>
|
||
</taxonomicName>
|
||
(35–42). (A) Geographical map of the area; (B) Distribution of the caves. Grey areas indicate rock outcrops containing limestone. Scale 1:500.000. Squares 10 x 10 km. For cave names, coordinates and altitudes see Appendix.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A70E36E19922793F16D80A2F3D0EFF68" blockId="6.[151,1436,1890,2033]" lastBlockId="7.[151,1436,152,252]" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
|
||
Externally, the new species is most similar to
|
||
<taxonomicName id="60B14D629922793E14EB0A2F3D17F8E8" ID-CoL="4QZLP" authority="Escalera, 1998" authorityName="Escalera" authorityYear="1998" box="[757,1134,1890,1916]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39922793E14EB0A2F3AE7F8EF" box="[757,926,1890,1915]" italics="true" pageId="6" pageNumber="31">Q. autumnalis</emphasis>
|
||
(Escalera, 1998)
|
||
</taxonomicName>
|
||
, a member of section III (
|
||
<bibRefCitation id="C3204B109922793E16810ACA383FF835" author="Jeannel" box="[159,326,1927,1953]" pageId="6" pageNumber="33" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel, 1924</bibRefCitation>
|
||
).
|
||
<taxonomicName id="60B14D629922793E17470ACA3B49F834" ID-CoL="4QZLX" box="[345,560,1927,1952]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39922793E17470ACA3B49F834" box="[345,560,1927,1952]" italics="true" pageId="6" pageNumber="31">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="60B14D629922793E15210ACA3A9CF834" ID-CoL="4QZLP" box="[831,997,1927,1952]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39922793E15210ACA3A9CF834" box="[831,997,1927,1952]" italics="true" pageId="6" pageNumber="31">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
based on the following characters: 1) in
|
||
<taxonomicName id="60B14D629922793E16A80AE23825F85C" ID-CoL="4QZLP" box="[182,348,1967,1992]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39922793E16A80AE23825F85C" box="[182,348,1967,1992]" italics="true" pageId="6" pageNumber="31">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
all antennomeres robust (slender in
|
||
<taxonomicName id="60B14D629922793E15120AE23AEDF85C" ID-CoL="4QZLX" box="[780,916,1967,1992]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39922793E15120AE23AEDF85C" box="[780,916,1967,1992]" italics="true" pageId="6" pageNumber="31">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
) and 11th antennomere almost twice as long as 10th (subequal in
|
||
<taxonomicName id="60B14D629922793E179A0A953B74F865" ID-CoL="4QZLX" box="[388,525,2008,2033]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39922793E179A0A953B74F865" box="[388,525,2008,2033]" italics="true" pageId="6" pageNumber="31">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
); 2) in
|
||
<taxonomicName id="60B14D629922793E14790A953A76F865" ID-CoL="4QZLP" box="[615,783,2008,2033]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39922793E14790A953A76F865" box="[615,783,2008,2033]" italics="true" pageId="6" pageNumber="31">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
ventral edge of mesoventral carina medially expanded (narrow throughout in
|
||
<taxonomicName id="60B14D629923793F17B00DD53B45FF25" ID-CoL="4QZLX" box="[430,572,152,177]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F17B00DD53B45FF25" box="[430,572,152,177]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
); 3) in
|
||
<taxonomicName id="60B14D629923793F14840DD53A3EFF25" ID-CoL="4QZLP" box="[666,839,152,177]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39923793F14840DD53A3EFF25" box="[666,839,152,177]" italics="true" pageId="7" pageNumber="32">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
protarsus always narrower than apex of protibia (clearly wider in
|
||
<taxonomicName id="60B14D629923793F177F0DF03890FF42" ID-CoL="4QZLX" box="[353,489,189,214]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F177F0DF03890FF42" box="[353,489,189,214]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
); 4)
|
||
<taxonomicName id="60B14D629923793F143D0DF03BB0FF42" ID-CoL="4QZLP" box="[547,713,189,214]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39923793F143D0DF03BB0FF42" box="[547,713,189,214]" italics="true" pageId="7" pageNumber="32">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
apex of median lobe blunt (pointed in
|
||
<taxonomicName id="60B14D629923793F12880DF03C64FF42" ID-CoL="4QZLX" box="[1174,1309,189,214]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F12880DF03C64FF42" box="[1174,1309,189,214]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
) and setae of parameres very long and undulate (much shorter and not undulate in
|
||
<taxonomicName id="60B14D629923793F15C00DAF3D1CFF6F" ID-CoL="4QZLX" box="[990,1125,226,251]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F15C00DAF3D1CFF6F" box="[990,1125,226,251]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F3CE66699923793F168909653ACDFBCB" httpUri="https://zenodo.org/record/293975/files/figure.png" pageId="7" pageNumber="32" targetBox="[188,1398,309,1040]" targetPageId="7">
|
||
<paragraph id="A70E36E19923793F168909653ACDFBCB" blockId="7.[151,1436,1064,1119]" pageId="7" pageNumber="32">
|
||
<emphasis id="95C5EAF39923793F16890965386CFBD4" bold="true" box="[151,277,1064,1088]" pageId="7" pageNumber="32">FIGURE 9</emphasis>
|
||
. Detailed map of study area indicating location of the five caves inhabited by
|
||
<taxonomicName id="60B14D629923793F126A09643C44FBD4" box="[1140,1341,1065,1088]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis" status="sp. nov.">
|
||
<emphasis id="95C5EAF39923793F126A09643C44FBD4" box="[1140,1341,1065,1088]" italics="true" pageId="7" pageNumber="32">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C5EAF39923793F135A09653CE5FBD4" bold="true" box="[1348,1436,1064,1088]" pageId="7" pageNumber="32">
|
||
<taxonomicNameLabel id="8EF657889923793F135A09653CE5FBD4" box="[1348,1436,1064,1088]" pageId="7" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(courtesy of the “Niphargus” Speleological Club). UTM grid 250 x 250 m.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EFAB656A9923793F16D809DC3D3AF801" pageId="7" pageNumber="32" type="description">
|
||
<paragraph id="A70E36E19923793F16D809DC3D4FFAF1" blockId="7.[151,1437,1166,2016]" pageId="7" pageNumber="32">
|
||
<taxonomicName id="60B14D629923793F16D809DC38E6FB3E" box="[198,415,1169,1194]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F16D809DC38E6FB3E" box="[198,415,1169,1194]" italics="true" pageId="7" pageNumber="32">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
is restricted to a small karstic area of some
|
||
<quantity id="60499B049923793F15B509DC3A91FB3F" box="[939,1000,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="32" unit="km" value="1.0">1 km</quantity>
|
||
2, and clearly isolated geographically from its most proximate species,
|
||
<taxonomicName id="60B14D629923793F143A09FB3BB2FB5B" box="[548,715,1206,1231]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="95C5EAF39923793F143A09FB3BB2FB5B" box="[548,715,1206,1231]" italics="true" pageId="7" pageNumber="32">Q. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
towards the north-west and
|
||
<taxonomicName id="60B14D629923793F123E09FB3DF2FB5B" box="[1056,1163,1206,1231]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="minos">
|
||
<emphasis id="95C5EAF39923793F123E09FB3DF2FB5B" box="[1056,1163,1206,1231]" italics="true" pageId="7" pageNumber="32">Q. minos</emphasis>
|
||
</taxonomicName>
|
||
towards the north-east by a strip of overthrusts.
|
||
<figureCitation id="3F8A2A649923793F17A009963883FB61" box="[446,506,1243,1269]" captionStart="FIGURES 8" captionStartId="6.[151,269,1761,1785]" captionTargetBox="[393,1193,608,1740]" captionTargetId="figure@6.[393,1194,608,1740]" captionTargetPageId="6" captionText="FIGURES 8. Location of the caves inhabited by Quaestus autumnalis (1 – 29), Q. pasensis sp. nov. (30 – 34) and Q. minos (35 – 42). (A) Geographical map of the area; (B) Distribution of the caves. Grey areas indicate rock outcrops containing limestone. Scale 1: 500.000. Squares 10 x 10 km. For cave names, coordinates and altitudes see Appendix." httpUri="https://zenodo.org/record/293974/files/figure.png" pageId="7" pageNumber="32">Fig 8</figureCitation>
|
||
B indicates the locations of the caves inhabited by the three species and clearly demonstrates gaps between the three species areas. The distribution area of
|
||
<taxonomicName id="60B14D629923793F1244084C3D95FA8E" box="[1114,1260,1281,1306]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F1244084C3D95FA8E" box="[1114,1260,1281,1306]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
appears to be restricted to the karst-fissured aquifer systems in the Upper Jurassic carbonate rocks within the Ocejo karst system that stretches through the municipal districts of Luena and Vega de Pas.
|
||
</paragraph>
|
||
<paragraph id="A70E36E19923793F16D8083C3827F9D1" blockId="7.[151,1437,1166,2016]" pageId="7" pageNumber="32">
|
||
Sampling was done in five caves: El Churrón, Relleno, Picón de Riolango, La Resaca and La Millajo (
|
||
<figureCitation id="3F8A2A649923793F1372083C39D1FA24" captionStart="FIGURE 9" captionStartId="7.[151,256,1064,1088]" captionTargetBox="[188,1398,309,1040]" captionTargetId="figure@7.[188,1398,309,1041]" captionTargetPageId="7" captionText="FIGURE 9. Detailed map of study area indicating location of the five caves inhabited by Quaestus pasensis sp. nov. (courtesy of the “ Niphargus ” Speleological Club). UTM grid 250 x 250 m." httpUri="https://zenodo.org/record/293975/files/figure.png" pageId="7" pageNumber="32">Fig. 9</figureCitation>
|
||
). El Churrón is about
|
||
<quantity id="60499B049923793F17AE08DB3B7AFA24" box="[432,515,1430,1456]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="32" unit="km" value="2.5">2.5 km</quantity>
|
||
long and it is the most important cave explored in the western part of the Pas Mountains. The cave is situated on a fault inscribed on a tectonical alignment controlled by a system of fractures in a general N 50–70º W and N 160–170º E direction. The other four caves are shorter, the longest of them being the Picón de Riolango Cave located at a roadside and consisting of a network of short galleries about
|
||
<quantity id="60499B049923793F16C10B663862F9D1" box="[223,283,1579,1605]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="32" unit="m" value="60.0">60 m</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="A70E36E19923793F16D80B1C3A2CF894" blockId="7.[151,1437,1166,2016]" pageId="7" pageNumber="32">
|
||
In all caves where
|
||
<taxonomicName id="60B14D629923793F17B60B1C3B48F9FE" box="[424,561,1617,1642]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F17B60B1C3B48F9FE" box="[424,561,1617,1642]" italics="true" pageId="7" pageNumber="32">Q. pasensis</emphasis>
|
||
</taxonomicName>
|
||
has been recorded it coexists with
|
||
<taxonomicName id="60B14D629923793F15C90B1C3CFBF9FF" authority="Jeannel, 1924" authorityName="Jeannel" authorityYear="1924" box="[983,1410,1617,1643]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="sharpi" subSpecies="nigricans">
|
||
<emphasis id="95C5EAF39923793F15C90B1C3DB9F9FE" box="[983,1216,1617,1642]" italics="true" pageId="7" pageNumber="32">Q. sharpi nigricans</emphasis>
|
||
(
|
||
<bibRefCitation id="C3204B109923793F12CE0B1C3C01F9FF" author="Jeannel" box="[1232,1400,1617,1643]" pageId="7" pageNumber="33" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63 (1), 1 - 436." type="journal article" year="1924">Jeannel, 1924</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, a subspecies widely distributed in the north eastern karstic area of Cantabria province and reported from almost one hundred caves (
|
||
<bibRefCitation id="C3204B109923793F179B0BD63B4AF921" author="Salgado" box="[389,563,1691,1717]" pageId="7" pageNumber="33" refString="Salgado, J. M. (1976) Origens e distribuicao geografica dos Bathysciinae (Col. Catopidae) cantabricos. Ciencia Biologica (Portugal), 1, 105 - 130." type="journal article" year="1976">Salgado, 1976</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="C3204B109923793F145E0BD63BF9F921" author="Salgado" box="[576,640,1691,1717]" pageId="7" pageNumber="33" refString="Salgado, J. M. (1994) Distribucion geografica de Speocharis sharpi (Escalera, 1898): consideraciones sobre la armadura del saco interno del edeago. Coexistencia con otros Bathysciinae (Coleoptera, Cholevidae). Nouvelle Revue d'Entomologie (N. S.), 11 (1), 5 - 17." type="journal article" year="1994">1994</bibRefCitation>
|
||
).
|
||
<taxonomicName id="60B14D629923793F148B0BD13A16F921" box="[661,879,1692,1717]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F148B0BD13A16F921" box="[661,879,1692,1717]" italics="true" pageId="7" pageNumber="32">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
was found only in samples from the central to the deepest parts of the caves (in greater numbers), whereas the specimens of
|
||
<taxonomicName id="60B14D629923793F127B0B8C3C21F94E" box="[1125,1368,1729,1754]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="sharpi" subSpecies="nigricans">
|
||
<emphasis id="95C5EAF39923793F127B0B8C3C21F94E" box="[1125,1368,1729,1754]" italics="true" pageId="7" pageNumber="32">Q. sharpi nigricans</emphasis>
|
||
</taxonomicName>
|
||
were collected in small series particulary near the cave entrances.
|
||
</paragraph>
|
||
<paragraph id="A70E36E19923793F16D80A413D3AF801" blockId="7.[151,1437,1166,2016]" pageId="7" pageNumber="32">
|
||
<taxonomicName id="60B14D629923793F16D80A4138D9F8B1" box="[198,416,1804,1829]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="pasensis">
|
||
<emphasis id="95C5EAF39923793F16D80A4138D9F8B1" box="[198,416,1804,1829]" italics="true" pageId="7" pageNumber="32">Quaestus pasensis</emphasis>
|
||
</taxonomicName>
|
||
was always collected in accumulations of organic matter. In El Churrón Cave, it has been found in wet remains of rotten wood; in the four other caves in old and mouldy bat manure. This species reaches the dark zone of the caves, near very wet areas; it has been found in deep zones where the air humidity is near saturation point, 95 to 99 %, and temperatures are between 10 and 14 ºC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFAB656A9923793F16D80AED38B8F874" pageId="7" pageNumber="32" type="etymology">
|
||
<paragraph id="A70E36E19923793F16D80AED38B8F874" blockId="7.[151,1437,1166,2016]" pageId="7" pageNumber="32">
|
||
<emphasis id="95C5EAF39923793F16D80AED3829F82E" bold="true" box="[198,336,1952,1978]" pageId="7" pageNumber="32">Etymology.</emphasis>
|
||
The specific name is derived from the Pas Mountains (Cantabria,
|
||
<collectingCountry id="DFA676719923793F129E0AEC3DBEF82F" box="[1152,1223,1953,1979]" name="Spain" pageId="7" pageNumber="32">Spain</collectingCountry>
|
||
), the karst region where the species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |