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<document id="8ADD2B911ACC268F7B51A03040F2AE7A" ID-CLB-Dataset="3695" ID-DOI="10.35929/RSZ.0050" ID-GBIF-Dataset="60a46419-793d-4cae-a0bc-a086fc235849" ID-ISSN="0035-418" ID-Zenodo-Dep="5640135" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635893751209" checkinUser="felipe" docAuthor="Balkenohl, Michael" docDate="2021" docId="014C8648FFE3BB3D324577C9FB66FC1B" docLanguage="en" docName="RevSuisseZool.128.2.357-374.pdf" docOrigin="Revue suisse de Zoologie 128 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Syleter sinepunctatus subsp. sinepunctatus sinepunctatus 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="367" masterDocId="FD75FE30FFEABB36313E7638FFFBFFB0" masterDocTitle="Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="374" masterPageNumber="357" pageNumber="365" updateTime="1699467467588" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0C6BEFDDDE94869C095E76C06F812C0F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB5460555956E79AF25207B424DAAC80">
<mods:title id="D0DC440D48E39A8450747E1A59B5615E">Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="01F5B33617EB30A3B219DA23C2C81637">Balkenohl, Michael</mods:namePart>
<mods:affiliation id="C5DDD4BFCC37AA7A0C3806A38223A586">Ligusterweg 9, CH- 8906 Bonstetten, Switzerland</mods:affiliation>
<mods:nameIdentifier id="C7252AB37465D1E51C5A033CFBD49544" type="email">mike.balkenohl@bluewin.ch</mods:nameIdentifier>
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<mods:title id="275DE3A5724942035C2840DE38240828">Revue suisse de Zoologie</mods:title>
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<mods:date id="0345527ADAFC70444B03A3A5005C540A">2021</mods:date>
<mods:detail id="F3DEEC7618D6E53303F87726273C0DAD" type="volume">
<mods:number id="98ABE950F6DEEACBFCDB2FD4E92A2601">128</mods:number>
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<mods:classification id="1900BE3BCCFC4741B9203CF6209B889A">journal article</mods:classification>
<mods:identifier id="FD62CBA40757016FC922AB22D43721E7" type="CLB-Dataset">3695</mods:identifier>
<mods:identifier id="6076D691889455A5D96CD7DC93F54D97" type="DOI">10.35929/RSZ.0050</mods:identifier>
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<mods:identifier id="6E4BA94B4BED0BFEC460089A65D141E2" type="ISSN">0035-418</mods:identifier>
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<treatment id="014C8648FFE3BB3D324577C9FB66FC1B" ID-DOI="http://doi.org/10.5281/zenodo.5646414" ID-GBIF-Taxon="189991363" ID-Zenodo-Dep="5646414" LSID="urn:lsid:plazi:treatment:014C8648FFE3BB3D324577C9FB66FC1B" httpUri="http://treatment.plazi.org/id/014C8648FFE3BB3D324577C9FB66FC1B" lastPageId="11" lastPageNumber="367" pageId="9" pageNumber="365">
<subSubSection id="C1FF64D5FFE3BB3F324577C9FA9AFDB9" box="[891,1377,497,521]" pageId="9" pageNumber="365" type="nomenclature">
<paragraph id="895A375EFFE3BB3F324577C9FA9AFDB9" blockId="9.[891,1377,497,552]" box="[891,1377,497,521]" pageId="9" pageNumber="365">
<heading id="D2128032FFE3BB3F324577C9FA9AFDB9" bold="true" box="[891,1377,497,521]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="365" reason="2">
<emphasis id="BB91EB4CFFE3BB3F324577C9FA9AFDB9" bold="true" box="[891,1377,497,521]" pageId="9" pageNumber="365">
<taxonomicName id="4EE54CDDFFE3BB3F324577C9FAFEFDB9" authority="Balkenohl, 2021" authorityName="sinepunctatus" authorityYear="2021" box="[891,1285,497,521]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="365" phylum="Arthropoda" rank="subSpecies" species="sinepunctatus" status="sp. nov." subSpecies="sinepunctatus">
<emphasis id="BB91EB4CFFE3BB3F324577C9FAFEFDB9" bold="true" box="[891,1285,497,521]" italics="true" pageId="9" pageNumber="365">Syleter sinepunctatus sinepunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE3BB3F343277C9FA9AFDB9" box="[1292,1377,497,521]" pageId="9" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE3BB3F35217428FB46FD98" box="[1055,1213,528,552]" pageId="9" pageNumber="365" type="description">
<paragraph id="895A375EFFE3BB3F35217428FB46FD98" blockId="9.[891,1377,497,552]" box="[1055,1213,528,552]" pageId="9" pageNumber="365">
<figureCitation id="11DE2BDBFFE3BB3F35217428FB95FD98" box="[1055,1134,528,552]" captionStart="Figs 7-13" captionStartId="8.[151,192,1967,1989]" captionTargetBox="[190,1390,262,1901]" captionTargetId="figure-4@8.[183,1410,256,1911]" captionTargetPageId="8" captionText="Figs 7-13. Syleter spp., habitus. (7) S. paradoxus (Putzeys, 1868), specimen from Bangkok, female. (8) S. acutipalpis (Putzeys, 1877), lectotype. (9) S. fulvaster (Andrewes, 1927), paratype, male. (10) S. papua Darlington, 1962, paratype, female. (11) S. sinepunctatus sinepunctatus sp. nov., holotype. (12) S. sinepunctatus kluangensis ssp. nov., holotype. (13) S. gradus sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.5640142" httpUri="https://zenodo.org/record/5640142/files/figure.png" pageId="9" pageNumber="365">Figs 11</figureCitation>
,
<figureCitation id="11DE2BDBFFE3BB3F35447428FB6CFD98" box="[1146,1175,528,552]" captionStart="Figs 14-20" captionStartId="9.[695,736,1848,1870]" captionTargetId="figure-270@9.[151,1431,769,2095]" captionTargetPageId="9" captionText="Figs 14-20. Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (14) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand. (15) S. acutipalpis (Putzeys, 1877), lectotype. (16) S. fulvaster (Andrewes, 1927), paratype. (17) S. papua Darlington, 1962, paratype.(18) S. sinepunctatus sinepunctatus sp. nov., holotype. (19) S. sinepunctatus kluangensis ssp. nov., holotype. (20) S. gradus sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.5646404" httpUri="https://zenodo.org/record/5646404/files/figure.png" pageId="9" pageNumber="365">18</figureCitation>
,
<figureCitation id="11DE2BDBFFE3BB3F359C7428FB46FD98" box="[1186,1213,528,552]" captionStart="Figs 21-27" captionStartId="10.[151,192,1236,1258]" captionTargetBox="[189,1399,262,1192]" captionTargetId="figure-388@10.[151,1431,257,1223]" captionTargetPageId="10" captionText="Figs 21-27. Female coxostyli with laterotergite. (21) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand, with latero- and mediotergite. (22) S. acutipalpis (Putzeys, 1877), specimen from Lilong, China. (23) S. fulvaster (Andrewes, 1927), holotype. (24) S. papua Darlington, 1962, paratype. (25) S. sinepunctatus sinepunctatus sp. nov., paratype. (26) S. sinepunctatus kluangensis sp. nov., paratype. (27) S. gradus sp. nov. paratype." figureDoi="http://doi.org/10.5281/zenodo.5640144" httpUri="https://zenodo.org/record/5640144/files/figure.png" pageId="9" pageNumber="365">25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE3BB3C32017477FDA9F89B" lastPageId="10" lastPageNumber="366" pageId="9" pageNumber="365" type="materials_examined">
<paragraph id="895A375EFFE3BB3F32017477FC07FD76" blockId="9.[831,1437,591,710]" pageId="9" pageNumber="365">
<materialsCitation id="398D3D03FFE3BB3F32017477FC07FD76" ID-GBIF-Occurrence="3398244310" collectingDate="28. VIII. - 5. IX. 82" collectionCode="BMNH" collectorName="B. E. Stork" country="Brunei" location="Nn hamunin" pageId="9" pageNumber="365" specimenCount="1" specimenCount-male="1" stateProvince="Bukit Sulang" typeStatus="holotype">
<emphasis id="BB91EB4CFFE3BB3F32017477FC49FDD7" bold="true" box="[831,946,591,615]" pageId="9" pageNumber="365">
<typeStatus id="565E89FCFFE3BB3F32017477FC57FDD7" box="[831,940,591,615]" pageId="9" pageNumber="365" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="EFF4AF9BFFE3BB3F32FF7477FBEEFDD6" box="[961,1045,591,614]" pageId="9" pageNumber="365">BMNH</collectionCode>
;
<specimenCount id="9FE3FCD7FFE3BB3F35167477FBC5FDD6" box="[1064,1086,591,614]" pageId="9" pageNumber="365" type="male">♂</specimenCount>
, with labels and data: white, handwritten in black ink “32
<collectingCountry id="F1F277CEFFE3BB3F35427457FB24FD36" box="[1148,1247,623,646]" name="Brunei" pageId="9" pageNumber="365">BRUNEI</collectingCountry>
<collectingRegion id="4B21F9BCFFE3BB3F35D97456FA8FFD36" box="[1255,1396,622,646]" pageId="9" pageNumber="365">Bukit Sulang</collectingRegion>
<location id="8C3A6185FFE3BB3F34427457FC64FD16" LSID="urn:lsid:plazi:treatment:014C8648FFE3BB3D324577C9FB66FC1B:8C3A6185FFE3BB3F34427457FC64FD16" country="Brunei" name="Nn hamunin" pageId="9" pageNumber="365" stateProvince="Bukit Sulang">Nn hamunin</location>
,
<collectingDate id="ED1FE876FFE3BB3F329274B6FBA5FD16" box="[940,1118,654,678]" pageId="9" pageNumber="365">28.VIII.-5.IX.82</collectingDate>
, B.E. Stork B.M. 1982-388”, printed “at light”.
</materialsCitation>
</paragraph>
<caption id="DD9A67D6FFE3BB3F33897100FBBCF7A8" ID-DOI="http://doi.org/10.5281/zenodo.5646404" ID-Zenodo-Dep="5646404" httpUri="https://zenodo.org/record/5646404/files/figure.png" pageId="9" pageNumber="365" startId="9.[695,736,1848,1870]" targetBox="[200,1437,791,2072]" targetPageId="9">
<paragraph id="895A375EFFE3BB3F33897100FBBCF7A8" blockId="9.[695,1437,1848,2072]" pageId="9" pageNumber="365">
Figs 14-20. Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (14)
<emphasis id="BB91EB4CFFE3BB3F35D0716DFA67F8DB" box="[1262,1436,1877,1899]" italics="true" pageId="9" pageNumber="365">Syleter paradoxus</emphasis>
(Putzeys, 1868), specimen from Tale Noi, Thailand. (15)
<emphasis id="BB91EB4CFFE3BB3F320271B7FC3CF815" box="[828,967,1935,1957]" italics="true" pageId="9" pageNumber="365">S. acutipalpis</emphasis>
(Putzeys, 1877), lectotype. (16)
<emphasis id="BB91EB4CFFE3BB3F341671B7FA67F815" box="[1320,1436,1935,1957]" italics="true" pageId="9" pageNumber="365">S. fulvaster</emphasis>
(Andrewes, 1927), paratype. (17)
<emphasis id="BB91EB4CFFE3BB3F35B17193FB1DF871" box="[1167,1254,1963,1985]" italics="true" pageId="9" pageNumber="365">S. papua</emphasis>
Darlington, 1962, paratype. (18)
<emphasis id="BB91EB4CFFE3BB3F32F871F0FB11F86E" box="[966,1258,1992,2014]" italics="true" pageId="9" pageNumber="365">S. sinepunctatus sinepunctatus</emphasis>
sp. nov., holotype. (19)
<emphasis id="BB91EB4CFFE3BB3F324F71DDFB69F84B" box="[881,1170,2021,2043]" italics="true" pageId="9" pageNumber="365">S. sinepunctatus kluangensis</emphasis>
ssp. nov., holotype. (20)
<emphasis id="BB91EB4CFFE3BB3F32027E3AFC6CF7A8" box="[828,919,2050,2072]" italics="true" pageId="9" pageNumber="365">S. gradus</emphasis>
sp. nov. holotype.
</paragraph>
</caption>
<caption id="DD9A67D6FFE0BB3C31A972ECFBD9FAF0" ID-DOI="http://doi.org/10.5281/zenodo.5640144" ID-Zenodo-Dep="5640144" httpUri="https://zenodo.org/record/5640144/files/figure.png" pageId="10" pageNumber="366" startId="10.[151,192,1236,1258]" targetBox="[189,1399,262,1192]" targetPageId="10">
<paragraph id="895A375EFFE0BB3C31A972ECFBD9FAF0" blockId="10.[151,1437,1236,1344]" pageId="10" pageNumber="366">
Figs 21-27. Female coxostyli with laterotergite. (21)
<taxonomicName id="4EE54CDDFFE0BB3C338E72ECFBFFFB5A" authority="(Putzeys, 1868)" baseAuthorityName="Putzeys" baseAuthorityYear="1868" box="[688,1028,1236,1258]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="BB91EB4CFFE0BB3C338E72ECFC9AFB5A" box="[688,865,1236,1258]" italics="true" pageId="10" pageNumber="366">Syleter paradoxus</emphasis>
(
<bibRefCitation id="ED744AAFFFE0BB3C324E72ECFC06FB5A" author="Putzeys J. A. A. H." box="[880,1021,1236,1258]" pageId="10" pageNumber="366" pagination="7 - 22" refId="ref14618" refString="Putzeys J. A. A. H. 1868. Supplement a la revision generale des Clivinides. Annales de la Societe Entomologique de Belgique 11: 7 - 22." type="journal article" year="1868">Putzeys, 1868</bibRefCitation>
)
</taxonomicName>
, specimen from Tale Noi, Thailand, with latero- and mediotergite. (22)
<taxonomicName id="4EE54CDDFFE0BB3C330272C9FCA6FAB7" authority="(Putzeys, 1877)" baseAuthorityName="Putzeys" baseAuthorityYear="1877" box="[572,861,1265,1287]" class="Insecta" family="Carabidae" genus="Psilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="acutipalpis">
<emphasis id="BB91EB4CFFE0BB3C330272C9FD45FAB7" box="[572,702,1265,1287]" italics="true" pageId="10" pageNumber="366">S. acutipalpis</emphasis>
(
<bibRefCitation id="ED744AAFFFE0BB3C33F572C9FCADFAB7" author="Putzeys J. A. A. H." box="[715,854,1265,1287]" pageId="10" pageNumber="366" refId="ref14683" refString="Putzeys J. A. A. H. 1877. Comptes-rendus des Seances de la Societe Entomologique de Belgique. Annales de la Societe Entomologique de Belgique 20: XL-XLVII." type="book" year="1877">Putzeys, 1877</bibRefCitation>
)
</taxonomicName>
, specimen from Lilong, China. (23)
<taxonomicName id="4EE54CDDFFE0BB3C358572C9FEADFA94" authority="(Andrewes, 1927)" baseAuthorityName="Andrewes" baseAuthorityYear="1927" class="Insecta" family="Carabidae" genus="Psilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="fulvaster">
<emphasis id="BB91EB4CFFE0BB3C358572C9FADCFAB7" box="[1211,1319,1265,1287]" italics="true" pageId="10" pageNumber="366">S. fulvaster</emphasis>
(
<bibRefCitation id="ED744AAFFFE0BB3C340A72C9FEB4FA94" author="Andrewes H. E." pageId="10" pageNumber="366" pagination="263 - 272" refId="ref13362" refString="Andrewes H. E. 1927. Papers on Oriental Carabidae. - XX. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, Ninth Series 20: 263 - 272." type="book chapter" year="1927">Andrewes, 1927</bibRefCitation>
)
</taxonomicName>
, holotype. (24)
<taxonomicName id="4EE54CDDFFE0BB3C30CF7336FD0FFA94" authority="Darlington, 1962" authorityName="Darlington" authorityYear="1962" box="[497,756,1294,1316]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="papua">
<emphasis id="BB91EB4CFFE0BB3C30CF7336FDBDFA94" box="[497,582,1294,1316]" italics="true" pageId="10" pageNumber="366">S. papua</emphasis>
<bibRefCitation id="ED744AAFFFE0BB3C33737336FD0FFA94" author="Darlington P. J." box="[589,756,1294,1316]" pageId="10" pageNumber="366" pagination="321 - 564" refId="ref14087" refString="Darlington P. J. 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology 126: 321 - 564." type="journal article" year="1962">Darlington, 1962</bibRefCitation>
</taxonomicName>
, paratype. (25)
<taxonomicName id="4EE54CDDFFE0BB3C32B07336FB4DFA93" authority="Balkenohl, 2021" authorityName="sinepunctatus" authorityYear="2021" box="[910,1206,1294,1316]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="subSpecies" species="sinepunctatus" status="sp. nov." subSpecies="sinepunctatus">
<emphasis id="BB91EB4CFFE0BB3C32B07336FB4DFA93" box="[910,1206,1294,1316]" italics="true" pageId="10" pageNumber="366">S. sinepunctatus sinepunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE0BB3C35837336FAF0FA93" box="[1213,1291,1294,1315]" pageId="10" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
, paratype. (26)
<taxonomicName id="4EE54CDDFFE0BB3C30227312FDD4FAF0" authorityName="Balkenohl" authorityYear="2021" box="[284,559,1322,1344]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="subSpecies" species="sinepunctatus" subSpecies="kluangensis">
<emphasis id="BB91EB4CFFE0BB3C30227312FDD4FAF0" box="[284,559,1322,1344]" italics="true" pageId="10" pageNumber="366">S. sinepunctatus kluangensis</emphasis>
</taxonomicName>
sp. nov., paratype. (27)
<taxonomicName id="4EE54CDDFFE0BB3C32297312FC89FAF0" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[791,882,1322,1344]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="gradus" status="sp. nov.">
<emphasis id="BB91EB4CFFE0BB3C32297312FC89FAF0" box="[791,882,1322,1344]" italics="true" pageId="10" pageNumber="366">S. gradus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE0BB3C32467313FC38FAF0" box="[888,963,1323,1344]" pageId="10" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
paratype.
</paragraph>
</caption>
<paragraph id="895A375EFFE0BB3C31A973B6FDA9F89B" blockId="10.[151,756,1421,1835]" pageId="10" pageNumber="366">
<materialsCitation id="398D3D03FFE0BB3C31A973B6FE99F9F8" ID-GBIF-Occurrence="3398244333" collectingDate="1961-07-28" collectingMethod="at light" collectionCode="BMNH" collectorName="R. O. S. Clarke." determinerName="Balkenohl" location="Kuching" pageId="10" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="SARAWAK" typeStatus="paratype">
<emphasis id="BB91EB4CFFE0BB3C31A973B6FEE9FA16" bold="true" box="[151,274,1422,1446]" pageId="10" pageNumber="366">
<typeStatus id="565E89FCFFE0BB3C31A973B6FEF6FA16" box="[151,269,1422,1446]" pageId="10" pageNumber="366" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="EFF4AF9BFFE0BB3C302273B6FE94FA15" box="[284,367,1422,1445]" pageId="10" pageNumber="366">BMNH</collectionCode>
;
<specimenCount id="9FE3FCD7FFE0BB3C304273B6FE5CFA15" box="[380,423,1422,1446]" pageId="10" pageNumber="366" type="female">1 ♀</specimenCount>
, with labels and data: orange, handwritten in black ink “
<location id="8C3A6185FFE0BB3C30C47396FDA3FA76" LSID="urn:lsid:plazi:treatment:014C8648FFE3BB3D324577C9FB66FC1B:8C3A6185FFE0BB3C30C47396FDA3FA76" box="[506,600,1454,1478]" name="Kuching" pageId="10" pageNumber="366" stateProvince="SARAWAK">Kuching</location>
<collectingRegion id="4B21F9BCFFE0BB3C334E7396FD0FFA76" box="[624,756,1454,1478]" pageId="10" pageNumber="366">SARAWAK</collectingRegion>
<date id="FD5B119EFFE0BB3C31A973F6FEFDFA56" box="[151,262,1486,1510]" pageId="10" pageNumber="366" value="1961-07-28">
<collectingDate id="ED1FE876FFE0BB3C31A973F6FEFDFA56" box="[151,262,1486,1510]" pageId="10" pageNumber="366" value="1961-07-28">28.7.1961</collectingDate>
</date>
”, “
<collectingMethod id="50A44F49FFE0BB3C307973F7FE54FA56" box="[327,431,1486,1510]" pageId="10" pageNumber="366">at light</collectingMethod>
”, white, black printed “R.O.S.Clarke. B.M. 1970-557.”, green, handwritten with pencil “
<taxonomicName id="4EE54CDDFFE0BB3C30117037FE85F997" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[303,382,1551,1575]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="genus" status="sp. nov.">Syleter</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE0BB3C30B27028FE44F997" box="[396,447,1552,1575]" pageId="10" pageNumber="366" rank="species">n.sp.</taxonomicNameLabel>
nr.
<taxonomicName id="4EE54CDDFFE0BB3C30C97037FD44F997" authority="PUTZ." authorityName="PUTZ." box="[503,703,1551,1575]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="paradoxus">paradoxus PUTZ.</taxonomicName>
det.
<determinerName id="7C86DB9AFFE0BB3C31A97008FEF2F9F8" box="[151,265,1584,1608]" pageId="10" pageNumber="366">Balkenohl</determinerName>
, 2000”
</materialsCitation>
. –
<materialsCitation id="398D3D03FFE0BB3C30AD7008FE71F939" ID-GBIF-Occurrence="3398244340" collectionCode="MFNB" collectorName="S. O. Borneo Grabowsky." location="Borneo" pageId="10" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Borneo" typeStatus="paratype">
<collectionCode id="EFF4AF9BFFE0BB3C30AD7008FE18F9F7" box="[403,483,1584,1607]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="366">MFNB</collectionCode>
;
<specimenCount id="9FE3FCD7FFE0BB3C30C87008FDD3F9F7" box="[502,552,1584,1608]" pageId="10" pageNumber="366" type="female">1 ♀</specimenCount>
, with labels and data: beige, black printed “S.O.
<collectingRegion id="4B21F9BCFFE0BB3C33387069FDAEF9D8" box="[518,597,1617,1640]" pageId="10" pageNumber="366">Borneo</collectingRegion>
Grabowsky.”, “Zool. Mus. Berlin”
</materialsCitation>
. –
<materialsCitation id="398D3D03FFE0BB3C30837049FDB9F95A" ID-GBIF-Occurrence="3398244348" collectingDate="1995-01-20" collectionCode="ADCW" collectorName="Seyfert &amp; Greindl" country="Indonesia" location="Tanjung Isuy" pageId="10" pageNumber="366" specimenCount="12" specimenCount-female="2" specimenCount-male="3" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="EFF4AF9BFFE0BB3C30837049FDE9F939" box="[445,530,1649,1673]" pageId="10" pageNumber="366">ADCW</collectionCode>
;
<specimenCount id="9FE3FCD7FFE0BB3C33197049FD8AF938" box="[551,625,1649,1673]" pageId="10" pageNumber="366" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9FE3FCD7FFE0BB3C33B97049FD34F938" box="[647,719,1649,1673]" pageId="10" pageNumber="366" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9FE3FCD7FFE0BB3C33D97049FEF6F919" pageId="10" pageNumber="366" type="generic">7 specimens</specimenCount>
., with labels and data: white, black printed “
<collectingCountry id="F1F277CEFFE0BB3C319D7089FEBCF979" box="[163,327,1713,1737]" name="Indonesia" pageId="10" pageNumber="366">INDONESIEN</collectingCountry>
:
<collectingRegion id="4B21F9BCFFE0BB3C305B7089FDC5F979" box="[357,574,1713,1737]" country="Indonesia" name="Kalimantan Timur" pageId="10" pageNumber="366">Kalimantan Timur</collectingRegion>
,
<location id="8C3A6185FFE0BB3C33697089FD0FF97A" LSID="urn:lsid:plazi:treatment:014C8648FFE3BB3D324577C9FB66FC1B:8C3A6185FFE0BB3C33697089FD0FF97A" box="[599,756,1713,1738]" country="Indonesia" name="Tanjung Isuy" pageId="10" pageNumber="366" stateProvince="Kalimantan Timur">Tanjung Isuy</location>
<date id="FD5B119EFFE0BB3C31A970EAFEF9F95A" box="[151,258,1746,1770]" pageId="10" pageNumber="366" value="1995-01-20">
<collectingDate id="ED1FE876FFE0BB3C31A970EAFEF9F95A" box="[151,258,1746,1770]" pageId="10" pageNumber="366" value="1995-01-20">20.1.1995</collectingDate>
</date>
leg.
<collectorName id="24105288FFE0BB3C300470EAFE7CF95A" box="[314,391,1746,1770]" pageId="10" pageNumber="366">Seyfert</collectorName>
&amp;
<collectorName id="24105288FFE0BB3C309370EAFE05F95A" box="[429,510,1746,1770]" pageId="10" pageNumber="366">Greindl</collectorName>
(34)”
</materialsCitation>
. –
<materialsCitation id="398D3D03FFE0BB3C335D70EAFDB5F89B" ID-GBIF-Occurrence="3398244315" collectingDate="1995-01-20" collectionCode="ADCW" collectorName="Zettel Wien" country="Indonesia" location="Tanjung Isuy" pageId="10" pageNumber="366" specimenCount="9" specimenCount-female="3" specimenCount-male="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="EFF4AF9BFFE0BB3C335D70EAFD4CF95A" box="[611,695,1746,1770]" pageId="10" pageNumber="366">ADCW</collectionCode>
;
<specimenCount id="9FE3FCD7FFE0BB3C33FD70EAFD14F959" box="[707,751,1746,1770]" pageId="10" pageNumber="366" type="male">1 ♂</specimenCount>
,
<specimenCount id="9FE3FCD7FFE0BB3C31A970CBFF2FF8BA" box="[151,212,1779,1803]" pageId="10" pageNumber="366" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="9FE3FCD7FFE0BB3C31DF70CAFE9CF8BA" box="[225,359,1778,1802]" pageId="10" pageNumber="366" type="generic">5 specimens</specimenCount>
, same data but “(33)” and “(34)” and orange, handwritten: “Coll.
<collectorName id="24105288FFE0BB3C30FC712BFDBFF89B" box="[450,580,1811,1835]" pageId="10" pageNumber="366">Zettel Wien</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE0BB3C31A9717BFB4DFA77" pageId="10" pageNumber="366" type="diagnosis">
<paragraph id="895A375EFFE0BB3C31A9717BFB4DFA77" blockId="10.[151,756,1858,2077]" lastBlockId="10.[831,1436,1421,1479]" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C31A9717BFEF4F8EB" bold="true" box="[151,271,1859,1883]" pageId="10" pageNumber="366">Diagnosis:</emphasis>
A medium sized species with seven-setose labrum, and the clypeus and frons isodiametrically reticulated. Differentiated from the similar species
<taxonomicName id="4EE54CDDFFE0BB3C31A9719CFED1F80B" box="[151,298,1955,1979]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="BB91EB4CFFE0BB3C31A9719CFED1F80B" box="[151,298,1955,1979]" italics="true" pageId="10" pageNumber="366">S. paradoxus</emphasis>
</taxonomicName>
and
<taxonomicName id="4EE54CDDFFE0BB3C3055719CFE2CF80B" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[363,471,1955,1979]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="gradus" status="sp. nov.">
<emphasis id="BB91EB4CFFE0BB3C3055719CFE2CF80B" box="[363,471,1955,1979]" italics="true" pageId="10" pageNumber="366">S. gradus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE0BB3C30DA719CFDC5F80B" box="[484,574,1956,1979]" pageId="10" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
by the missing punctures on the disk of the pronotum and the missing clypeo-frontal furrow.
<taxonomicName id="4EE54CDDFFE0BB3C30AC71DDFDD9F84C" box="[402,546,2020,2044]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="BB91EB4CFFE0BB3C30AC71DDFDD9F84C" box="[402,546,2020,2044]" italics="true" pageId="10" pageNumber="366">S. paradoxus</emphasis>
</taxonomicName>
differs in addition by the smooth clypeus.
<taxonomicName id="4EE54CDDFFE0BB3C30907E3EFDE1F7AD" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[430,538,2053,2077]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="366" phylum="Arthropoda" rank="species" species="gradus" status="sp. nov.">
<emphasis id="BB91EB4CFFE0BB3C30907E3EFDE1F7AD" box="[430,538,2053,2077]" italics="true" pageId="10" pageNumber="366">S. gradus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A25637FFE0BB3C33197E3EFD7BF7AD" box="[551,640,2054,2077]" pageId="10" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
differs in addition by the complete and step-like developed neck constriction, and by the larger size.
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE0BB3C320173D8FC5AF98A" pageId="10" pageNumber="366" type="etymology">
<paragraph id="895A375EFFE0BB3C320173D8FC5AF98A" blockId="10.[831,1436,1503,1594]" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C320173D8FC3FFA48" bold="true" box="[831,964,1504,1528]" pageId="10" pageNumber="366">Etymology:</emphasis>
The name refers to the completely missing punctures on the disc of the pronotum (Latin: sine – without).
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE0BB3D3201706BFB89FCEC" lastPageId="11" lastPageNumber="367" pageId="10" pageNumber="366" type="description">
<paragraph id="895A375EFFE0BB3C3201706BFC38F9DB" blockId="10.[831,1437,1619,2077]" box="[831,963,1619,1643]" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C3201706BFC38F9DB" bold="true" box="[831,963,1619,1643]" pageId="10" pageNumber="366">Description</emphasis>
</paragraph>
<paragraph id="895A375EFFE0BB3C3201704DFB1BF940" blockId="10.[831,1437,1619,2077]" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C3201704DFC21F93C" box="[831,986,1653,1676]" italics="true" pageId="10" pageNumber="366">Measurements</emphasis>
: Body length
<quantity id="4E1D9ABBFFE0BB3C3555704CFB0AF93C" box="[1131,1265,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="3.585" metricValueMax="3.8" metricValueMin="3.37" pageId="10" pageNumber="366" unit="mm" value="3.585" valueMax="3.8" valueMin="3.37">3.37-3.8 mm</quantity>
(x =
<quantity id="4E1D9ABBFFE0BB3C341D704CFA7EF93C" box="[1315,1413,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="3.64" pageId="10" pageNumber="366" unit="mm" value="3.64">3.64 mm</quantity>
*), width
<quantity id="4E1D9ABBFFE0BB3C32BA70ADFBE0F91D" box="[900,1051,1685,1709]" metricMagnitude="-3" metricUnit="m" metricValue="1.2149999999999999" metricValueMax="1.27" metricValueMin="1.16" pageId="10" pageNumber="366" unit="mm" value="1.2149999999999999" valueMax="1.27" valueMin="1.16">1.16-1.27 mm</quantity>
(x =
<quantity id="4E1D9ABBFFE0BB3C356670ADFB45F91D" box="[1112,1214,1685,1709]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="10" pageNumber="366" unit="mm" value="1.22">1.22 mm</quantity>
*), ratio length/width of pronotum 1.01-1.05 (x = 1.03*), ratio length/width of elytra 1.58-1.66 (x = 1.62*); (*n = 10).
</paragraph>
<paragraph id="895A375EFFE0BB3C320170C2FB25F883" blockId="10.[831,1437,1619,2077]" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C320170C2FC70F8A2" box="[831,907,1786,1810]" italics="true" pageId="10" pageNumber="366">Colour</emphasis>
: Piceous to fuscous. Legs fuscous. Antenna, tarsomeres and palpomeres testaceous.
</paragraph>
<paragraph id="895A375EFFE0BB3D32017104FD3DFCA1" blockId="10.[831,1437,1619,2077]" lastBlockId="11.[151,757,257,2077]" lastPageId="11" lastPageNumber="367" pageId="10" pageNumber="366">
<emphasis id="BB91EB4CFFE0BB3C32017104FC82F8E4" box="[831,889,1852,1876]" italics="true" pageId="10" pageNumber="366">Head</emphasis>
: A third narrower than pronotum, frons and clypeus anterior eye-level conspicuously elongated. Clypeus straight anteriorly, with lateral angle obtuse rounded; completely fused with clypeal wing and supraantennal plate, all reflexed margined, clypeus convex, separated from elongated convex supraantennal plate by parallel running deep and conspicuously wide clypeal furrows, extending up to anterior eye-level; supraantennal plate extended up to mid-eye level; clypeus separated from frons by indistinct transverse nearly not traceable change of the direction of the convexity. Frons moderately convex, with wide frontal furrows, with distinct lateral carina separating frontal furrow from supraorbital furrow, frontal furrow and lateral carina wide, slightly diverging and extending nearly up to neck constriction. Clypeal, frontal, and supraorbital furrow not connected. Clypeus and frons with distinct isodiametric reticulation, clypeal wing and supraantennal plate with longitudinal reticulation. Neck at middle at same level as frons, constriction developed laterally as row of punctures. Eye moderately projecting laterally. Gena indistinct. Antenna reaching over posterior angle of pronotum, antennomeres four to ten elongate (around 1.8 times longer than wide). Labrum straight anteriorly, reticulated, seven setose.
</paragraph>
<paragraph id="895A375EFFE1BB3D31A97521FE6EFB59" blockId="11.[151,757,257,2077]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D31A97521FEF1FC80" box="[151,266,793,816]" italics="true" pageId="11" pageNumber="367">Pronotum:</emphasis>
Disk in lateral view moderately convex, convex in frontal view. Outline trapezoid. Reflexed lateral margin smooth, nearly straight attenuated in anterior half, widest at posterior third; anterior angle slightly projecting; posterior angle developed as distinct rounded tooth, tooth laterally much less projecting than widest part of lateral margin. Lateral channel as wide as reflexed lateral margin, anterior setigerous punctures removed from channel by diameter of pore, posterior setigerous puncture situated in the channel close to the tooth of the posterior angle. Median line moderately narrow; anterior transverse line slightly wider than median line, joining with median line. Surface with isodiametric to transverse reticulation; basal constriction distinct, flange 2.5 times wider than constriction.
</paragraph>
<paragraph id="895A375EFFE1BB3D31A972C9FDBBF8B1" blockId="11.[151,757,257,2077]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D31A972C9FF14FAB9" box="[151,239,1265,1289]" italics="true" pageId="11" pageNumber="367">Elytron:</emphasis>
Disk slightly convex in anterior half in lateral view, convex on frontal view. Around 1.6 times as long as wide, slightly dilated in apical half, maximum width slightly behind middle. Reflexed lateral margin smooth. Scutellar striole short but visible. Base of elytron with setigerous tubercle at base of second interval, with a small tubercle at base of third interval. Striae moderately deep, crenulate-striate, four to seven reaching base, the seventh at level of the humerus, humerus marked by distinct tooth. Stria one running to apex, six and seven joining apically, the others partly joining at apex (variable). Intervals moderately convex. Third interval with three setigerous punctures, first distinctly removed from base and approaching third stria, the second situated at middle of the interval, the third approaching second stria. Interval one to seven smooth, glossy, interval eight with indistinct longitudinal reticulation.
</paragraph>
<paragraph id="895A375EFFE1BB3D31A97130FE2FF890" blockId="11.[151,757,257,2077]" box="[151,468,1800,1824]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D31A97130FEEAF890" box="[151,273,1800,1824]" italics="true" pageId="11" pageNumber="367">Hind wings</emphasis>
: Fully developed.
</paragraph>
<paragraph id="895A375EFFE1BB3D31A97111FEE3F82E" blockId="11.[151,757,257,2077]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D31A97111FECAF8F0" box="[151,305,1832,1856]" italics="true" pageId="11" pageNumber="367">Lower surface</emphasis>
: Proepisternum covered with isodiametric reticulation and few transverse wrinkles. Sternites of abdomen transversely sulcate, with transverse reticulation.
</paragraph>
<paragraph id="895A375EFFE1BB3D31A9719FFEA8F7AD" blockId="11.[151,757,257,2077]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D31A9719FFF32F80E" box="[151,201,1959,1982]" italics="true" pageId="11" pageNumber="367">Legs</emphasis>
: Profemur dorsally sulcate. Protibia covered with longitudinal reticulation, with two slender spines of moderate length, the third one obtuse, movable spur regularly arcuate.
</paragraph>
<paragraph id="895A375EFFE1BB3D32017739FAACFE0A" blockId="11.[831,1437,257,860]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D32017739FC20FEA9" box="[831,987,257,281]" italics="true" pageId="11" pageNumber="367">Male genitalia</emphasis>
(
<figureCitation id="11DE2BDBFFE1BB3D32D67739FBCCFEA9" box="[1000,1079,257,281]" captionStart="Figs 14-20" captionStartId="9.[695,736,1848,1870]" captionTargetId="figure-270@9.[151,1431,769,2095]" captionTargetPageId="9" captionText="Figs 14-20. Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (14) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand. (15) S. acutipalpis (Putzeys, 1877), lectotype. (16) S. fulvaster (Andrewes, 1927), paratype. (17) S. papua Darlington, 1962, paratype.(18) S. sinepunctatus sinepunctatus sp. nov., holotype. (19) S. sinepunctatus kluangensis ssp. nov., holotype. (20) S. gradus sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.5646404" httpUri="https://zenodo.org/record/5646404/files/figure.png" pageId="11" pageNumber="367">Fig. 18</figureCitation>
): Median lobe regularly curved in basal half, gently curved towards apex, apically distinctly acute in ventral view, in lateral view widened at middle, developed apically as asymmetric distorted spatula, broadly rounded at tip of apex; endophallus with two longitudinal groups of bristles; parameres asetose.
</paragraph>
<paragraph id="895A375EFFE1BB3D327E77FAFB8BFD4B" blockId="11.[831,1437,257,860]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D327E77FAFC06FE6A" box="[832,1021,450,474]" italics="true" pageId="11" pageNumber="367">Female genitalia</emphasis>
(
<figureCitation id="11DE2BDBFFE1BB3D352C77FAFB90FE6A" box="[1042,1131,450,474]" captionStart="Figs 21-27" captionStartId="10.[151,192,1236,1258]" captionTargetBox="[189,1399,262,1192]" captionTargetId="figure-388@10.[151,1431,257,1223]" captionTargetPageId="10" captionText="Figs 21-27. Female coxostyli with laterotergite. (21) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand, with latero- and mediotergite. (22) S. acutipalpis (Putzeys, 1877), specimen from Lilong, China. (23) S. fulvaster (Andrewes, 1927), holotype. (24) S. papua Darlington, 1962, paratype. (25) S. sinepunctatus sinepunctatus sp. nov., paratype. (26) S. sinepunctatus kluangensis sp. nov., paratype. (27) S. gradus sp. nov. paratype." figureDoi="http://doi.org/10.5281/zenodo.5640144" httpUri="https://zenodo.org/record/5640144/files/figure.png" pageId="11" pageNumber="367">Fig. 25</figureCitation>
): Coxostylus one and two partly fused; coxostylus two gently curved at apex, ventromedially conspicuously narrowed and with conspicuously wide and long nematiform seta, apically with moderately large cone-like seta, with twelve long and one short nematiform setae, coxostylus one with two long nematiform apical setae, ventromedially with four moderately long setae, with two shorter setae at middle; laterotergite with five moderately long nematiform setae, one of them situated basally.
</paragraph>
<paragraph id="895A375EFFE1BB3D327E753CFB89FCEC" blockId="11.[831,1437,257,860]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D327E753CFC4AFCAC" box="[832,945,772,796]" italics="true" pageId="11" pageNumber="367">Variability</emphasis>
: The reticulation on interval eight is more or less distinct. At the apex of the elytron the striae two to five are more or less joining.
</paragraph>
</subSubSection>
<subSubSection id="C1FF64D5FFE1BB3D327E754CFB66FC1B" pageId="11" pageNumber="367" type="distribution">
<paragraph id="895A375EFFE1BB3D327E754CFB66FC1B" blockId="11.[832,1436,883,939]" pageId="11" pageNumber="367">
<emphasis id="BB91EB4CFFE1BB3D327E754CFC2DFC3C" bold="true" box="[832,982,884,908]" pageId="11" pageNumber="367">Distribution:</emphasis>
Known from two locations in Borneo (
<collectingCountry id="F1F277CEFFE1BB3D327975ABFC6AFC1B" box="[839,913,915,939]" name="Brunei" pageId="11" pageNumber="367">Brunei</collectingCountry>
and South East Borneo).
</paragraph>
</subSubSection>
</treatment>
</document>
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