treatments-xml/data/1A/40/05/1A4005F3A3258B383069F3A7EDCCB0A6.xml
2024-06-21 12:30:36 +02:00

350 lines
31 KiB
XML

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. &amp; Dowling, Ashley P. G." docDate="2017" docId="1A4005F3A3258B383069F3A7EDCCB0A6" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola irapalpa Fisher &amp; Dowling, sp. n." docType="treatment" docUuid="FEAF955F-6EC2-4A5C-B97F-6168103C5003" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Danielle M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nelson, Whitney A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>701</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>496</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="134839657" LSID="urn:lsid:zoobank.org:act:FEAF955F-6EC2-4A5C-B97F-6168103C5003" httpUri="http://treatment.plazi.org/id/1A4005F3A3258B383069F3A7EDCCB0A6" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/FEAF955F-6EC2-4A5C-B97F-6168103C5003" authority="Fisher &amp; Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa Fisher &amp; Dowling</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">HOLOTYPE (♀): from New Brunswick, Canada, York County, Stanley, Nashwaak River, Stanley Municipal Park, 19 Jun 2012, by IM Smith, IMS120031, DNA 2956.</paragraph>
<paragraph pageId="0" pageNumber="1">
PARATYPES (39 ♀; 26 ♂): Arkansas, USA: 1 ♀ from Newton County, Buffalo National River, Cecil Creek (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="36.0877">36°5'15.72&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-93.22314">93°13'23.28&quot;W</geoCoordinate>
), 13 Jun 2012, by TD Edwards, TDE 12-0613-010
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Newton County, Ozark-St. Francis National Forest, Little Buffalo River, 11 Jun 2012, by TD Edwards, TDE 12-0711-004
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ from Polk County, beside Forest Route 38, north of Shady Lake Recreation Area, East Saline Creek (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.37316">34°22'23.39&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-94.0309">94°1'51.22&quot;W</geoCoordinate>
), 30 Jul 2011, by IM Smith, IMS110041
<normalizedToken originalValue="•">*</normalizedToken>
Kentucky, USA: 2 ♀ and 1 ♂ from McCreary County, White Oak Junction, Rock Creek, beside Forest Route 556, 2.3 kilometers south of Route 1363, 8 Jul 1990, by IM Smith, IMS900082A &amp; IMS900082B
<normalizedToken originalValue="•">*</normalizedToken>
Illinois, USA: 3 ♀ and 1 ♂ from Kankakee County, Kankakee River State Park, beside Route 102, Rock Creek (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.2">41°12'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-88.0">88°0'W</geoCoordinate>
), 9 Sep 1991, by IM Smith, IMS910027A
<normalizedToken originalValue="•">*</normalizedToken>
Indiana, USA: 1 ♀ from Wayne County, south of intersection of Interstate 70 and Route 1 (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="39.85361">39°51'13&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-85.134445">85°8'4&quot;W</geoCoordinate>
), 31 Jul 2014, by MJ Skvarla, MS 14-0731-001
<normalizedToken originalValue="•">*</normalizedToken>
Missouri, USA: 3 ♀ Wayne County, beside Road 143 near Sam A Baker State Park, 8 Jul 1960, by DR Cook, DRC600010
<normalizedToken originalValue="•">*</normalizedToken>
New Brunswick, Canada: 2 ♀ and 1 ♂ from Charlotte County, Rollingdam, Digdeguash River, beside Highway 770 at covered bridge, 30 Jun 1989, by IM Smith, IMS890053
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Charlotte County, Digdeguash River, beside Sorrel Ridge Road, west of Whittier Road, 10 Jun 2012, by IM Smith, IMS120015
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 1 ♂ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS110118
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ (ALLOTYPE) from York County, Stanley, Nashwaak River, Stanley Municipal Park, 19 Jun 2012, by IM Smith, IMS120031, DNA 2957
<normalizedToken originalValue="•">*</normalizedToken>
Ohio, USA: 1 ♂ from Montgomery County, Engelwood Metro Park (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="39.882774">39°52'58&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.292496">84°17'33&quot;W</geoCoordinate>
), 31 Jul 2014, by MJ Skvarla, MS 14-0731-002
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 2 ♂ from Pickaway County, beside Scioto-Darby Road, just north of Darby, Big Darby Creek (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="40.033333">40°2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-83.25">83°15'W</geoCoordinate>
), 30 Jun 1997, by IM Smith &amp; M MacKenzie, IMS970016
<normalizedToken originalValue="•">*</normalizedToken>
Ontario, Canada: 1 ♀ and 1 ♂ from Grey County, Durham, Saugeen River, beside County Road 27 near Durham Conservation Area, 9 Jun 1989, by IM Smith, IMS890028A
<normalizedToken originalValue="•">*</normalizedToken>
Pennsylvania, USA: 1 ♀ from Bedford County, Chaneysville, Sweet Root Picnic Area beside Route 326, 18 Jul 1990, by IM Smith, IMS900105
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Huntingdon County, Alan Seeger Natural Area, Stone Creek, beside road from McAlevys Fort to Route 322, 19 Jul 1990, by IM Smith, IMS900107
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Tioga County, Straight Run, beside Straight Run Road, 2.1 kilometers north of Route 6 between Ansonia &amp; Wellsboro, 20 Jul1990, by IM Smith, IMS900111
<normalizedToken originalValue="•">*</normalizedToken>
Saskatchewan, Canada: 2 ♀ and 2 ♂ from Torch River beside Highway 106 at access road, 30 kilometers north of Highway 55 at Smeaton, 30 Jul 1988, by IM Smith, IMS880054A
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Torch River at end of forest access road, 10 kilometers north of Highway 55 at Love, 30 Jul 1988, by IM Smith, IMS880056
<normalizedToken originalValue="•">*</normalizedToken>
Texas, USA: 2 ♀ from Bandera County, Vanderpool, beside Route 187, Sabinal River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="29.802776">29°48'10&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-99.575">99°34'30&quot;W</geoCoordinate>
), 2 May 2009, by IM Smith, IMS090007
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Kinney County, Brackettville, beside Route 90, 12.1 kilometers west of Route 131, Pinto Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="29.335001">29°20'6&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-100.53472">100°32'5&quot;W</geoCoordinate>
), 28 Sep 1994, by IM Smith, IMS940025
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Uvalde County, Garner State Park, Frio River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="29.589445">29°35'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-99.736664">99°44'12&quot;W</geoCoordinate>
), 28 May 1998, by IM Smith, IMS980027A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Val Verde County, Bakers Crossing Campground off Route 163, Devils River (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.966667">29°58'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-101.15">101°9'W</geoCoordinate>
), 5 Oct 1999, by IM Smith, IMS990061A
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ from Val Verde County, Dolan Falls Preserve, Devils River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="29.886667">29°53'12&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-100.99361">100°59'37&quot;W</geoCoordinate>
), 24 May 2011, by IM Smith, IMS110013
<normalizedToken originalValue="•">*</normalizedToken>
4 ♀ and 4 ♂ from Val Verde County, Dolan Falls Preserve, Snake Spring (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="29.895277">29°53'43&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-100.98278">100°58'58&quot;W</geoCoordinate>
), 25 May 2011, by IM Smith, IMS110015
<normalizedToken originalValue="•">*</normalizedToken>
Virginia, USA: 1 ♀ and 1 ♂ from Bath County, Jackson River, beside Forest Route 1843 (continuation of Route 623), 3.5 kilometers south of Route 220, 16 Jul 1990, by IM Smith, IMS900100
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Scott County, North Fork Holston River, beside Route 58/421, 0.9 kilometers east of Route 709 at Hiltons, 7 Jul 1990, by IM Smith, IMS900080
<normalizedToken originalValue="•">*</normalizedToken>
West Virginia, USA: 1 ♀ and 1 ♂ from Pendleton County, South Branch of the Potomac River, beside Route 28/55, 20.8 kilometers southwest of Route 42, 17 Jul 1990, by IM Smith, IMS900104.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type deposition">
<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and most paratypes (34 ♀; 20 ♂) deposited in the CNC; other paratypes (5 ♀; 5 ♂) deposited in ACUA.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa</taxonomicName>
are similar to other members of the Raptor Group (
<taxonomicName lsidName="T. gnoma" pageId="0" pageNumber="1" rank="species" species="gnoma">T. gnoma</taxonomicName>
,
<taxonomicName lsidName="T. longitibia" pageId="0" pageNumber="1" rank="species" species="longitibia">T. longitibia</taxonomicName>
,
<taxonomicName lsidName="T. mjolniri" pageId="0" pageNumber="1" rank="species" species="mjolniri">T. mjolniri</taxonomicName>
,
<taxonomicName lsidName="T. elusiva" pageId="0" pageNumber="1" rank="species" species="elusiva">T. elusiva</taxonomicName>
,
<taxonomicName lsidName="T. racupalpa" pageId="0" pageNumber="1" rank="species" species="racupalpa">T. racupalpa</taxonomicName>
,
<taxonomicName lsidName="T. raptor" pageId="0" pageNumber="1" rank="species" species="raptor">T. raptor</taxonomicName>
,
<taxonomicName lsidName="T. danielleae" pageId="0" pageNumber="1" rank="species" species="danielleae">T. danielleae</taxonomicName>
,
<taxonomicName lsidName="T. daemon" pageId="0" pageNumber="1" rank="species" species="daemon">T. daemon</taxonomicName>
, and
<taxonomicName lsidName="T. ivyae" pageId="0" pageNumber="1" rank="species" species="ivyae">T. ivyae</taxonomicName>
) in having round bodies; Dgl-4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae.
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. longitibia" pageId="0" pageNumber="1" rank="species" species="longitibia">T. longitibia</taxonomicName>
,
<taxonomicName lsidName="T. mjolniri" pageId="0" pageNumber="1" rank="species" species="mjolniri">T. mjolniri</taxonomicName>
,
<taxonomicName lsidName="T. elusiva" pageId="0" pageNumber="1" rank="species" species="elusiva">T. elusiva</taxonomicName>
,
<taxonomicName lsidName="T. racupalpa" pageId="0" pageNumber="1" rank="species" species="racupalpa">T. racupalpa</taxonomicName>
, and
<taxonomicName lsidName="T. ivyae" pageId="0" pageNumber="1" rank="species" species="ivyae">T. ivyae</taxonomicName>
by having Dgl-4 closer to the dorsal edge (dorsal width/distance between Dgl-4 = 1.81-2.09 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 2.19-2.77 in others) and a less elongate rostrum (length/width = 2.66-3.39 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
; 3.56-4.32 in others).
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. gnoma" pageId="0" pageNumber="1" rank="species" species="gnoma">T. gnoma</taxonomicName>
by having Dgl-4 closer to the dorsal edge (dorsal width/distance between Dgl-4 ♀ = 1.81-2.09 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 2.65-3.29 in
<taxonomicName lsidName="T. gnoma" pageId="0" pageNumber="1" rank="species" species="gnoma">T. gnoma</taxonomicName>
; ♂ = 1.58-1.86 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 2.06-2.73 in
<taxonomicName lsidName="T. gnoma" pageId="0" pageNumber="1" rank="species" species="gnoma">T. gnoma</taxonomicName>
) and by dorsal coloration and pattern.
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. raptor" pageId="0" pageNumber="1" rank="species" species="raptor">T. raptor</taxonomicName>
by having less elongate tibiae (length/width ♀ = 4.09-5.67 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 6.00-7.54 in
<taxonomicName lsidName="T. raptor" pageId="0" pageNumber="1" rank="species" species="raptor">T. raptor</taxonomicName>
; ♂ = 4.25-4.75 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 5.29-5.63 in
<taxonomicName lsidName="T. raptor" pageId="0" pageNumber="1" rank="species" species="raptor">T. raptor</taxonomicName>
); less elongate subcapitulum (ventral length/height = 2.52-2.90 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 2.98-3.27 in
<taxonomicName lsidName="T. raptor" pageId="0" pageNumber="1" rank="species" species="raptor">T. raptor</taxonomicName>
); and by dorsal pattern.
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. danielleae" pageId="0" pageNumber="1" rank="species" species="danielleae">T. danielleae</taxonomicName>
by having Dgl-4 further from the dorsal edge (dorsal width/distance between Dgl-4 = ♀ = 1.81-2.09 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 1.57-1.70 in
<taxonomicName lsidName="T. danielleae" pageId="0" pageNumber="1" rank="species" species="danielleae">T. danielleae</taxonomicName>
; ♂ = 1.58-1.86 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 1.42-1.52 in
<taxonomicName lsidName="T. danielleae" pageId="0" pageNumber="1" rank="species" species="danielleae">T. danielleae</taxonomicName>
); a less elongate rostrum (length/width = 2.66-3.39 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 3.43-3.75 in
<taxonomicName lsidName="T. danielleae" pageId="0" pageNumber="1" rank="species" species="danielleae">T. danielleae</taxonomicName>
); and by dorsal coloration and pattern. Female
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from female
<taxonomicName lsidName="T. daemon" pageId="0" pageNumber="1" rank="species" species="daemon">T. daemon</taxonomicName>
by having Dgl-4 further from the dorsal edge (dorsal width/distance between Dgl-4 ♀ = 1.81-2.09 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 1.59-1.67 in
<taxonomicName lsidName="T. daemon" pageId="0" pageNumber="1" rank="species" species="daemon">T. daemon</taxonomicName>
) and a less elongate gnathosomal bay (length/width ♀ = 1.35-1.86 in
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
, 1.95-2.42 in
<taxonomicName lsidName="T. daemon" pageId="0" pageNumber="1" rank="species" species="daemon">T. daemon</taxonomicName>
). Male
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
can be differentiated from male
<taxonomicName lsidName="T. daemon" pageId="0" pageNumber="1" rank="species" species="daemon">T. daemon</taxonomicName>
by dorsal coloration and pattern.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female (Figure 103) (n = 9) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (575-710 (610) long; 465-580 (475) wide) circular with navy blue to bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (132.5-160 (137.5) long; 52.5-67.5 (62.5) wide). Anterio-lateral platelets (155-197.5 (177.5) long; 67.5-87.5 (77.5) wide) free from dorsal plate. Dgl-4 midway between muscle scars and dorsum edge (distance between Dgl-4 465-580 (475)). Dorsal plate proportions: dorsum length/width 1.20-1.28 (1.28); dorsal width/distance between Dgl-4 1.81-2.09 (1.90); anterio-medial platelet length/width 2.12-2.52 (2.20); anterio-lateral platelet length/width 2.17-2.39 (2.29); anterio-lateral/anterio-medial length 1.17-1.42 (1.29).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (300-360 (340) long (ventral); 219-270 (270) long (dorsal); 125-157.5 (145) tall) with faint navy blue to bluish-purple coloration, sometimes colorless. Rostrum (126.25-152.5 (142.5) long; 55-76 (65) wide) elongate. Chelicerae (245-360 (360) long) with curved fangs (55-75 (65) long). Subcapitular proportions: ventral length/height 2.29-2.57 (2.34); rostrum length/width 2.66-3.39 (3.17). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45-52.5 (45) long); femur (115-132.5 (132.5) long); genu (65-80 (72.5) long); tibia (100-117.5 (102.5) long; 17.5-20 (20) wide); tarsus (17.5-20 (20) long). Palpomere proportions: femur/genu 1.66-1.89 (1.83); tibia/femur 0.77-0.89 (0.77); tibia length/width 4.09-5.67 (4.82).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (690-870 (770) long; 508-743 (550) wide) with navy blue to bluish purple coloration. Gnathosomal bay (132.5-165 (165) long; 87.5-120 (90) wide). Cxgl-4 far from apex. Medial suture (10-22.5 (15) long). Genital plates (155-175 (175) long; 140-162.5 (150) wide). Additional measurements: Cx-1 (266-334 (310) long (total); 103-156 (140) long (medial)); Cx-3 (332-487 (375) wide); anterior venter (156-197.5 (182.5) long). Ventral proportions: gnathosomal bay length/width 1.35-1.86 (1.83); anterior venter/genital field length 0.99-1.18 (1.04); anterior venter length/genital field width 1.06-1.27 (1.22); anterior venter/medial suture 8.56-15.63 (12.17).</paragraph>
<paragraph pageId="0" pageNumber="1">Male (Figure 104) (n = 8) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (520-620 (560) long; 400-490 (435) wide) circular with navy blue to bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (115-150 (140) long; 55-67.5 (62.5) wide). Anterio-lateral platelets (160-188.75 (187.5) long; 62.5-77.5 (75) wide) free from dorsal plate. Dgl-4 midway between muscle scars and dorsum edge (distance between Dgl-4 220-310 (250)). Dorsal plate proportions: dorsum length/width 1.26-1.30 (1.29); dorsal width/distance between Dgl-41.58-1.86 (1.74); anterio-medial platelet length/width 2.00-2.48 (2.24); anterio-lateral platelet length/width 2.34-2.60 (2.50); anterio-lateral/anterio-medial length 1.22-1.41 (1.34).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (275-340 (310) long (ventral); 207-240 (240) long (dorsal); 105-132.5 (107.5) tall) with faint navy blue to bluish-purple coloration, sometimes colorless. Rostrum (112.5-137.5 (135) long; 40-47.5 (40) wide) elongate. Chelicerae (252-305 (305) long) with curved fangs (51-60 (55) long). Subcapitular proportions: ventral length/height 2.52-2.90 (2.88); rostrum length/width 2.81-3.38 (3.38). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35-50 (42.5) long); femur (107.5-123.75 (115) long); genu (60-70 (67.5) long); tibia (87.5-108.75 (95) long; 20-25 (20) wide); tarsus (18.75-20 (20) long). Palpomere proportions: femur/genu 1.70-1.83 (1.70); tibia/femur 0.81-0.91 (0.83); tibia length/width 4.25-4.75 (4.75).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (630-705 (690) long; 490-540 (500) wide) with navy blue to bluish purple coloration. Gnathosomal bay (135-177.5 (146.25) long; 75-95 (75) wide). Cxgl-4 far from apex. Medial suture (50-75 (75) long). Genital plates (127.5-158.75 (143.75) long; 100-137.5 (122.5) wide). Additional measurements: Cx-1 (260-290 (290) long (total); 137-155 (155) long (medial)); Cx-3 (323-360 (345) wide); anterior venter (215-255 (245) long). Ventral proportions: gnathosomal bay length/width 1.42-2.06 (1.95); anterior venter/genital field length 1.51-1.75 (1.70); anterior venter length/genital field width 1.78-2.20 (2.00); anterior venter/medial suture 3.27-4.40 (3.27).</paragraph>
<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Specific epithet (
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">irapalpa</taxonomicName>
) refers to the pedipalps of this species, which resemble ferocious weapons of wrath (ira, L. fury, rage, wrath; palpus, L. hand, feeler).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Primarily eastern (Figure 102), but note that
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
is one of few
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
found in the Great Plains, at least in the south (Texas) and north (Saskatchewan).
</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 102.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa</taxonomicName>
sp. n. distribution.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 103.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa</taxonomicName>
sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 104.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa</taxonomicName>
sp. n. male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola irapalpa" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irapalpa">Torrenticola irapalpa</taxonomicName>
groups with other members of the Raptor Complex with high support and specimens of this species are less than 2% different in COI sequence from each other. In the combined analysis,
<taxonomicName lsidName="T. irapalpa" pageId="0" pageNumber="1" rank="species" species="irapalpa">T. irapalpa</taxonomicName>
groups with
<taxonomicName lsidName="T. gnoma" pageId="0" pageNumber="1" rank="species" species="gnoma">T. gnoma</taxonomicName>
with high support, but the position of this clade was not recovered. These species are greater than 9% from each other. Based upon overall similarity, distribution, and phylogenetic position, this species is placed within the Raptor Identification Group.
</paragraph>
<paragraph pageId="0" pageNumber="1">This species hypothesis is supported by low COI divergence within the species (0-2%) and high divergence between species (3-15%), and by the morphological characters outlined in the diagnosis.</paragraph>
</subSubSection>
</treatment>
</document>