496 lines
76 KiB
XML
496 lines
76 KiB
XML
<document id="5B06574BC66F1E6AEB716EDBF76012AD" ID-DOI="10.11646/zootaxa.3846.3.7" ID-GBIF-Dataset="8736c350-248a-459d-99a6-5ec8b3e33649" ID-ISSN="1175-5326" ID-Zenodo-Dep="226466" ID-ZooBank="E13CCEE1-662A-4CF1-9169-60EB6ED7C2F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460466975566" checkinUser="plazi" docAuthor="Bruce, A. J." docDate="2014" docId="5C1A3024FFF1FFE7ECBE9A66FC05EB06" docLanguage="en" docName="zt03846p446.pdf" docOrigin="Zootaxa 3846 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laomenes albonigrus Bruce, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="445" masterDocId="A023485CFFF0FFE1EC299B40FFD2E83E" masterDocTitle="Laomenes albonigrus sp. nov., (Crustacea: Decapoda; Pontoniinae) from Ashmore Reef, Western Australia" masterLastPageNumber="446" masterPageNumber="439" pageNumber="440" updateTime="1698305133603" updateUser="plazi">
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<mods:title id="22E8B150EFEA43F9D662836875773F53">Laomenes albonigrus sp. nov., (Crustacea: Decapoda; Pontoniinae) from Ashmore Reef, Western Australia</mods:title>
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<mods:name id="A4DAA87795D744938FCB5504CA875D94" type="personal">
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<mods:namePart id="AA9480D0DB9988730C0EECB8AB9ABBE2">Bruce, A. J.</mods:namePart>
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<mods:typeOfResource id="F5A351FD1E1B2FC33C4DCBBD1DBA4433">text</mods:typeOfResource>
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<mods:title id="F4D1CEDD11DE52DF4F6D2AB5F5CBED3E">Zootaxa</mods:title>
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<mods:part id="12DE3EB4D1F2BFEEE66EB86F8D6433BA">
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<mods:date id="98AA8F64AF27CF42AE22B45546A5F56F">2014</mods:date>
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<mods:detail id="DA485E790E91AE021FC7077927497706" type="volume">
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<mods:number id="41218B9BCA095924E9F727F0B893233D">3846</mods:number>
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<mods:detail id="FEC5EC6BE169DDEA10BCA8839DC82B70" type="issue">
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<mods:number id="67CCDDEC301D7EF11A2B22E84E685C25">3</mods:number>
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<mods:classification id="7BD6B8439BDB826DDAC9E7DD3DB2BDD9">journal article</mods:classification>
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<mods:identifier id="FCD3AF3D5103E20255FB6C9DBD15FCDD" type="DOI">10.11646/zootaxa.3846.3.7</mods:identifier>
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<mods:identifier id="AC7D54D680A10085DD90CC0D1A6AABD8" type="GBIF-Dataset">8736c350-248a-459d-99a6-5ec8b3e33649</mods:identifier>
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<treatment id="5C1A3024FFF1FFE7ECBE9A66FC05EB06" ID-DOI="http://doi.org/10.5281/zenodo.6125475" ID-GBIF-Taxon="119514404" ID-Zenodo-Dep="6125475" LSID="urn:lsid:plazi:treatment:5C1A3024FFF1FFE7ECBE9A66FC05EB06" httpUri="http://treatment.plazi.org/id/5C1A3024FFF1FFE7ECBE9A66FC05EB06" lastPageId="6" lastPageNumber="445" pageId="1" pageNumber="440">
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<subSubSection id="9CA9D2B9FFF1FFE0ECBE9A66FED9E95F" pageId="1" pageNumber="440" type="nomenclature">
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<paragraph id="D40C8132FFF1FFE0ECBE9A66FDD3E97E" blockId="1.[151,513,294,354]" box="[151,513,294,320]" pageId="1" pageNumber="440">
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<heading id="8F44365EFFF1FFE0ECBE9A66FDD3E97E" bold="true" box="[151,513,294,320]" fontSize="11" level="1" pageId="1" pageNumber="440" reason="1">
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<emphasis id="E6C75D20FFF1FFE0ECBE9A66FDD3E97E" bold="true" box="[151,513,294,320]" pageId="1" pageNumber="440">
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<taxonomicName id="13B3FAB1FFF1FFE0ECBE9A66FE4EE97E" box="[151,412,294,320]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="albonigrus" status="sp. nov.">
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<emphasis id="E6C75D20FFF1FFE0ECBE9A66FE4EE97E" bold="true" box="[151,412,294,320]" italics="true" pageId="1" pageNumber="440">Laomenes albonigrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDF4E05BFFF1FFE0ED8D9A66FDD3E97E" box="[420,513,294,320]" pageId="1" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECBE9A09FED9E95F" blockId="1.[151,513,294,354]" box="[151,267,329,354]" pageId="1" pageNumber="440">
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(
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<figureCitation id="4C889DB7FFF1FFE0ECB69A09FED0E95F" box="[159,258,329,354]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1477,1499]" captionStartId-1="3.[151,250,1976,1998]" captionStartId-2="4.[151,250,1683,1705]" captionStartId-3="5.[151,250,1079,1101]" captionTargetBox-0="[168,1420,271,1424]" captionTargetBox-1="[168,1426,850,1945]" captionTargetBox-2="[173,1436,212,1596]" captionTargetBox-3="[150,1435,193,1057]" captionTargetId-0="figure@2.[151,1436,265,1456]" captionTargetId-1="figure@3.[151,1436,841,1955]" captionTargetId-2="figure@4.[151,1436,193,1661]" captionTargetId-3="figure@5.[150,1436,193,1058]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." captionText-1="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." captionText-2="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." captionText-3="FIGURE 4. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. Photograph, Andrew Hosie." httpUri-0="https://zenodo.org/record/226467/files/figure.png" httpUri-1="https://zenodo.org/record/226468/files/figure.png" httpUri-2="https://zenodo.org/record/226469/files/figure.png" httpUri-3="https://zenodo.org/record/226470/files/figure.png" pageId="1" pageNumber="440">Figs 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA9D2B9FFF1FFE0ECBE9AD1FBD7E9F0" pageId="1" pageNumber="440" type="materials_examined">
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<paragraph id="D40C8132FFF1FFE0ECBE9AD1FBD7E9F0" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECBE9AD1FE53E994" bold="true" box="[151,385,401,426]" pageId="1" pageNumber="440">Material examined.</emphasis>
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1 ♀,
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<typeStatus id="0B083F90FFF1FFE0EDE89AD1FDF4E994" box="[449,550,401,426]" pageId="1" pageNumber="440" type="holotype">holotype</typeStatus>
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,
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<collectingCountry id="ACA4C1A2FFF1FFE0EE189AD1FD45E994" box="[561,663,401,426]" name="Ashmore and Cartier Islands" pageId="1" pageNumber="440">Ashmore</collectingCountry>
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Reef, Western
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<collectingCountry id="ACA4C1A2FFF1FFE0EF6C9AD1FC7DE994" box="[837,943,401,426]" name="Australia" pageId="1" pageNumber="440">Australia</collectingCountry>
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, north side, lagoon middle entrance,
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<geoCoordinate id="B187E7F5FFF1FFE0E9679AD1FF2DE9F0" direction="south" orientation="latitude" pageId="1" pageNumber="440" precision="1" value="-12.19448">12° 11' 40.128"S</geoCoordinate>
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<geoCoordinate id="B187E7F5FFF1FFE0ED2F9AF6FE7EE9F0" box="[262,428,437,462]" direction="east" orientation="longitude" pageId="1" pageNumber="440" precision="1" value="123.050156">123° 3' 0.54"E</geoCoordinate>
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,
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<quantity id="134B2CD7FFF1FFE0ED9E9AF6FE36E9F3" box="[439,484,438,461]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="440" unit="m" value="7.0">7 m</quantity>
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,
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<date id="A00DA7F2FFF1FFE0EDD99AF6FD73E9F0" box="[496,673,437,462]" pageId="1" pageNumber="440" value="2013-10-01">1 October 2013</date>
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, coll. M. Ekins,
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<collectionCode id="B2A219F7FFF1FFE0EF709AF6FC4CE9F0" LSID="urn:lsid:biocol.org:col:34981" box="[857,926,438,462]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="440">WAM</collectionCode>
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C55074.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA9D2B9FFF1FFE0ECEE9A99FB02EA60" pageId="1" pageNumber="440" type="diagnosis">
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<paragraph id="D40C8132FFF1FFE0ECEE9A99FB02EA60" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE9A99FE93E9CC" bold="true" box="[199,321,473,498]" pageId="1" pageNumber="440">Diagnosis.</emphasis>
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A typical species of the genus
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<taxonomicName id="13B3FAB1FFF1FFE0EEF39A9AFC38E9CC" authority="Clark, 1915" authorityName="Clark" authorityYear="1915" box="[730,1002,473,498]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="440" phylum="Arthropoda" rank="genus">
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<emphasis id="E6C75D20FFF1FFE0EEF39A9AFC9EE9CF" box="[730,844,474,497]" italics="true" pageId="1" pageNumber="440">Laomenes</emphasis>
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Clark, 1915
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</taxonomicName>
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, of the
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<taxonomicName id="13B3FAB1FFF1FFE0E87E9A99FB31E9CC" box="[1111,1251,473,498]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="amboinensis">
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<emphasis id="E6C75D20FFF1FFE0E87E9A99FB31E9CC" box="[1111,1251,473,498]" italics="true" pageId="1" pageNumber="440">amboinensis</emphasis>
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</taxonomicName>
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species group, characterised by its hemispherical cornea without a distal papilla, second pereiopods strongly unequal, dissimilar, and the third ambulatory dactyl with shorter erect accessory tooth and propod with longer distoventral spines and seven ventral spines, dorsal telson spines of normal length, projecting over lateral telson margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA9D2B9FFF1FFE4ECEE9929FC3AEEC9" lastPageId="5" lastPageNumber="444" pageId="1" pageNumber="440" type="description">
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<paragraph id="D40C8132FFF1FFE0ECEE9929FBA5EABC" blockId="1.[151,1437,401,2010]" box="[199,1143,617,642]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE9929FE8BEABC" bold="true" box="[199,345,617,642]" pageId="1" pageNumber="440">Description.</emphasis>
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Medium-sized shrimp with slender subcylindrical anterior body shape.
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECEE99CEFDD9EB2C" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE99CEFEF4EA9B" box="[199,294,654,677]" italics="true" pageId="1" pageNumber="440">Rostrum</emphasis>
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(
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<figureCitation id="4C889DB7FFF1FFE0ED1F99CDFEA4EA98" box="[310,374,653,678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
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B) deep, compressed, acute, reaching to end of antennular peduncle, with five well developed acute, anteroverted subequal dorsal teeth on distal half, proximal half convex, unarmed, interdental spaces sparsely setose proximally, ventral margin convex, with two acute teeth distally, lateral carinae distinct posteriorly, confluent with supraorbital teeth.
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECEE985EFE36EB9C" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE985EFEE7EB0B" box="[199,309,798,821]" italics="true" pageId="1" pageNumber="440">Carapace</emphasis>
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(
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<figureCitation id="4C889DB7FFF1FFE0ED61985DFE58EB08" box="[328,394,797,822]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
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A) about twice as long as deep in lateral view, compressed, with large supraorbital teeth, without epigastric tooth, well developed rounded inferior orbital angle, orbit well developed, deep, antennal tooth and hepatic tooth subequal (
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<figureCitation id="4C889DB7FFF1FFE0EDF39825FDCEEB40" box="[474,540,869,894]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
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C, 3A), antennal tooth slightly exceeding carapace margin, anterolateral angle anteriorly produced, rounded.
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECEE98EDFAF0EBD7" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE98EDFEE3EBF8" box="[199,305,941,966]" italics="true" pageId="1" pageNumber="440">Abdomen</emphasis>
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slender, pleura small, rounded, third segment not posterodorsally produced, sixth segment 1.6 times longer than fifth, 0.78 of telson length, with posterolateral angle and posteroventral angle similar, acute.
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECEE98B5FDD0ECA0" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE98B5FEDFEC30" box="[199,269,1013,1038]" italics="true" pageId="1" pageNumber="440">Telson</emphasis>
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(
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<figureCitation id="4C889DB7FFF1FFE0ED0998B5FEB2EC30" box="[288,352,1013,1038]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
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J) 0.6 of CL, 2.6 times longer than anterior width, tapering posteriorly, lateral margins straight, posterior margin broadly rounded (
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<figureCitation id="4C889DB7FFF1FFE0EE099F59FDB1EC0C" box="[544,611,1049,1074]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
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J), without median point, with two pairs of similar marginal dorsal spines at 0.46 and 0.73 of telson length, about 0.06 of telson length, overhanging lateral telson margin, lateral posterior spines as dorsal spines, intermediate spines 0.25 of telson length, robust, submedian spines slender, non-setulose, 0.5 of intermediate spine length.
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</paragraph>
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<paragraph id="D40C8132FFF1FFE0ECEE9FE9FB89ED6C" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
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<emphasis id="E6C75D20FFF1FFE0ECEE9FE9FEEBECFC" box="[199,313,1193,1218]" italics="true" pageId="1" pageNumber="440">Antennule</emphasis>
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(
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<figureCitation id="4C889DB7FFF1FFE0ED619FE9FE55ECFC" box="[328,391,1193,1218]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
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D) with peduncle slightly exceeding rostrum, proximal segment 1.5 times longer than broad, medial margin straight, setose, with large ventral tooth (
|
||
<figureCitation id="4C889DB7FFF1FFE0EF3C9F8DFC8AECD8" box="[789,856,1229,1254]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
|
||
B) at near half length, anterolateral region strongly produced, with distolateral tooth exceeding intermediate segment, with well developed lobe medially (
|
||
<figureCitation id="4C889DB7FFF1FFE0E91D9FB1FAABED34" box="[1332,1401,1265,1290]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
|
||
C), slightly angular distally, sparsely setose, stylocerite acute, reaching half proximal segment length, intermediate and distal segments of subequal length, 0.5 of proximal segment length, flagella damaged.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE0ECEE9E1EFC94EDDC" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9E1EFEF6ED4B" box="[199,292,1374,1397]" italics="true" pageId="1" pageNumber="440">Antenna</emphasis>
|
||
(
|
||
<figureCitation id="4C889DB7FFF1FFE0ED1D9E1DFEA1ED48" box="[308,371,1373,1398]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
|
||
E) with robust basicerite with stout distolateral tooth, carpocerite subcylindrical, twice as long as wide, reaching to half scaphocerite length, merocerite and ischiocerite normal, scaphocerite exceeding antennular peduncle, 2.4 times longer than broad, lateral margin straight, with acute distolateral tooth at 0.8 of length, lamella distally rounded, well exceeding lateral tooth.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE0ECEE9EAEFE9EEE4C" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9EAEFF23EE3B" box="[199,241,1518,1541]" italics="true" pageId="1" pageNumber="440">Eye</emphasis>
|
||
(
|
||
<figureCitation id="4C889DB7FFF1FFE0ED299EADFEEDEE38" box="[256,319,1517,1542]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
|
||
F) with globular cornea, slightly oblique, without terminal papilla, diameter about 0.22 of CL, well pigmented, with posterolateral pigment spot, reaching to level of second rostral tooth, stalk stout, subcylindrical, 1.5 times longer than broad. Ophthalmic somite unarmed, with low median carina anteriorly, median black pigment spot posteriorly.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE0ECEE9D3EFB95EEA8" blockId="1.[151,1437,401,2010]" box="[199,1095,1661,1686]" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9D3EFEFDEEAB" box="[199,303,1662,1685]" italics="true" pageId="1" pageNumber="440">Epistome</emphasis>
|
||
with well developed acute convergent submedian processes (
|
||
<figureCitation id="4C889DB7FFF1FFE0EFC99D3DFBF0EEA8" box="[992,1058,1661,1686]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE0ECEE9DE1FD5BEF3C" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9DE1FEE1EE84" box="[199,307,1697,1722]" italics="true" pageId="1" pageNumber="440">Mandible</emphasis>
|
||
with molar process robust, subcylindrical, with two large ventral teeth (
|
||
<figureCitation id="4C889DB7FFF1FFE0E8569DE1FB16EE84" box="[1151,1220,1697,1722]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
|
||
E); incisor process (
|
||
<figureCitation id="4C889DB7FFF1FFE0ECB69D85FF32EEE0" box="[159,224,1733,1758]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="1" pageNumber="440">Fig. 3</figureCitation>
|
||
F) distally expanded, oblique distally with large feebly bilobed medial tooth and eight smaller similar simple acute teeth. Other mouthparts not examined.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE0ECEE9C4DFD37EF74" blockId="1.[151,1437,401,2010]" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9C4DFE44EF1B" box="[199,406,1805,1830]" italics="true" pageId="1" pageNumber="440">Thoracic sternites</emphasis>
|
||
narrow, fifth with transverse carina with well marked median notch posterior to second pereiopod attachments, posterior segments unarmed.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF1FFE3ECEE9C16FDDBE8EB" blockId="1.[151,1437,401,2010]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440">
|
||
<emphasis id="E6C75D20FFF1FFE0ECEE9C16FEA4EF50" box="[199,374,1877,1902]" italics="true" pageId="1" pageNumber="440">First pereiopod</emphasis>
|
||
(
|
||
<figureCitation id="4C889DB7FFF1FFE0EDAE9C15FE14EF50" box="[391,454,1877,1902]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
|
||
G) exceeding scaphocerite by chela; chela (
|
||
<figureCitation id="4C889DB7FFF1FFE0EF939C15FC28EF50" box="[954,1018,1877,1902]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="1" pageNumber="440">Fig. 1</figureCitation>
|
||
H) slender, about 0.33 of CL, feebly bowed, palm oval in section, about 2.5 times longer than deep, with several short rows of cleaning setae proximally, fingers slender, tapering to fine simple hooked tips, about 0.75 of palm length, cutting edges entire; carpus 1.6 times chela length, 7.5 times longer than distal width, tapering proximally, with few cleaning setae distoventrally; merus 1.45 times chela length, 8.0 times longer than central depth, ischium and basis normal; coxa with strong setose lobe ventrally.
|
||
</paragraph>
|
||
<caption id="80CCD1BAFFF2FFE3ECBE9E85FE22EE26" httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="2" pageNumber="441" targetBox="[168,1420,271,1424]" targetPageId="2">
|
||
<paragraph id="D40C8132FFF2FFE3ECBE9E85FE22EE26" blockId="2.[151,1436,1477,1560]" pageId="2" pageNumber="441">
|
||
<emphasis id="E6C75D20FFF2FFE3ECBE9E85FECBEDE4" bold="true" box="[151,281,1477,1499]" pageId="2" pageNumber="441">FIGURE 1.</emphasis>
|
||
<taxonomicName id="13B3FAB1FFF2FFE3ED0C9E86FDD4EDE5" box="[293,518,1477,1499]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="albonigrus" status="sp. nov.">
|
||
<emphasis id="E6C75D20FFF2FFE3ED0C9E86FDD4EDE5" box="[293,518,1477,1499]" italics="true" pageId="2" pageNumber="441">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E6C75D20FFF2FFE3EE3B9E86FDBBEDE5" bold="true" box="[530,617,1478,1499]" pageId="2" pageNumber="441">
|
||
<taxonomicNameLabel id="FDF4E05BFFF2FFE3EE3B9E86FDBBEDE5" box="[530,617,1478,1499]" pageId="2" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, Ashmore Reef, WAM C55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D40C8132FFF2FFE2ECEE9D00FE8EE95B" blockId="2.[151,1437,1600,2021]" lastBlockId="3.[151,1437,151,789]" lastPageId="3" lastPageNumber="442" pageId="2" pageNumber="441">
|
||
<emphasis id="E6C75D20FFF2FFE3ECEE9D00FE4EEE67" box="[199,412,1600,1625]" italics="true" pageId="2" pageNumber="441">Second pereiopods</emphasis>
|
||
well developed, unequal, dissimilar. Major second pereiopod (
|
||
<figureCitation id="4C889DB7FFF2FFE3E84C9D00FB7AEE66" box="[1125,1192,1600,1625]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
|
||
A) robust, chela (
|
||
<figureCitation id="4C889DB7FFF2FFE3E9469D00FF74EE42" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
|
||
B) about 1.2 of CL, palm smooth, glabrous, 0.85 of CL, oval in section, 2.6 times longer than central depth, tapering proximally and distally, fingers (
|
||
<figureCitation id="4C889DB7FFF2FFE3EE529DC8FD10EE9E" box="[635,706,1672,1697]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
|
||
C) about 0.3 of palm length, incurved; dactyl robust, medially subspatulate, 3.0 times longer than proximal depth, strongly curved, acute, cutting edge distally entire, without distinct cutting edge, central third laminar with larger distal tooth, three smaller proximal teeth, proximal third with flatted plaque, sparsely setose, with strongly hooked tip occluding into deep preterminal notch on fixed finger; fixed finger 3.5 times longer than central depth, raised, curved, carinate, with broad low distal tooth, separated by deep notch from feebly curved tip, proximal cutting edge with three low subacute teeth; carpus short, 0.2 of palm length, distally excavate, distal width subequal to length, unarmed; merus 0.45 of palm length, 3.5 times longer than central depth, straight, tapering proximally, with large acute distoventral tooth, ventral margin unarmed; ischium 0.25 of palm length, 2.4 times longer than distal width, tapering proximally, ventral margin unarmed; basis and coxa normal. Minor second pereiopod (
|
||
<figureCitation id="4C889DB7FFF2FFE3EE559C8CFD6CEFDA" box="[636,702,1996,2021]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
|
||
D) slender, chela (
|
||
<figureCitation id="4C889DB7FFF2FFE3EFA09C8CFC18EFDA" box="[905,970,1996,2021]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
|
||
E) about 0.72 of major palm length, about 0.76 of CL, palm smooth, glabrous, 0.45 of CL, oval in section, 2.7 times longer than central depth, subcylindrical, dactyl (
|
||
<figureCitation id="4C889DB7FFF3FFE2ECC59BFCFEFDE8EA" box="[236,303,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="3" pageNumber="442">Fig. 2</figureCitation>
|
||
F) 0.7 of palm length, feebly curved, tip hooked, cutting edge feebly concave with three small acute teeth on central third; fixed finger (
|
||
<figureCitation id="4C889DB7FFF3FFE2EE1E9B9FFDACE8C6" box="[567,638,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[168,1426,850,1945]" captionTargetId="figure@3.[151,1436,841,1955]" captionTargetPageId="3" captionText="FIGURE 2. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers." httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="3" pageNumber="442">Fig. 2</figureCitation>
|
||
F) straight, 4.0 times longer than proximal depth with stout strongly hooked tip separated by deep notch from distal cutting edge, cutting edge proximally blunt with two small acute teeth, distal third with larger proximal tooth, distal cutting edge sharp; carpus and proximal segments as in major chela but smaller.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF3FFE2ECEE9A2FFC9CEA5E" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="442">
|
||
<emphasis id="E6C75D20FFF3FFE2ECEE9A2FFE1FE9B6" box="[199,461,367,392]" italics="true" pageId="3" pageNumber="442">Ambulatory pereiopods</emphasis>
|
||
slender, third pereiopod (
|
||
<figureCitation id="4C889DB7FFF3FFE2EEC59A2FFCFEE9B6" box="[748,812,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1477,1499]" captionTargetBox="[168,1420,271,1424]" captionTargetId="figure@2.[151,1436,265,1456]" captionTargetPageId="2" captionText="FIGURE 1. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, anterior carapace and appendages. B, rostrum. C, carapace, antennal region. D, antennule. E, antenna. F, eye, lateral. G, first pereiopod. H, same, chela. I, third pereiopod. J, telson." httpUri="https://zenodo.org/record/226467/files/figure.png" pageId="3" pageNumber="442">
|
||
Fig.
|
||
<date id="A00DA7F2FFF3FFE2EF089A30FCFEE9B6" box="[801,812,368,392]" pageId="3" pageNumber="442">1</date>
|
||
</figureCitation>
|
||
I) with dactyl (
|
||
<figureCitation id="4C889DB7FFF3FFE2EFFF9A2FFBC5E9B6" box="[982,1047,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="3" pageNumber="442">Fig. 3</figureCitation>
|
||
H) short, compressed, about 0.2 of propod length, unguis distinct from corpus, dorsal margin about 2.5 times distal propod depth, unguial portion about 0.6 of dorsal margin length, ventral margin of corpus straight with small slender straight erect acute distal accessory tooth, about 0.5 of unguis length, proximal margin straight, unarmed; propod (
|
||
<figureCitation id="4C889DB7FFF3FFE2E8BA9A9CFB04E9CA" box="[1171,1238,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="3" pageNumber="442">Fig. 3</figureCitation>
|
||
G) slender, about 0.27 of CL, 4.5 times longer than wide, with pair of subequal similar simple slender distoventral spines, longer spine 0.45 of propod width, with seven well spaced small spines along ventral margin, several long simple distolateral setae; proximal segments without special features.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF3FFE2ECEE992CFAD8EA96" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="442">
|
||
<emphasis id="E6C75D20FFF3FFE2ECEE992CFEFDEABB" box="[199,303,620,645]" italics="true" pageId="3" pageNumber="442">Pleopods</emphasis>
|
||
of normal format. Second pleopod (
|
||
<figureCitation id="4C889DB7FFF3FFE2EEF6992CFCF7EABA" box="[735,805,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="3" pageNumber="442">
|
||
Fig.
|
||
<date id="A00DA7F2FFF3FFE2EF31992CFCF7EABA" box="[792,805,620,644]" pageId="3" pageNumber="442">3</date>
|
||
</figureCitation>
|
||
I) with endopod subequal to exopod length, 5.0 times longer than wide, with appendix interna 0.4 of exopod length attached at 0.3 of medial margin length.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF3FFE2ECEE99F4FBDFEB2B" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="442">
|
||
<emphasis id="E6C75D20FFF3FFE2ECEE99F4FEFAEAF3" box="[199,296,692,717]" italics="true" pageId="3" pageNumber="442">Uropods</emphasis>
|
||
with rami exceeding telson, protopod posterolaterally unarmed non-setose; exopod 3.3 times longer than broad, lateral margin straight, non-setose, posterolateral angles with small acute tooth with large fossa medially (
|
||
<figureCitation id="4C889DB7FFF3FFE2ED2199BCFE98EB2A" box="[264,330,764,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1683,1705]" captionTargetBox="[173,1436,212,1596]" captionTargetId="figure@4.[151,1436,193,1661]" captionTargetPageId="4" captionText="FIGURE 3. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod." httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="3" pageNumber="442">Fig. 3</figureCitation>
|
||
K), both distolateral spines lost; endopod 0.9 of exopod length.
|
||
</paragraph>
|
||
<caption id="80CCD1BAFFF3FFE2ECBE9CF8FBE2EFD3" httpUri="https://zenodo.org/record/226468/files/figure.png" pageId="3" pageNumber="442" targetBox="[168,1426,850,1945]" targetPageId="3">
|
||
<paragraph id="D40C8132FFF3FFE2ECBE9CF8FBE2EFD3" blockId="3.[151,1435,1976,2029]" pageId="3" pageNumber="442">
|
||
<emphasis id="E6C75D20FFF3FFE2ECBE9CF8FEC6EFF3" bold="true" box="[151,276,1976,1998]" pageId="3" pageNumber="442">FIGURE 2.</emphasis>
|
||
<taxonomicName id="13B3FAB1FFF3FFE2ED359CF9FE2BEFF0" box="[284,505,1976,1998]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="albonigrus" status="sp. nov.">
|
||
<emphasis id="E6C75D20FFF3FFE2ED359CF9FE2BEFF0" box="[284,505,1976,1998]" italics="true" pageId="3" pageNumber="442">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E6C75D20FFF3FFE2EE289CF8FD81EFF3" bold="true" box="[513,595,1976,1997]" pageId="3" pageNumber="442">
|
||
<taxonomicNameLabel id="FDF4E05BFFF3FFE2EE289CF8FD81EFF3" box="[513,595,1976,1997]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, Ashmore Reef, WAM C55074. A, major second pereiopod. B, same, chela. C, same, fingers. D, minor second pereiopod. E, same, chela. F, same, fingers.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="80CCD1BAFFF4FFE5ECBE9DD3FBA7EF3A" httpUri="https://zenodo.org/record/226469/files/figure.png" pageId="4" pageNumber="443" targetBox="[173,1436,212,1596]" targetPageId="4">
|
||
<paragraph id="D40C8132FFF4FFE5ECBE9DD3FBA7EF3A" blockId="4.[151,1436,1683,1796]" pageId="4" pageNumber="443">
|
||
<emphasis id="E6C75D20FFF4FFE5ECBE9DD3FEC7EE96" bold="true" box="[151,277,1683,1705]" pageId="4" pageNumber="443">FIGURE 3.</emphasis>
|
||
<taxonomicName id="13B3FAB1FFF4FFE5ED379DD4FE29EE97" box="[286,507,1683,1705]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="albonigrus" status="sp. nov.">
|
||
<emphasis id="E6C75D20FFF4FFE5ED379DD4FE29EE97" box="[286,507,1683,1705]" italics="true" pageId="4" pageNumber="443">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E6C75D20FFF4FFE5EE2D9DD3FD84EE96" bold="true" box="[516,598,1683,1704]" pageId="4" pageNumber="443">
|
||
<taxonomicNameLabel id="FDF4E05BFFF4FFE5EE2D9DD3FD84EE96" box="[516,598,1683,1704]" pageId="4" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, Ashmore Reef, WAM C55074. A, antennal and hepatic teeth, dorsal. B, proximal segment of antennular peduncle and epistomal horn, medial. C, proximal antennular segment, distolateral angle. D, epistome. E, mandible, molar process. F, same, incisor process. G, third pereiopod, propod and dactyl. H, same, distal propod and dactyl. I, second pleopod. J, posterior telson spines. K, uropod, posterolateral exopod.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D40C8132FFF4FFE5ECEE9C71FE57EF88" blockId="4.[151,1436,1841,2010]" pageId="4" pageNumber="443">
|
||
<emphasis id="E6C75D20FFF4FFE5ECEE9C71FE07EF74" bold="true" box="[199,469,1841,1866]" pageId="4" pageNumber="443">Measurements (mms).</emphasis>
|
||
<typeStatus id="0B083F90FFF4FFE5EDC89C71FD9BEF74" box="[481,585,1841,1866]" pageId="4" pageNumber="443" type="holotype">Holotype</typeStatus>
|
||
female, rostrum and carapace, 4.3; postorbital carapace, 3.7; total body length (approx.), 11.0; major second pereiopod chela, 4.8; minor second pereiopod chela, 3.0.
|
||
<emphasis id="E6C75D20FFF4FFE5ECEE9C39FE9CEFAC" bold="true" box="[199,334,1913,1938]" pageId="4" pageNumber="443">Etymology.</emphasis>
|
||
Named after the black and white colouration of the live animal. From
|
||
<emphasis id="E6C75D20FFF4FFE5E8A89C39FB6CEFAC" box="[1153,1214,1913,1938]" italics="true" pageId="4" pageNumber="443">albus</emphasis>
|
||
(Latin), white, and
|
||
<emphasis id="E6C75D20FFF4FFE5ECBE9CDEFF0DEF8B" box="[151,223,1950,1973]" italics="true" pageId="4" pageNumber="443">nigrus</emphasis>
|
||
(Latin), black.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF4FFE5ECEE9C81FC2BEFE4" blockId="4.[151,1436,1841,2010]" box="[199,1017,1985,2010]" pageId="4" pageNumber="443">
|
||
<emphasis id="E6C75D20FFF4FFE5ECEE9C81FED4EFE4" bold="true" box="[199,262,1985,2010]" pageId="4" pageNumber="443">Host.</emphasis>
|
||
<taxonomicName id="13B3FAB1FFF4FFE5ED249C81FC82EFE4" authority="Muller, 1841 (Crinoidacea)" authorityName="Muller, 1841 (Crinoidacea" box="[269,848,1985,2010]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="4" pageNumber="443" phylum="Echinodermata" rank="species" species="bennetti">
|
||
<emphasis id="E6C75D20FFF4FFE5ED249C81FDC6EFE4" box="[269,532,1985,2010]" italics="true" pageId="4" pageNumber="443">Oxycomanthus bennetti</emphasis>
|
||
Müller, 1841 (Crinoidacea)
|
||
</taxonomicName>
|
||
, det. L. Marsh.
|
||
</paragraph>
|
||
<caption id="80CCD1BAFFF5FFE4ECBE9F77FAB6EC73" box="[151,1380,1079,1102]" httpUri="https://zenodo.org/record/226470/files/figure.png" pageId="5" pageNumber="444" targetBox="[150,1435,193,1057]" targetPageId="5">
|
||
<paragraph id="D40C8132FFF5FFE4ECBE9F77FAB6EC73" blockId="5.[151,1380,1079,1102]" box="[151,1380,1079,1102]" pageId="5" pageNumber="444">
|
||
<emphasis id="E6C75D20FFF5FFE4ECBE9F77FEC0EC72" bold="true" box="[151,274,1079,1101]" pageId="5" pageNumber="444">FIGURE 4.</emphasis>
|
||
<taxonomicName id="13B3FAB1FFF5FFE4ED309F79FE26EC73" box="[281,500,1079,1102]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="albonigrus" status="sp. nov.">
|
||
<emphasis id="E6C75D20FFF5FFE4ED309F79FE26EC73" box="[281,500,1079,1102]" italics="true" pageId="5" pageNumber="444">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E6C75D20FFF5FFE4EDD39F78FD98EC73" bold="true" box="[506,586,1080,1101]" pageId="5" pageNumber="444">
|
||
<taxonomicNameLabel id="FDF4E05BFFF5FFE4EDD39F78FD98EC73" box="[506,586,1080,1101]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, Ashmore Reef, WAM C55074. Photograph, Andrew Hosie.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D40C8132FFF5FFE4ECEE9F3AFF05ED1D" blockId="5.[151,1437,1146,2035]" pageId="5" pageNumber="444">
|
||
<emphasis id="E6C75D20FFF5FFE4ECEE9F3AFEBCECAD" bold="true" box="[199,366,1146,1171]" pageId="5" pageNumber="444">Colouration. (</emphasis>
|
||
From colour photograph,
|
||
<figureCitation id="4C889DB7FFF5FFE4EEA49F3AFD18ECAD" box="[653,714,1146,1171]" captionStart="FIGURE 4" captionStartId="5.[151,250,1079,1101]" captionTargetBox="[150,1435,193,1057]" captionTargetId="figure@5.[150,1436,193,1058]" captionTargetPageId="5" captionText="FIGURE 4. Laomenes albonigrus sp. nov., female holotype, Ashmore Reef, WAM C 55074. Photograph, Andrew Hosie." httpUri="https://zenodo.org/record/226470/files/figure.png" pageId="5" pageNumber="444">Fig 4</figureCitation>
|
||
). Generally black and white. Body and appendages white except for proximal halves of antennae, antero-dorsal carapace, ventral branchiostegite, most of first to fourth abdominal segments except posterior part of each pleuron, antero-ventral half of fifth segment, second pereiopods with distal carpus and proximal chela, bases of third to fifth pereiopods and pleopods. Second pereiopod chelae with golden tinge.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF5FFE4ECEE9E6EFCF1ED54" blockId="5.[151,1437,1146,2035]" pageId="5" pageNumber="444">
|
||
<emphasis id="E6C75D20FFF5FFE4ECEE9E6EFE6BED79" bold="true" box="[199,441,1326,1351]" pageId="5" pageNumber="444">Systematic position.</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF5FFE4EDED9E6EFCE5ED79" box="[452,823,1326,1351]" pageId="5" pageNumber="444" refString="Marin, I., Chan, T. - Y. & Okuno, J. (2012) Crinoid associated pontoniine shrimps of the genus Laomenes Clark, 1919 (Decapoda, Caridea, Palaemonidae) from Panglao 2004 and Kumejima 2009 Expeditions, with descriptions of two new species. Zootaxa, 3367, 103 - 114, figs. 1 - 8." type="journal article">Marin, Chan and Okuno (2012)</bibRefCitation>
|
||
noted that the only species of
|
||
<taxonomicName id="13B3FAB1FFF5FFE4E8839E6FFAC9ED78" box="[1194,1307,1327,1350]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6C75D20FFF5FFE4E8839E6FFAC9ED78" box="[1194,1307,1327,1350]" italics="true" pageId="5" pageNumber="444">Laomenes</emphasis>
|
||
</taxonomicName>
|
||
that had a cornea lacking a terminal papilla was
|
||
<taxonomicName id="13B3FAB1FFF5FFE4EE169E13FCCEED55" box="[575,796,1362,1387]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="gymnophthalmus">
|
||
<emphasis id="E6C75D20FFF5FFE4EE169E13FCCEED55" box="[575,796,1362,1387]" italics="true" pageId="5" pageNumber="444">L. gymnophthalmus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF5FFE4ECEE9E37FE12ED8D" blockId="5.[151,1437,1146,2035]" pageId="5" pageNumber="444">
|
||
<taxonomicName id="13B3FAB1FFF5FFE4ECEE9E37FE6BEDB1" box="[199,441,1398,1423]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="albonigrus">
|
||
<emphasis id="E6C75D20FFF5FFE4ECEE9E37FE6BEDB1" box="[199,441,1398,1423]" italics="true" pageId="5" pageNumber="444">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from
|
||
<taxonomicName id="13B3FAB1FFF5FFE4EEDF9E37FC01EDB1" box="[758,979,1398,1423]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="gymnophthalmus">
|
||
<emphasis id="E6C75D20FFF5FFE4EEDF9E37FC01EDB1" box="[758,979,1398,1423]" italics="true" pageId="5" pageNumber="444">L. gymnophthalmus</emphasis>
|
||
</taxonomicName>
|
||
as indicated below (
|
||
<taxonomicName id="13B3FAB1FFF5FFE4E8979E37FA49EDB1" box="[1214,1435,1398,1423]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="gymnophthalmus">
|
||
<emphasis id="E6C75D20FFF5FFE4E8979E37FA49EDB1" box="[1214,1435,1398,1423]" italics="true" pageId="5" pageNumber="444">L. gymnophthalmus</emphasis>
|
||
</taxonomicName>
|
||
characters in parentheses):
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF5FFE4ECEE9EFEFC3AEEC9" blockId="5.[151,1437,1146,2035]" pageId="5" pageNumber="444">
|
||
Rostral dentition 5/2 (
|
||
<emphasis id="E6C75D20FFF5FFE4ED979EFFFE07EDE8" box="[446,469,1471,1494]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
5/1, with additional minute preterminal denticle), antennal and hepatic teeth subequal (
|
||
<emphasis id="E6C75D20FFF5FFE4EC899EA3FF7EEDC4" box="[160,172,1507,1530]" italics="true" pageId="5" pageNumber="444">v</emphasis>
|
||
s hepatic tooth larger than antennal), second pereiopods unequal, minor chela half length of major chela (
|
||
<emphasis id="E6C75D20FFF5FFE4E9AD9EA3FA49EDC4" box="[1412,1435,1507,1530]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
subequal), distoventral merus acutely produced (
|
||
<emphasis id="E6C75D20FFF5FFE4EEFA9D47FD38EE20" box="[723,746,1543,1566]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
bluntly produced), major chela with fingers 0.33 of palm length, strongly dentate, (
|
||
<emphasis id="E6C75D20FFF5FFE4ED919D6BFE1DEE7C" box="[440,463,1579,1602]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
about 0.66 of palm length, feebly dentate), minor chela fingers 0.7 of palm length (
|
||
<emphasis id="E6C75D20FFF5FFE4E9AD9D6BFA49EE7C" box="[1412,1435,1579,1602]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
about 0.66 of palm length, feebly dentate), third pereiopod dactylar unguis about 0.7 of corpus length (
|
||
<emphasis id="E6C75D20FFF5FFE4E9139D0FFA83EE58" box="[1338,1361,1615,1638]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
0.59), accessory tooth 0.1 of unguis length (
|
||
<emphasis id="E6C75D20FFF5FFE4EE139D33FD83EEB4" box="[570,593,1651,1674]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
0.18), ventral corpus straight (
|
||
<emphasis id="E6C75D20FFF5FFE4EF809D33FC12EEB4" box="[937,960,1651,1674]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
sinuous), propod with pair of long slender distoventral spines (
|
||
<emphasis id="E6C75D20FFF5FFE4ED529DD7FE40EE90" box="[379,402,1687,1710]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
short spines), ventral margin with 7 small spines along whole length (
|
||
<emphasis id="E6C75D20FFF5FFE4E89C9DD7FB1EEE90" box="[1205,1228,1687,1710]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
2 distal spinules), telson with two pairs of well developed dorsal spines, 0.06 of telson length, projecting over lateral margin of telson (
|
||
<emphasis id="E6C75D20FFF5FFE4EC899D9FFF65EEC8" box="[160,183,1759,1782]" italics="true" pageId="5" pageNumber="444">vs</emphasis>
|
||
minute dorsal spines, 0.015 of telson length, not projecting over margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA9D2B9FFF5FFE4ECEE9C42FD89EF95" pageId="5" pageNumber="444" type="discussion">
|
||
<paragraph id="D40C8132FFF5FFE4ECEE9C42FD89EF95" blockId="5.[151,1437,1146,2035]" pageId="5" pageNumber="444">
|
||
<emphasis id="E6C75D20FFF5FFE4ECEE9C42FEE9EF25" bold="true" box="[199,315,1794,1819]" pageId="5" pageNumber="444">Remarks.</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF5FFE4ED6B9C42FE0AEF25" author="Marin" box="[322,472,1794,1819]" pageId="5" pageNumber="444" refString="Marin, I. (2009) Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity. Zootaxa, 1971, 1 - 49, figs. 1 - 36." type="journal article" year="2009">Marin (2009)</bibRefCitation>
|
||
divided the then known species of
|
||
<taxonomicName id="13B3FAB1FFF5FFE4EF4F9C43FC05EF24" box="[870,983,1795,1818]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6C75D20FFF5FFE4EF4F9C43FC05EF24" box="[870,983,1795,1818]" italics="true" pageId="5" pageNumber="444">Laomenes</emphasis>
|
||
</taxonomicName>
|
||
into three species groups centered on
|
||
<emphasis id="E6C75D20FFF5FFE4E9AC9C43FE02EF01" italics="true" pageId="5" pageNumber="444">
|
||
<taxonomicName id="13B3FAB1FFF5FFE4E9AC9C43FEF4EF01" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="amboinensis">L. amboinensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="13B3FAB1FFF5FFE4ED1A9C67FE02EF01" box="[307,464,1830,1855]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="jackhintoni">L. jackhintoni</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="E6C75D20FFF5FFE4EE239C67FD3CEF01" box="[522,750,1830,1855]" italics="true" pageId="5" pageNumber="444">
|
||
<taxonomicName id="13B3FAB1FFF5FFE4EE239C67FD38EF01" box="[522,746,1830,1855]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="ceratophthalmus">L. ceratophthalmus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The taxonomically important characters in these groups are presence or absence of ventral rostral teeth, hemispherical or papillate cornea, relative development of third pereiopod dactylar accessory tooth, presence or absence of distolateral spines on third pereiopod propod, presence or absence of tooth on uropodal exopod.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA9D2B9FFF5FFE7ECBE9CF6FC05EB06" lastPageId="6" lastPageNumber="445" pageId="5" pageNumber="444" type="reference_group">
|
||
<paragraph id="D40C8132FFF5FFE7ECBE9CF6FD7CE8C6" blockId="5.[151,1437,1146,2035]" lastBlockId="6.[151,1437,151,824]" lastPageId="6" lastPageNumber="445" pageId="5" pageNumber="444">
|
||
The
|
||
<taxonomicName id="13B3FAB1FFF5FFE4ECE69CF7FEADEFF1" ID-CoL="0d88ed20-0800-4b1d-8595-f5f5f0d45f4e" box="[207,383,1974,1999]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="amboinensis">
|
||
<emphasis id="E6C75D20FFF5FFE4ECE69CF7FEADEFF1" box="[207,383,1974,1999]" italics="true" pageId="5" pageNumber="444">L. amboinensis</emphasis>
|
||
</taxonomicName>
|
||
group is distinguished by the presence of ventral rostral teeth, cornea hemispherical or papillate, distoventral spines on third pereiopod propod, and a fixed tooth on distolateral margin of uropodal exopod. Included species are
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EDC69BD9FCB1E88E" ID-CoL="0d88ed20-0800-4b1d-8595-f5f5f0d45f4e" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[495,867,151,176]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="amboinensis">
|
||
<emphasis id="E6C75D20FFF6FFE7EDC69BD9FD4EE88E" box="[495,668,151,176]" italics="true" pageId="6" pageNumber="445">L. amboinensis</emphasis>
|
||
(
|
||
<collectingCountry id="ACA4C1A2FFF6FFE7EE879BD8FD02E88E" box="[686,720,152,176]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="445">De</collectingCountry>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7EEF29BD8FC88E88E" author="Man" box="[731,858,152,176]" pageId="6" pageNumber="445" refString="Man, J. G. de (1888) Bericht uber die von Herrn Dr. J. Brock im Indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschiche, 53, 215 - 600, figs 1 - 22 a." type="journal article" year="1888">
|
||
<collectingCountry id="ACA4C1A2FFF6FFE7EEF29BD8FCC3E891" box="[731,785,152,175]" name="Isle of Man" pageId="6" pageNumber="445">Man</collectingCountry>
|
||
, 1888
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EF5A9BD9FB06E88E" ID-CoL="ecad49c8-56ed-421e-907a-e75efd09f7b1" authority="Borradaile, 1915" authorityName="Borradaile" authorityYear="1915" box="[883,1236,151,176]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="E6C75D20FFF6FFE7EF5A9BD9FC24E88E" box="[883,1014,153,176]" italics="true" pageId="6" pageNumber="445">L. cornutus</emphasis>
|
||
(
|
||
<bibRefCitation id="B022FCC3FFF6FFE7E8219BD7FB1EE88E" author="Borradaile" box="[1032,1228,151,176]" pageId="6" pageNumber="445" refString="Borradaile, L. A. (1915) Notes on Carides. The Annals and Magazine of Natural History, Series 8, 15, 205 - 213. http: // dx. doi. org / 10.1080 / 00222931508693629" type="journal article" year="1915">Borradaile, 1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13B3FAB1FFF6FFE7E8CD9BD9FF00E8EA" ID-CoL="ff7ab485-870b-43eb-8f16-2d465316db28" authority="Marin, 2009" authorityName="Marin" authorityYear="2009" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="clarki">
|
||
<emphasis id="E6C75D20FFF6FFE7E8CD9BD9FA94E88E" box="[1252,1350,151,176]" italics="true" pageId="6" pageNumber="445">L. clarki</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7E9799BD7FF00E8EA" author="Marin" pageId="6" pageNumber="445" refString="Marin, I. (2009) Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity. Zootaxa, 1971, 1 - 49, figs. 1 - 36." type="journal article" year="2009">Marin, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECF49BFDFE0DE8EA" ID-CoL="0fa03b9d-5947-4c45-ad52-2c6786ac13b2" authority="Marin, 2009" authorityName="Marin" authorityYear="2009" box="[221,479,188,213]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="pardus">
|
||
<emphasis id="E6C75D20FFF6FFE7ECF49BFDFE98E8EB" box="[221,330,188,213]" italics="true" pageId="6" pageNumber="445">L. pardus</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7ED789BFCFE0DE8EA" author="Marin" box="[337,479,188,213]" pageId="6" pageNumber="445" refString="Marin, I. (2009) Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity. Zootaxa, 1971, 1 - 49, figs. 1 - 36." type="journal article" year="2009">Marin, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E6C75D20FFF6FFE7EDC29BFDFDB9E8EB" box="[491,619,188,213]" italics="true" pageId="6" pageNumber="445">
|
||
L,
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EE209BFCFDB9E8EB" ID-CoL="9240a32d-da9a-442b-95bb-885a3b8e2b8c" authority="Marin & Okuni, 2010" authorityName="Marin & Okuni" authorityYear="2010" box="[521,619,188,213]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="holthuisi">holthuisi</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<bibRefCitation id="B022FCC3FFF6FFE7EE539BFCFCA1E8EA" author="Marin" box="[634,883,188,213]" pageId="6" pageNumber="445" refString="Marin, I. & Okuno, J. (2010) Laomenes holthuisi sp. nov., a new species of crinoid-associated pontoniine shrimp (Decapoda, Caridea, Palaemonidae) from the Izu Islands, Japan, with some remarks on Japanese species of the genus Laomenes. In: Fransen, C. H. J. M., de Grave, S. & Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 461 - 472, figs. 1 - 6. http: // dx. doi. org / 10.1163 / 9789047427759 _ 031" type="journal article" year="2010">Marin & Okuni, 2010</bibRefCitation>
|
||
,
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EF569BFDFABAE8EA" ID-CoL="1b05c598-e9f4-44cf-8afa-faa63d1335c9" authority="Marin, Chan & Okuno, 2012" authorityName="Marin, Chan & Okuno" authorityYear="2012" box="[895,1384,188,213]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="pestrushka">
|
||
<emphasis id="E6C75D20FFF6FFE7EF569BFDFBC4E8EB" box="[895,1046,188,213]" italics="true" pageId="6" pageNumber="445">L. pestrushka</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7E8379BFCFABAE8EA" author="Marin" box="[1054,1384,188,213]" pageId="6" pageNumber="445" refString="Marin, I., Chan, T. - Y. & Okuno, J. (2012) Crinoid associated pontoniine shrimps of the genus Laomenes Clark, 1919 (Decapoda, Caridea, Palaemonidae) from Panglao 2004 and Kumejima 2009 Expeditions, with descriptions of two new species. Zootaxa, 3367, 103 - 114, figs. 1 - 8." type="journal article" year="2012">Marin, Chan & Okuno, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECBE9BA1FD79E8C6" ID-CoL="931e0101-130f-4db6-905a-9dc0ea2b37fc" authority="Marin, Chan & Okuno, 2012" authorityName="Marin, Chan & Okuno" authorityYear="2012" box="[151,683,223,248]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="gyrophthalmus">
|
||
<emphasis id="E6C75D20FFF6FFE7ECBE9BA1FE8EE8C6" box="[151,348,223,248]" italics="true" pageId="6" pageNumber="445">L. gyrophthalmus</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7ED4A9B9FFD79E8C6" author="Marin" box="[355,683,223,248]" pageId="6" pageNumber="445" refString="Marin, I., Chan, T. - Y. & Okuno, J. (2012) Crinoid associated pontoniine shrimps of the genus Laomenes Clark, 1919 (Decapoda, Caridea, Palaemonidae) from Panglao 2004 and Kumejima 2009 Expeditions, with descriptions of two new species. Zootaxa, 3367, 103 - 114, figs. 1 - 8." type="journal article" year="2012">Marin, Chan & Okuno, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF6FFE7ECEE9A44FF04E9B6" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="445">
|
||
The
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECD59A45FE48E923" ID-CoL="4055477c-0294-4f2e-957f-b08b6c312529" box="[252,410,260,285]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="jackhintoni">
|
||
<emphasis id="E6C75D20FFF6FFE7ECD59A45FE48E923" box="[252,410,260,285]" italics="true" pageId="6" pageNumber="445">L. jackhintoni</emphasis>
|
||
</taxonomicName>
|
||
group is characterized by the absence of ventral rostral teeth, extremely elongated apical corneal papilla, third pereiopod propod without distoventral spines, with large accessory tooth on dactyl, and a fixed tooth on exopod of uropod. Included species are
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EF399A0DFB63E95B" ID-CoL="4055477c-0294-4f2e-957f-b08b6c312529" authority="Bruce, 2006" authorityName="Bruce" authorityYear="2006" box="[784,1201,332,357]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="jackhintoni">
|
||
<emphasis id="E6C75D20FFF6FFE7EF399A0DFBDBE95B" box="[784,1033,332,357]" italics="true" pageId="6" pageNumber="445">Laomenes jackhintoni</emphasis>
|
||
(
|
||
<bibRefCitation id="B022FCC3FFF6FFE7E8339A0CFB7BE95B" author="Bruce" box="[1050,1193,332,357]" pageId="6" pageNumber="445" refString="Bruce, A. J. (2006) Periclimenes jackhintoni sp. nov. (Crustacea: Decapoda: Palaemonidae), a new pontoniine shrimp and crinoid associate from Tonga. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 22, 23 - 29, figs. 1 - 5." type="journal article" year="2006">Bruce, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13B3FAB1FFF6FFE7E8C59A0DFF01E9B6" ID-CoL="691f48ab-3c29-4ed9-b812-b801ba88bde3" authority="Marin, 2009" authorityName="Marin" authorityYear="2009" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="tigris">
|
||
<emphasis id="E6C75D20FFF6FFE7E8C59A0DFA9AE95A" box="[1260,1352,333,356]" italics="true" pageId="6" pageNumber="445">L. tigris</emphasis>
|
||
<bibRefCitation id="B022FCC3FFF6FFE7E9799A0CFF01E9B6" author="Marin" pageId="6" pageNumber="445" refString="Marin, I. (2009) Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity. Zootaxa, 1971, 1 - 49, figs. 1 - 36." type="journal article" year="2009">Marin, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF6FFE7ECEE9AD4FC41EA26" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="445">
|
||
The
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ED299AD5FE32E993" ID-CoL="4a11748e-6dd0-47e5-a909-6d50036fde82" box="[256,480,404,429]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="E6C75D20FFF6FFE7ED299AD5FE32E993" box="[256,480,404,429]" italics="true" pageId="6" pageNumber="445">L. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
group is characterized by the absence of ventral rostral teeth, cornea papillate, distoventral spines on propod and accessory tooth on dactyl of third pereiopod small, tiny marginal dorsal spines on telson and the absence of a fixed distolateral tooth on uropodal exopod. Included species are
|
||
<taxonomicName id="13B3FAB1FFF6FFE7E9039A9DFDC9EA26" ID-CoL="ebead364-e4a5-4ae4-9774-e9654e54d3ea" authority="Borradaile1915" authorityName="Borradaile" authorityYear="1915" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="E6C75D20FFF6FFE7E9039A9DFE81EA26" italics="true" pageId="6" pageNumber="445">Laomenes ceratophthalmus</emphasis>
|
||
(
|
||
<bibRefCitation id="B022FCC3FFF6FFE7ED4B9ABFFDC6EA26" author="Borradaile" box="[354,532,511,536]" pageId="6" pageNumber="445" refString="Borradaile, L. A. (1915) Notes on Carides. The Annals and Magazine of Natural History, Series 8, 15, 205 - 213. http: // dx. doi. org / 10.1080 / 00222931508693629" type="journal article" year="1915">Borradaile1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EE789941FC5CEA26" ID-CoL="7d97cf11-095f-4d06-ad51-366e12555634" authority="Bruce, 1968" authorityName="Bruce" authorityYear="1968" box="[593,910,511,536]" class="Malacostraca" family="Palaemonidae" genus="Parapontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="nudirostris">
|
||
<emphasis id="E6C75D20FFF6FFE7EE789941FD3BEA26" box="[593,745,511,536]" italics="true" pageId="6" pageNumber="445">L. nudirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="B022FCC3FFF6FFE7EED19940FC54EA26" author="Bruce" box="[760,902,511,536]" pageId="6" pageNumber="445" refString="Bruce, A. J. (1968) A report on some pontoniid shrimps from New Caledonia. Bulletin du Museum National d'Histoire Naturelle, 39 (6), 1148 - 1171." type="journal article" year="1968">Bruce, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D40C8132FFF6FFE7ECEE9965FC05EB06" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="445">
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECEE9965FE6CEA03" box="[199,446,548,573]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="albonigrus">
|
||
<emphasis id="E6C75D20FFF6FFE7ECEE9965FE6CEA03" box="[199,446,548,573]" italics="true" pageId="6" pageNumber="445">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EE509965FCF5EA03" ID-CoL="0d88ed20-0800-4b1d-8595-f5f5f0d45f4e" box="[633,807,548,573]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="amboinensis">
|
||
<emphasis id="E6C75D20FFF6FFE7EE509965FCF5EA03" box="[633,807,548,573]" italics="true" pageId="6" pageNumber="445">L. amboinensis</emphasis>
|
||
</taxonomicName>
|
||
species group as it has ventral rostral teeth, a robust antennal tooth, a hemispherical cornea, epistomal horns well developed, acute, second pereiopods unequal, major fingers less than half palm length, very strongly dentate, minor feebly dentate, third pereiopod dactyl with small anteroverted accessory tooth, third pereiopod propod with setiform distoventral spines, uropodal exopod with distolateral tooth, dorsal telson spines normal.
|
||
<collectingCountry id="ACA4C1A2FFF6FFE7EE9E99F4FD04EAF2" box="[695,726,692,716]" name="American Samoa" pageId="6" pageNumber="445">As</collectingCountry>
|
||
noted by
|
||
<bibRefCitation id="B022FCC3FFF6FFE7EF7999F4FBFBEAF2" box="[848,1065,692,717]" pageId="6" pageNumber="445" refString="Marin, I., Chan, T. - Y. & Okuno, J. (2012) Crinoid associated pontoniine shrimps of the genus Laomenes Clark, 1919 (Decapoda, Caridea, Palaemonidae) from Panglao 2004 and Kumejima 2009 Expeditions, with descriptions of two new species. Zootaxa, 3367, 103 - 114, figs. 1 - 8." type="journal article">
|
||
Marin
|
||
<emphasis id="E6C75D20FFF6FFE7EFB699F5FC00EAF3" box="[927,978,692,717]" italics="true" pageId="6" pageNumber="445">et al</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
<taxonomicName id="13B3FAB1FFF6FFE7E81A99F5FB29EAF3" ID-CoL="931e0101-130f-4db6-905a-9dc0ea2b37fc" box="[1075,1275,692,717]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="gyrophthalmus">
|
||
<emphasis id="E6C75D20FFF6FFE7E81A99F5FB29EAF3" box="[1075,1275,692,717]" italics="true" pageId="6" pageNumber="445">L. gyrophthalmus</emphasis>
|
||
</taxonomicName>
|
||
it is the only species in this group, and in the genus
|
||
<emphasis id="E6C75D20FFF6FFE7EE639999FD11EACE" box="[586,707,729,752]" italics="true" pageId="6" pageNumber="445">
|
||
<taxonomicName id="13B3FAB1FFF6FFE7EE639999FD6DEACE" ID-CoL="478b2f61-861b-4312-8a50-abdeab12ca67" box="[586,703,729,752]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="genus">Laomenes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
that has a globular cornea lacking any vestige of a distal papilla.
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECBE99BDFE5AEB2B" box="[151,392,764,789]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="albonigrus">
|
||
<emphasis id="E6C75D20FFF6FFE7ECBE99BDFE5AEB2B" box="[151,392,764,789]" italics="true" pageId="6" pageNumber="445">Laomenes albonigrus</emphasis>
|
||
</taxonomicName>
|
||
is a second species with this character and is closely similar morphologically. The species of the
|
||
<taxonomicName id="13B3FAB1FFF6FFE7ECE9985FFE9EEB06" ID-CoL="0d88ed20-0800-4b1d-8595-f5f5f0d45f4e" box="[192,332,799,824]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="amboinensis">
|
||
<emphasis id="E6C75D20FFF6FFE7ECE9985FFE9EEB06" box="[192,332,799,824]" italics="true" pageId="6" pageNumber="445">amboinensis</emphasis>
|
||
</taxonomicName>
|
||
species group may be distinguished by the following key.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |