170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="60466F5EB21FB5126D95882C9992A8B4" ID="10.11646/zootaxa.4178.3.6" ID-DOI="10.11646/zootaxa.4178.3.6" ID-GBIF-Dataset="44b8171c-8531-4192-af43-fadab952741d" ID-ISSN="1175-5326" ID-Zenodo-Dep="254783" ID-ZooBank="3278A5A0-B5FE-4C05-8C09-BAA31E87B9B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477475351260" checkinUser="plazi" docAuthor="Alekseev, Vitalii I. & Bukejs, Andris" docDate="2016" docId="F8765132CF60FF92FF68FEA71B2C7B83" docLanguage="en" docName="zootaxa.4178.3.6.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xylolaemus richardklebsi Alekseev & Bukejs, sp. nov." docType="treatment" docUuid="31270420-4171-4B3E-A8A6-50451CAB8EC7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="413" masterDocId="044F294ACF63FF96FFFFFF801E357B56" masterDocTitle="New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber" masterLastPageNumber="427" masterPageNumber="409" pageNumber="412" updateTime="1698390869414" updateUser="plazi">
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<mods:mods id="ACAB4468282E3A67FB4D3F2138C08852" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6E91E74D042E1646BAD1D8B88BFB15F1">
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<mods:title id="C2C1A2164F53B603A1094711EC9A37B4">New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber</mods:title>
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</mods:titleInfo>
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<mods:name id="78FEC03D9CF301BFAA604CA6D89C8BA9" type="personal">
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<mods:role id="76977D0D9A0CEA6F1D2260650574C25F">
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<mods:roleTerm id="25F94FEC09E60FCD44A3375E6C3E62FA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="93444FED914B01EA0AE79287FF91B6A9">Alekseev, Vitalii I.</mods:namePart>
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</mods:name>
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<mods:name id="1B0D6460992C98662F886F77D652B391" type="personal">
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<mods:role id="3DD63C5A20B157941C0A0324DA39E1A4">
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<mods:roleTerm id="3FF3935C8E0AE2027EEBF5F373EC5DF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BAC91D926C7B0336E80930B341DB5375">Bukejs, Andris</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="29B9925B8D83F6939A452A49A2457A35">text</mods:typeOfResource>
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<mods:relatedItem id="CA18B00932162BB2A7690E34BC1B7093" type="host">
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<mods:titleInfo id="DAFDC00DB835EFA934BA4423BD9B0BF4">
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<mods:title id="66E11705C879111077CBD93C3AD910EA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="463948572659CAB1946B0BEDF1991CE7">
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<mods:date id="F79F49491523FF3B05CAC340BB2AEDE6">2016</mods:date>
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<mods:detail id="3F268BDBBB0C1665A62A0B2520269EA5" type="volume">
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<mods:number id="3BF12DB70AD39A57B9BCC725D545E057">4178</mods:number>
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</mods:detail>
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<mods:detail id="C82FB40283BB0A4F9A7EAD6A63ADB4EA" type="issue">
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<mods:number id="850070FA86F935C0C43944166265AF47">3</mods:number>
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</mods:detail>
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<mods:extent id="E9009C8618CCD97F08F40B54F1D94934" unit="page">
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<mods:end id="7B1E7B87B474515ED6D1581163C408CD">427</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="45DA3CD6CD42436C1A9D28FB64A0BCF0">journal article</mods:classification>
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<mods:identifier id="999FF11EF4CC15A55E0A2ABEA89B2E2E" type="DOI">10.11646/zootaxa.4178.3.6</mods:identifier>
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<mods:identifier id="0B0B110542500065639233D07D86368E" type="GBIF-Dataset">44b8171c-8531-4192-af43-fadab952741d</mods:identifier>
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<mods:identifier id="A0EA67797E100A7A919D1C8E27B2B6D2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3A8FA5A8B571DB8D8513A7F90674264C" type="Zenodo-Dep">254783</mods:identifier>
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<mods:identifier id="EB93EC78F211BDAE1C3374B1D715E661" type="ZooBank">3278A5A0-B5FE-4C05-8C09-BAA31E87B9B9</mods:identifier>
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</mods:mods>
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<treatment id="F8765132CF60FF92FF68FEA71B2C7B83" ID-DOI="http://doi.org/10.5281/zenodo.6068524" ID-GBIF-Taxon="124558193" ID-Zenodo-Dep="6068524" LSID="urn:lsid:zoobank.org:act:31270420-4171-4B3E-A8A6-50451CAB8EC7" httpUri="http://treatment.plazi.org/id/F8765132CF60FF92FF68FEA71B2C7B83" lastPageId="4" lastPageNumber="413" pageId="3" pageNumber="412">
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<subSubSection id="38C5B3AFCF60FF95FF68FEA71F017A35" pageId="3" pageNumber="412" type="nomenclature">
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<paragraph id="7060E024CF60FF95FF68FEA71C847A14" blockId="3.[151,689,295,355]" box="[151,689,295,322]" pageId="3" pageNumber="412">
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<heading id="2B285748CF60FF95FF68FEA71C847A14" bold="true" box="[151,689,295,322]" fontSize="11" level="1" pageId="3" pageNumber="412" reason="1">
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<emphasis id="42AB3C36CF60FF95FF68FEA71C847A14" bold="true" box="[151,689,295,322]" pageId="3" pageNumber="412">
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<taxonomicName id="B7DF9BA7CF60FF95FF68FEA71C7E7A14" authority="Alekseev & Bukejs" authorityName="Alekseev & Bukejs" box="[151,587,295,322]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="richardklebsi" status="sp. nov.">
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<emphasis id="42AB3C36CF60FF95FF68FEA71F6C7A17" bold="true" box="[151,345,295,321]" italics="true" pageId="3" pageNumber="412">X. richardklebsi</emphasis>
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Alekseev & Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="5998814DCF60FF95FDACFEA81C847A14" box="[595,689,296,322]" pageId="3" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7060E024CF60FF95FF68FECA1F017A35" blockId="3.[151,689,295,355]" box="[151,308,330,355]" pageId="3" pageNumber="412">
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(
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<figureCitation id="E8E4FCA1CF60FF95FF60FECA1F317A35" box="[159,260,330,355]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[225,1361,267,1903]" captionTargetId="figure@4.[225,1362,267,1904]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Xylolaemus richardklebsi sp. nov., holotype: 4 — habitus, dorsal view; 5 — habitus, ventral view; 6 — habitus, lateral view." httpUri="https://zenodo.org/record/254785/files/figure.png" pageId="3" pageNumber="412">Figs 4–6</figureCitation>
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,
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<figureCitation id="E8E4FCA1CF60FF95FEF0FECB1F197A35" box="[271,300,331,355]" captionStart="FIGURES 22 – 23" captionStartId="11.[151,264,1556,1579]" captionTargetBox="[150,1427,746,1518]" captionTargetId="figure@11.[150,1436,735,1523]" captionTargetPageId="11" captionText="FIGURES 22 – 23. Reconstructions of habitus, dorsal view: 22 — Xylolaemus legalovi sp. nov.; 23 — X. richardklebsi sp. nov.; 24 — Diodesma slipinskii sp. nov." httpUri="https://zenodo.org/record/254790/files/figure.png" pageId="3" pageNumber="412">23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="38C5B3AFCF60FF95FF68FE121EDC79F1" pageId="3" pageNumber="412" type="materials_examined">
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<paragraph id="7060E024CF60FF95FF68FE121D1E796D" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF68FE121F707AFD" bold="true" box="[151,325,402,427]" pageId="3" pageNumber="412">Type material.</emphasis>
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Holotype: Nr. 138-3 [CCHH] “Holotype /
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<taxonomicName id="B7DF9BA7CF60FF95FCDAFE121A767AFD" box="[805,1091,402,427]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="richardklebsi" status="sp. nov.">
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<emphasis id="42AB3C36CF60FF95FCDAFE121A767AFD" box="[805,1091,402,427]" italics="true" pageId="3" pageNumber="412">Xylolaemus richardklebsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5998814DCF60FF95FBB5FE131A957AFC" box="[1098,1184,403,426]" pageId="3" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
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/ des. Alekseev V.I. & Bukejs A.” [red hand-written label]; sex unknown. Complete beetle with partially exposed hind wings and distal portion of genitalia visible, included in small, transparent yellow amber piece embedded in GTS-polyester resin block with dimensions 15 × 7 ×
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<quantity id="B7274DC1CF60FF95FDE0FE7E1C527940" box="[543,615,510,535]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="412" unit="mm" value="6.0">6 mm</quantity>
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. Syninclusions (except one gas bubble) are absent—amber piece with inclusion may have been thermally processed in autoclave.
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38FDC61B467908" blockId="3.[151,1437,402,2011]" box="[199,1395,582,607]" pageId="3" pageNumber="412">
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<materialsCitation id="C0B7EA79CF60FF95FF38FDC61B467908" ID-GBIF-Occurrence="1324200994" box="[199,1395,582,607]" county="Estimated" location="Baltic" municipality="Prussian Formation" pageId="3" pageNumber="412" specimenCount="1" typeStatus="holotype">
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<emphasis id="42AB3C36CF60FF95FF38FDC61F607909" bold="true" box="[199,341,582,607]" pageId="3" pageNumber="412">
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<typeStatus id="AF645E86CF60FF95FF38FDC61F377909" box="[199,258,582,607]" pageId="3" pageNumber="412">Type</typeStatus>
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strata.
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</emphasis>
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<location id="7500B6FFCF60FF95FEA3FDC61FAA7909" LSID="urn:lsid:plazi:treatment:F8765132CF60FF92FF68FEA71B2C7B83:7500B6FFCF60FF95FEA3FDC61FAA7909" box="[348,415,582,607]" county="Estimated" municipality="Prussian Formation" name="Baltic" pageId="3" pageNumber="412">Baltic</location>
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amber,
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<location id="7500B6FFCF60FF95FE06FDC71CAE7908" LSID="urn:lsid:plazi:treatment:F8765132CF60FF92FF68FEA71B2C7B83:7500B6FFCF60FF95FE06FDC71CAE7908" box="[505,667,583,606]" county="Estimated" municipality="Prussian Formation" name="Upper Eocene" pageId="3" pageNumber="412">Upper Eocene</location>
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,
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<collectingMunicipality id="90047A5ECF60FF95FD59FDC61DB47909" box="[678,897,582,607]" pageId="3" pageNumber="412">Prussian Formation</collectingMunicipality>
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(Priabonian).
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<collectingCounty id="990198A8CF60FF95FBDDFDC61AA77909" box="[1058,1170,582,607]" pageId="3" pageNumber="412">Estimated</collectingCounty>
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age: 37.2–33.9 Ma.
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</materialsCitation>
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38FDEA1EDC79F1" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<materialsCitation id="C0B7EA79CF60FF95FF38FDEA1EDC79F1" ID-GBIF-Occurrence="1324201002" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="3" pageNumber="412" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="42AB3C36CF60FF95FF38FDEA1F5079D5" bold="true" box="[199,357,618,643]" pageId="3" pageNumber="412">
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<typeStatus id="AF645E86CF60FF95FF38FDEA1F3779D5" box="[199,258,618,643]" pageId="3" pageNumber="412">Type</typeStatus>
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locality.
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</emphasis>
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<location id="7500B6FFCF60FF95FE93FDEB1FE479D5" LSID="urn:lsid:plazi:treatment:F8765132CF60FF92FF68FEA71B2C7B83:7500B6FFCF60FF95FE93FDEB1FE479D5" box="[364,465,619,643]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="3" pageNumber="412" stateProvince="Kaliningrad">Yantarny</location>
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settlement (formerly Palmnicken),
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<collectingMunicipality id="90047A5ECF60FF95FC9EFDEA1DF679D5" box="[865,963,618,643]" pageId="3" pageNumber="412">Sambian</collectingMunicipality>
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(Samland)
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<collectingCounty id="990198A8CF60FF95FBB9FDEA1A8379D5" box="[1094,1206,618,643]" pageId="3" pageNumber="412">Peninsula</collectingCounty>
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,
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<collectingRegion id="B21B2EC6CF60FF95FB3EFDEA1B7279D5" box="[1217,1351,618,643]" country="Russia" name="Kaliningrad" pageId="3" pageNumber="412">Kaliningrad</collectingRegion>
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region,
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<collectingCountry id="08C8A0B4CF60FF95FF68FD0E1ED379F1" box="[151,230,654,679]" name="Russia" pageId="3" pageNumber="412">Russia</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="38C5B3AFCF60FF95FF38FD321DA17860" pageId="3" pageNumber="412" type="diagnosis">
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<paragraph id="7060E024CF60FF95FF38FD321DA17860" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF38FD321FF8799D" bold="true" box="[199,461,690,715]" pageId="3" pageNumber="412">Differential diagnosis.</emphasis>
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<taxonomicName id="B7DF9BA7CF60FF95FE2CFD321CC4799D" box="[467,753,690,715]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="richardklebsi" status="sp. nov.">
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<emphasis id="42AB3C36CF60FF95FE2CFD321CC4799D" box="[467,753,690,715]" italics="true" pageId="3" pageNumber="412">Xylolaemus richardklebsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5998814DCF60FF95FD08FD321D65799D" box="[759,848,690,715]" pageId="3" pageNumber="412" rank="species">
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<emphasis id="42AB3C36CF60FF95FD08FD321D65799D" bold="true" box="[759,848,690,715]" pageId="3" pageNumber="412">sp. nov.</emphasis>
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</taxonomicNameLabel>
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differs from extant and fossil species of the genus in the following combination of characters: pronotum with longitudinal impression medially; setae on elytra long and strongly curved; pronotal granulation dense; prosternal process dilated; fascicules (setose patches) on elytra absent; antennae 11-segmented with antennomere 10 dilated and transverse.
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</paragraph>
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</subSubSection>
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<subSubSection id="38C5B3AFCF60FF95FF38FCC21ACF7C8D" pageId="3" pageNumber="412" type="description">
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<paragraph id="7060E024CF60FF95FF38FCC21D307829" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF38FCC21F6C780D" bold="true" box="[199,345,834,859]" pageId="3" pageNumber="412">Description.</emphasis>
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Body length about
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<quantity id="B7274DC1CF60FF95FDBCFCC31CAB780C" box="[579,670,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="412" unit="mm" value="4.1">4.1 mm</quantity>
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, maximal width
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<quantity id="B7274DC1CF60FF95FC9BFCC31DF4780C" box="[868,961,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="412" unit="mm" value="1.4">1.4 mm</quantity>
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; elongate, nearly parallel-sided, flattened dorsally; body and appendages appear uniformly black.
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38FC0A1F837F95" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF38FC0A1F3178F5" box="[199,260,906,931]" italics="true" pageId="3" pageNumber="412">Head</emphasis>
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. Longer than wide; widest between antennae; covered with moderately large, round, unisetose granules. Anterior clypeal margin straight, covered with thin setae. Lateral margins convex above antennal insertions and narrowing to anterior margin. Eyes large, prominent, hemispherical, with coarse facets; interfacetal setae apparent. Maxillary palpus 4-segmented, short; palpomere 4 oval-shaped, slightly longer and narrower than palpomere 3. Labial palpus 3-segmented, palpomere 3 oval, as long as penultimate palpomere. Antennae short, extending to basal half of pronotum; 11-segmented with distinct, 2-segmented, finely pubescent antennal club, sparsely covered with dark, semierect setae. Scape, pedicel and antennomeres 3–4 elongate, conical; antennomeres 5–9 transverse; antennomere 10 transverse, dilated distally, 0.7× as long as wide; antennomere 11 widely ovate with rounded apex, slightly longer than wide.
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38FB4E1CEA7D50" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF38FB4E1F2D7FB1" box="[199,280,1230,1255]" italics="true" pageId="3" pageNumber="412">Thorax</emphasis>
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. Pronotum transverse, 1.3× as wide as long, widest at middle; pronotal disc convex with longitudinal impression at middle, lateral sides explanate; densely covered with unisetose granules (slightly smaller and distinctly denser than granules on frons). Anterior margin arcuate; lateral margins nearly parallel-sided, finely denticulate (about 12 small denticles), with each denticle bearing short seta; posterior margin rounded; anterior angles triangular, acute, prominent, reaching posterior half of eye; posterior angles obtuse. Prosternum densely covered with granules (sparser than pronotal granulation); procoxa nearly round, shagreened. Granules of meso- and metasternum flatter than those of prosternum. Procoxal cavities externally open, meso- and metacoxal cavities closed; all coxae narrowly separated from each other. Prosternal process oblong parallel-sided between procoxae and then strongly dilated,with widely rounded apex.
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38F9921CF77C55" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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Scutellum small, rounded. Elytra elongate (length
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<quantity id="B7274DC1CF60FF95FCF3F9931D527D7C" box="[780,871,1555,1579]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="412" unit="mm" value="2.7">2.7 mm</quantity>
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, maximum width
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<quantity id="B7274DC1CF60FF95FBC5F9931AA27D7D" box="[1082,1175,1555,1579]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="412" unit="mm" value="1.4">1.4 mm</quantity>
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), nearly parallel-sided, flattened dorsally, slightly wider than pronotum, jointly rounded apically. Humeral angles rounded. Each elytron with 10 rows of small, round punctures, distance between strial punctures equal to diameter of one puncture; intervals flat, distance between striae approximately 1.5–2.0× diameter of single puncture. Striae with small regular tubercles, bearing long, strongly curved setae. Scutellary striole apparent, consisting of four punctures. Epipleura well-developed, reaching apex of elytra, widest at humeral angle. Hind wings apparent. Ratio of lengths of mesoventrite to metaventrite to abdomen:
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<date id="0461C6E4CF60FF95FD8EF96A1C8B7C55" box="[625,702,1770,1795]" pageId="3" pageNumber="412" value="2014-05-03">3-5-14</date>
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.
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</paragraph>
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<paragraph id="7060E024CF60FF95FF38F88E1BA97C38" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
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<emphasis id="42AB3C36CF60FF95FF38F88E1F047C71" box="[199,305,1806,1831]" italics="true" pageId="3" pageNumber="412">Abdomen</emphasis>
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. With five visible, similarly articulated ventrites. Ventrite length ratios: 6-4-5-3-3. Setation of ventrites 1–4 not apparent (possibly absent); ventrite 5 rounded apically, finely pubescent, with two long, erect setae near apical margin, length of setae approximately
|
||
<quantity id="B7274DC1CF60FF95FCF8F8D71D5B7C38" box="[775,878,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="412" unit="mm" value="0.14">0.14 mm</quantity>
|
||
. Intercoxal process of abdominal ventrite 1 acute.
|
||
</paragraph>
|
||
<paragraph id="7060E024CF60FF95FF38F8FC1ACF7C8D" blockId="3.[151,1437,402,2011]" pageId="3" pageNumber="412">
|
||
<emphasis id="42AB3C36CF60FF95FF38F8FC1EC97CC5" box="[199,252,1916,1939]" italics="true" pageId="3" pageNumber="412">Legs</emphasis>
|
||
. Profemora thickened medially; tibiae covered with curved setae. Tarsi tetramerous. Metatarsomere 4 as long as metatarsomeres 1–3 combined; pro- and mesotarsomeres 4 slightly shorter than combined length of tarsomeres 1–3. Tarsal claws simple, large, equal in size, with length about one-third of tarsomere 4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="38C5B3AFCF67FF92FF38FF171B2C7B83" pageId="4" pageNumber="413" type="etymology">
|
||
<paragraph id="7060E024CF67FF92FF38FF171B2C7B83" blockId="4.[151,1436,151,213]" pageId="4" pageNumber="413">
|
||
<emphasis id="42AB3C36CF67FF92FF38FF171F7B7BE6" bold="true" box="[199,334,151,176]" pageId="4" pageNumber="413">Etymology.</emphasis>
|
||
Patronymic, this new species is dedicated to Professor Dr. Richard Hermann Erdmann Klebs (
|
||
<date id="0461C6E4CF67FF92FF60FF3C1F2A7B83" box="[159,287,188,213]" pageId="4" pageNumber="413" value="1850" valueMax="1911">1850–1911</date>
|
||
) from Königsberg, a world-renowned East-Prussian expert of Baltic amber and inclusions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |