376 lines
53 KiB
XML
376 lines
53 KiB
XML
<document id="88DBA473DD748CC489953B749F9CAFF1" ID-DOI="10.15407/zoo2024.01.19" ID-ISSN="2707-7268" ID-Zenodo-Dep="10788414" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709735874948" checkinUser="felipe" docAuthor="Satria, R., Nugraha, F. A. D., Achyar, A., Zulyetti, D. & Eguchi, K." docDate="2024" docId="C32C87F9FFCF143CB09FD5506C3AE156" docLanguage="en" docName="Zoodiversity.58.1.19-26.pdf" docOrigin="Zoodiversity 58 (1)" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Leptogenys borneensis Wheeler 1919" docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="3F15FF81FFCE1439B031D3586E69E504" masterDocTitle="Description Of The Male Caste Of Leptogenys Borneensis (Hymenoptera, Formicidae), With Its First Discovery In Sumatra" masterLastPageNumber="26" masterPageNumber="19" pageNumber="20" updateTime="1709739080448" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="758C9B56E2149877410B83C0CA5D8FE2">Description Of The Male Caste Of Leptogenys Borneensis (Hymenoptera, Formicidae), With Its First Discovery In Sumatra</mods:title>
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<mods:namePart id="82D045BC16D579880E0C36612DC5CD9B">Satria, R.</mods:namePart>
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<mods:affiliation id="9DDC70075A8B1340EE5284360AABCAE7">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia & Biodiversity of Sumatra Research Group, Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia</mods:affiliation>
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<mods:namePart id="5529D4CC0D1682FE7D8F432BA994F883">Nugraha, F. A. D.</mods:namePart>
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<mods:affiliation id="8AF7AD1FB217E550DB590CD80304B3E7">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia & Biodiversity of Sumatra Research Group, Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia</mods:affiliation>
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<mods:namePart id="F802A6AD665BD53BE73F077D4F6CBDFB">Achyar, A.</mods:namePart>
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<mods:affiliation id="27BA5B583112EF0BE5E0E3DB903F3EF8">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia & Biodiversity of Sumatra Research Group, Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, West Sumatra, 35171, Indonesia</mods:affiliation>
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<mods:namePart id="79A3BCB787A573110EF62C5090B32536">Zulyetti, D.</mods:namePart>
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<mods:affiliation id="ACB6FE0610700D4E46E1A23E8217122A">CTKIP Yayasan Abdi Pendidikan Payakumbuh, Payakumbuh, West Sumatra, 26213, Indonesia</mods:affiliation>
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<mods:namePart id="EC8D713B81B0208AB4181626947194C7">Eguchi, K.</mods:namePart>
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<mods:affiliation id="74A18BCEE9DBA2A9DB15EED4B4C61E3D">Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Tokyo, 1920397, Japan</mods:affiliation>
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<mods:title id="10C20D1A5F74C826596D8854A0E59B60">Zoodiversity</mods:title>
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<mods:date id="61FC7DF2098690CCEDE201443C9B499C">2024</mods:date>
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<mods:number id="E5A5204BBE503907D4B29ABB85FE6C7B">58</mods:number>
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<mods:start id="8B9F71BFCAA8AB54D0D9FA542A16608F">19</mods:start>
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<mods:end id="A00CA1AEFDB04C854C939217A3E7DEE8">26</mods:end>
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<mods:identifier id="BFC69C12B0ADA0A999F6899DE3DA48F5" type="DOI">10.15407/zoo2024.01.19</mods:identifier>
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<mods:identifier id="4F0A7585CA7060C1973BEF73CCB1D5C3" type="ISSN">2707-7268</mods:identifier>
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<treatment id="C32C87F9FFCF143CB09FD5506C3AE156" LSID="urn:lsid:plazi:treatment:C32C87F9FFCF143CB09FD5506C3AE156" httpUri="http://treatment.plazi.org/id/C32C87F9FFCF143CB09FD5506C3AE156" lastPageId="5" lastPageNumber="24" pageId="1" pageNumber="20">
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<subSubSection id="039F6564FFCF1438B09FD5506CB4E326" box="[174,733,1544,1571]" pageId="1" pageNumber="20" type="nomenclature">
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<paragraph id="4B3A36EFFFCF1438B09FD5506CB4E326" blockId="1.[174,733,1544,1571]" box="[174,733,1544,1571]" pageId="1" pageNumber="20">
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<heading id="10728183FFCF1438B09FD5506CB4E326" box="[174,733,1544,1571]" fontSize="36" level="2" pageId="1" pageNumber="20" reason="3">
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<taxonomicName id="8C854D6CFFCF1438B09FD5506C01E326" ID-CoL="3TPD3" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[174,616,1544,1571]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="borneensis">
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<emphasis id="79F1EAFDFFCF1438B09FD5506FD1E327" bold="true" box="[174,440,1544,1571]" italics="true" pageId="1" pageNumber="20">Leptogenys borneensis</emphasis>
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<bibRefCitation id="2F144B1EFFCF1438B18FD5506C01E326" author="Wheeler, W. M." box="[446,616,1544,1570]" pageId="1" pageNumber="20" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="D3BE2A6AFFCF1438B247D5506CBDE326" box="[630,724,1544,1570]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[166,199,925,947]" captionStartId-1="2.[579,612,1769,1791]" captionStartId-2="3.[174,207,971,993]" captionTargetBox-0="[180,1212,175,898]" captionTargetBox-1="[166,1226,1159,1854]" captionTargetBox-2="[191,1207,174,944]" captionTargetId-0="figure-168@2.[180,1212,175,898]" captionTargetId-1="figure-214@2.[576,1225,1237,1752]" captionTargetId-2="figure-163@3.[191,1207,174,950]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Leptogenys borneensis Wheeler, 1919 (worker, individual code: SEMUT13xii2021-A, colony code: RS- SAGO-02):A — head in full-face view;B — body in lateral view; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view." captionText-1="Fig. 2. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS-SAGO-02): habitus in lateral view." captionText-2="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi-0="http://doi.org/10.5281/zenodo.10788416" figureDoi-1="http://doi.org/10.5281/zenodo.10788418" figureDoi-2="http://doi.org/10.5281/zenodo.10788422" httpUri-0="https://zenodo.org/record/10788416/files/figure.png" httpUri-1="https://zenodo.org/record/10788418/files/figure.png" httpUri-2="https://zenodo.org/record/10788422/files/figure.png" pageId="1" pageNumber="20">figs 1–3</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="039F6564FFCF1438B09FD5746F0EE36C" pageId="1" pageNumber="20" type="reference_group">
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<paragraph id="4B3A36EFFFCF1438B09FD5746F0EE36C" blockId="1.[174,1232,1579,1640]" pageId="1" pageNumber="20">
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<treatmentCitationGroup id="6B9511C1FFCF1438B09FD5746F0EE36C" pageId="1" pageNumber="20">
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<treatmentCitation id="CA2410FEFFCF1438B09FD5746D73E342" author="Wheeler, W. M." box="[174,794,1579,1606]" page="59" pageId="1" pageNumber="20" year="1919">
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<taxonomicName id="8C854D6CFFCF1438B09FD5746D73E342" ID-CoL="3TPD3" authority="Wheeler, 1919: 59" authorityName="Wheeler" authorityPageNumber="59" authorityYear="1919" box="[174,794,1579,1606]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="borneensis" subGenus="Lobopelta">
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<emphasis id="79F1EAFDFFCF1438B09FD5746F44E341" box="[174,301,1580,1605]" italics="true" pageId="1" pageNumber="20">Leptogenys</emphasis>
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(
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<emphasis id="79F1EAFDFFCF1438B10ED5736FD8E341" box="[319,433,1579,1605]" italics="true" pageId="1" pageNumber="20">Lobopelta</emphasis>
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)
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<emphasis id="79F1EAFDFFCF1438B1F3D5736C52E341" box="[450,571,1579,1605]" italics="true" pageId="1" pageNumber="20">borneensis</emphasis>
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||
<materialsCitation id="FBED3CB2FFCF1438B272D5736D73E342" box="[579,794,1579,1606]" country="Malaysia" location="Malaysia" pageId="1" pageNumber="20" specimenCount="1">
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||
<bibRefCitation id="2F144B1EFFCF1438B272D5736D73E342" author="Wheeler, W. M." box="[579,794,1579,1606]" pageId="1" pageNumber="20" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919: 59</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, worker,
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<typeStatus id="943E884DFFCF1438B3B9D5746DD3E342" box="[904,954,1580,1606]" pageId="1" pageNumber="20">type</typeStatus>
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locality:
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<collectingCountry id="3392767FFFCF1438B417D5736AE7E341" box="[1062,1166,1579,1605]" name="Malaysia" pageId="1" pageNumber="20">Malaysia</collectingCountry>
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(Sarawak: Kuching)
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="039F6564FFCF143DB09FD5DD6FE1E0C8" lastPageId="4" lastPageNumber="23" pageId="1" pageNumber="20" type="diagnosis">
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||
<paragraph id="4B3A36EFFFCF1438B09FD5DD6F9EE39B" blockId="1.[174,1234,1669,1836]" box="[174,503,1669,1695]" pageId="1" pageNumber="20">Diagnosis of the worker</paragraph>
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<paragraph id="4B3A36EFFFCF143BB0D2D5F06DD1E15E" blockId="1.[174,1234,1669,1836]" lastBlockId="2.[166,1225,1053,1115]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">Head in frontal view with its posterior margin slightly convex; mandible triangular; clypeus strongly carinate. Mesosoma in lateral view, long and slender; dorsal margin of mesothorax slightly concave; posterior margin of petiole weakly convex. Petiolar node in frontal and lateral views with a bluntly pointed apex, in lateral view with anterior slope broadly convex and posterior slope weakly convex or almost straight; subpetiolar process not developed. The body dark brown with legs and antennae paler.</paragraph>
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<caption id="1FFA6667FFCC143BB097D0C56C51E6E9" ID-DOI="http://doi.org/10.5281/zenodo.10788416" ID-Zenodo-Dep="10788416" httpUri="https://zenodo.org/record/10788416/files/figure.png" pageId="2" pageNumber="21" startId="2.[166,199,925,947]" targetBox="[180,1212,175,898]" targetPageId="2" targetType="figure">
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<paragraph id="4B3A36EFFFCC143BB097D0C56C51E6E9" blockId="2.[165,1225,925,1005]" pageId="2" pageNumber="21">
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Fig. 1.
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<taxonomicName id="8C854D6CFFCC143BB0DAD0C66C26E6B6" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[235,591,925,947]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="borneensis">
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<emphasis id="79F1EAFDFFCC143BB0DAD0C66FD4E6B6" box="[235,445,925,947]" italics="true" pageId="2" pageNumber="21">Leptogenys borneensis</emphasis>
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<bibRefCitation id="2F144B1EFFCC143BB1F3D0C56C26E6B6" author="Wheeler, W. M." box="[450,591,925,947]" pageId="2" pageNumber="21" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919</bibRefCitation>
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</taxonomicName>
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(worker, individual code: SEMUT13xii2021-A, colony code: RS- SAGO-02): A — head in full-face view; B — body in lateral view; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view.
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</paragraph>
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</caption>
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<caption id="1FFA6667FFCC143BB272D5B16C1EE23D" ID-DOI="http://doi.org/10.5281/zenodo.10788418" ID-Zenodo-Dep="10788418" httpUri="https://zenodo.org/record/10788418/files/figure.png" pageId="2" pageNumber="21" startId="2.[579,612,1769,1791]" targetBox="[166,1226,1159,1854]" targetPageId="2" targetType="figure">
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||
<paragraph id="4B3A36EFFFCC143BB272D5B16C1EE23D" blockId="2.[579,1226,1769,1849]" pageId="2" pageNumber="21">
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Fig. 2.
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<taxonomicName id="8C854D6CFFCC143BB2B5D5B26D8FE3FA" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[644,998,1769,1791]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="borneensis">
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||
<emphasis id="79F1EAFDFFCC143BB2B5D5B26D3CE3FA" box="[644,853,1769,1791]" italics="true" pageId="2" pageNumber="21">Leptogenys borneensis</emphasis>
|
||
<bibRefCitation id="2F144B1EFFCC143BB36AD5B16D8FE3FA" author="Wheeler, W. M." box="[859,998,1769,1791]" pageId="2" pageNumber="21" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919</bibRefCitation>
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</taxonomicName>
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(male, individual code: SEMUT13xii2021-B, colony code: RS-SAGO-02): habitus in lateral view.
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</paragraph>
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</caption>
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<caption id="1FFA6667FFCD143AB09FD0936A49E11F" ID-DOI="http://doi.org/10.5281/zenodo.10788422" ID-Zenodo-Dep="10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="3" pageNumber="22" startId="3.[174,207,971,993]" targetBox="[191,1207,174,944]" targetPageId="3" targetType="figure">
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<paragraph id="4B3A36EFFFCD143AB09FD0936A49E11F" blockId="3.[174,1232,971,1051]" pageId="3" pageNumber="22">
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Fig. 3.
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<taxonomicName id="8C854D6CFFCD143AB0C4D0946C09E6E4" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[245,608,971,993]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="borneensis">
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||
<emphasis id="79F1EAFDFFCD143AB0C4D0946FA0E6E4" box="[245,457,971,993]" italics="true" pageId="3" pageNumber="22">Leptogenys borneensis</emphasis>
|
||
<bibRefCitation id="2F144B1EFFCD143AB1E3D0936C09E6E4" author="Wheeler, W. M." box="[466,608,971,993]" pageId="3" pageNumber="22" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919</bibRefCitation>
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</taxonomicName>
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(male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing.
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</paragraph>
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</caption>
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<caption id="1FFA6667FFCD143AB09FD5F86C93E3B2" ID-DOI="http://doi.org/10.5281/zenodo.10788424" ID-Zenodo-Dep="10788424" box="[174,762,1696,1718]" httpUri="https://zenodo.org/record/10788424/files/figure.png" pageId="3" pageNumber="22" startId="3.[174,207,1696,1718]" targetBox="[173,1233,1100,1833]" targetPageId="3" targetType="figure">
|
||
<paragraph id="4B3A36EFFFCD143AB09FD5F86C93E3B2" blockId="3.[173,763,1696,1833]" box="[174,762,1696,1718]" pageId="3" pageNumber="22">
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Fig. 4.
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<taxonomicName id="8C854D6CFFCD143AB0DED5F96C39E3B1" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[239,592,1696,1718]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="borneensis">
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<emphasis id="79F1EAFDFFCD143AB0DED5F96FA9E3B1" box="[239,448,1696,1718]" italics="true" pageId="3" pageNumber="22">Leptogenys borneensis</emphasis>
|
||
<bibRefCitation id="2F144B1EFFCD143AB1F4D5F86C39E3B1" author="Wheeler, W. M." box="[453,592,1696,1718]" pageId="3" pageNumber="22" pagination="43 - 147" refId="ref4886" refString="Wheeler, W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology, 63, 43 - 147." type="journal article" year="1919">Wheeler, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
(male, individual
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4B3A36EFFFCA143DB097D3F76F0EE5E9" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">ly long, tapering apically. Clypeus without longitudinal median carina; anterior margin broadly convex.</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD3AE6F2DE72D" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">
|
||
Mesoscutum in dorsal view longer than wide, widest posterior to mid-length, broadly convex in lateral view; notaulus scrobiculate; parapsidal line longer than half length of mesoscutal sulcus; transscutal line sinuous. Mesopleuron distinct and complete. Metapleuropropodeal suture present. Propodeal dorsum in lateral view straight, longer than length of declivity; propodeal declivity broadly convex; propodeal spiracle circular. Wing venation as in
|
||
<figureCitation id="D3BE2A6AFFCA143DB0D9D2FE6F40E4C4" box="[232,297,422,448]" captionStart="Fig" captionStartId="2.[579,612,1769,1791]" captionTargetBox="[166,1226,1159,1854]" captionTargetId="figure-214@2.[576,1225,1237,1752]" captionTargetPageId="2" captionText="Fig. 2. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS-SAGO-02): habitus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.10788418" httpUri="https://zenodo.org/record/10788418/files/figure.png" pageId="4" pageNumber="23">figs 2</figureCitation>
|
||
, G, H, with stigma; fore wing approximately 3 times as long as wide, approximately 1.3 times as long as hind wing length; hind wing approximately 3.5 times as long as wide. Fore tibia with single spur, each middle and hind tibia with two spurs. Pretarsal claw multidentate.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD16A6F02E647" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">
|
||
Petiole in lateral view longer than height, in dorsal view longer than wide (
|
||
<figureCitation id="D3BE2A6AFFCA143DB46DD16A6AF3E749" box="[1116,1178,562,589]" captionStart="Fig" captionStartId="3.[174,207,971,993]" captionTargetBox="[191,1207,174,944]" captionTargetId="figure-163@3.[191,1207,174,950]" captionTargetPageId="3" captionText="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi="http://doi.org/10.5281/zenodo.10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="4" pageNumber="23">fig. 3</figureCitation>
|
||
, F); lateral margin straight in dorsal view; petiolar node in lateral view subtrapezoidal, anteri- or margin inclined posterad, posterior margin slightly inclined anterad. Anterior lobe of subpetiolar process in lateral view short, with postero-ventral angle sharpened; posteri- or extension in lateral view long and low, separated from anterior lobe by shallow notch, with ventral margin broadly convex. Pygostyle digitiform with long setae on apical half (
|
||
<figureCitation id="D3BE2A6AFFCA143DB09ED05E6E84E624" box="[175,237,774,800]" captionStart="Fig" captionStartId="3.[174,207,971,993]" captionTargetBox="[191,1207,174,944]" captionTargetId="figure-163@3.[191,1207,174,950]" captionTargetPageId="3" captionText="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi="http://doi.org/10.5281/zenodo.10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="4" pageNumber="23">fig. 3</figureCitation>
|
||
, A). Abdominal sternite 9 with elongate, subtriangular apical lobe of which apex is round (
|
||
<figureCitation id="D3BE2A6AFFCA143DB131D0716F55E647" box="[256,316,809,835]" captionStart="Fig" captionStartId="3.[174,207,971,993]" captionTargetBox="[191,1207,174,944]" captionTargetId="figure-163@3.[191,1207,174,950]" captionTargetPageId="3" captionText="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi="http://doi.org/10.5281/zenodo.10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="4" pageNumber="23">fig. 3</figureCitation>
|
||
, B).
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD0146F82E112" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">
|
||
Genital capsule longer than broad. Gonopod apex in lateral view longer than high (
|
||
<figureCitation id="D3BE2A6AFFCA143DB09ED0376E82E68D" box="[175,235,879,905]" captionStart="Fig" captionStartId="3.[174,207,971,993]" captionTargetBox="[191,1207,174,944]" captionTargetId="figure-163@3.[191,1207,174,950]" captionTargetPageId="3" captionText="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi="http://doi.org/10.5281/zenodo.10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="4" pageNumber="23">fig. 3</figureCitation>
|
||
, C). Lateropenite in lateral view rectangular with rounded corners and long, its posterior margin as long as cuspis. Cuspis in lateral view elongate and weakly arched dorsal of lateropenite, apically slightly swollen, rounded. Apicoventral apex of valviceps strongly produced; basiventral corner of valviceps distinctly produced; ventral margin of valviceps with 13 denticles (
|
||
<figureCitation id="D3BE2A6AFFCA143DB14DD0A46FD0E112" box="[380,441,1020,1046]" captionStart="Fig" captionStartId="3.[174,207,971,993]" captionTargetBox="[191,1207,174,944]" captionTargetId="figure-163@3.[191,1207,174,950]" captionTargetPageId="3" captionText="Fig. 3. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS- SAGO-02): A — head in full-face view; B — ocelli; C — mesosoma in lateral view; D — mesosoma in dorsal view; E — petiole in lateral view; F — petiole in dorsal view; G — forewing; H — hind wing." figureDoi="http://doi.org/10.5281/zenodo.10788422" httpUri="https://zenodo.org/record/10788422/files/figure.png" pageId="4" pageNumber="23">fig. 3</figureCitation>
|
||
, D).
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD7476EB4E1A7" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">Head entirely smooth and shiny, except on area between median and lateral ocelli finely striate. Mandible and clypeus weakly sculptured. Pronotum, mesoscutum, mesopleuron, propodeum, petiole and gaster smooth and shiny; mesoscutellum with longitudinal striation.</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD7F46D5AE1ED" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">Head, mesosoma, legs, petiole and gaster with fine dense subdecumbent to decumbent various length of hairs; hairs on head longer than other.</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD7AB6C60E034" blockId="4.[166,1226,175,1328]" pageId="4" pageNumber="23">
|
||
For color pattern see in
|
||
<figureCitation id="D3BE2A6AFFCA143DB234D7AA6C21E009" box="[517,584,1266,1293]" captionStart="Fig" captionStartId="2.[579,612,1769,1791]" captionTargetBox="[166,1226,1159,1854]" captionTargetId="figure-214@2.[576,1225,1237,1752]" captionTargetPageId="2" captionText="Fig. 2. Leptogenys borneensis Wheeler, 1919 (male, individual code: SEMUT13xii2021-B, colony code: RS-SAGO-02): habitus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.10788418" httpUri="https://zenodo.org/record/10788418/files/figure.png" pageId="4" pageNumber="23">figs 2</figureCitation>
|
||
, A–F; body basically dark brown; clypeus, mandible, antennal scape, and legs paler.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB097D6116CFFE067" blockId="4.[166,1225,1353,1485]" box="[166,662,1353,1379]" pageId="4" pageNumber="23">M e a s u r e m e n t s o f t h e M a l e s (n= 3)</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD6356FE1E0C8" blockId="4.[166,1225,1353,1485]" pageId="4" pageNumber="23">
|
||
HW
|
||
<quantity id="8C7D9B0AFFCA143DB124D6346FD7E083" box="[277,446,1388,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.56" metricValueMin="1.54" pageId="4" pageNumber="23" unit="mm" value="1.55" valueMax="1.56" valueMin="1.54">1.54–1.56 mm</quantity>
|
||
, HL
|
||
<quantity id="8C7D9B0AFFCA143DB1C7D6346CF6E083" box="[502,671,1388,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.32" metricValueMin="1.31" pageId="4" pageNumber="23" unit="mm" value="1.315" valueMax="1.32" valueMin="1.31">1.31–1.32 mm</quantity>
|
||
, EL
|
||
<quantity id="8C7D9B0AFFCA143DB2E3D6346D52E083" box="[722,827,1388,1415]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="23" unit="mm" value="0.82">0.82 mm</quantity>
|
||
, EW
|
||
<quantity id="8C7D9B0AFFCA143DB34BD6346A4AE083" box="[890,1059,1388,1415]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="4" pageNumber="23" unit="mm" value="0.615" valueMax="0.62" valueMin="0.61">0.61–0.62 mm</quantity>
|
||
, OL
|
||
<quantity id="8C7D9B0AFFCA143DB46BD6346AADE083" box="[1114,1220,1388,1415]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="4" pageNumber="23" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, OED
|
||
<quantity id="8C7D9B0AFFCA143DB0DBD6D76F3CE0AE" box="[234,341,1423,1450]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="23" unit="mm" value="0.24">0.24 mm</quantity>
|
||
, SL
|
||
<quantity id="8C7D9B0AFFCA143DB1B6D6D76F9BE0AE" box="[391,498,1423,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="23" unit="mm" value="1.17">1.17 mm</quantity>
|
||
, WL
|
||
<quantity id="8C7D9B0AFFCA143DB202D6D76CF7E0AE" box="[563,670,1423,1450]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="23" unit="mm" value="3.3">3.30 mm</quantity>
|
||
, PTH
|
||
<quantity id="8C7D9B0AFFCA143DB2DAD6D76DFFE0AE" box="[747,918,1423,1450]" metricMagnitude="-4" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.0" metricValueMin="6.8999999999999995" pageId="4" pageNumber="23" unit="mm" value="0.695" valueMax="0.7" valueMin="0.69">0.69–0.70 mm</quantity>
|
||
, PTL
|
||
<quantity id="8C7D9B0AFFCA143DB3ECD6D76A21E0AE" box="[989,1096,1423,1450]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="23" unit="mm" value="0.8">0.80 mm</quantity>
|
||
, CI 84–85, SI 88, PTHI 86–87.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="039F6564FFCA143CB097D6A16C3AE156" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23" type="discussion">
|
||
<paragraph id="4B3A36EFFFCA143DB097D6A16F7AE317" blockId="4.[166,275,1529,1555]" box="[166,275,1529,1555]" pageId="4" pageNumber="23">
|
||
<emphasis id="79F1EAFDFFCA143DB097D6A16F7AE317" bold="true" box="[166,275,1529,1555]" pageId="4" pageNumber="23">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143DB0EAD5186D7EE3A4" blockId="4.[165,1225,1599,1837]" pageId="4" pageNumber="23">
|
||
<taxonomicName id="8C854D6CFFCA143DB0EAD5186FB3E35D" authorityName="Wheeler" authorityYear="1919" box="[219,474,1599,1625]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="79F1EAFDFFCA143DB0EAD5186FB3E35D" box="[219,474,1599,1625]" italics="true" pageId="4" pageNumber="23">Leptogenys borneensis</emphasis>
|
||
</taxonomicName>
|
||
is recorded for the first time in
|
||
<collectingRegion id="8941F80DFFCA143DB360D5676DDDE35D" box="[849,948,1599,1625]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="23">Sumatra</collectingRegion>
|
||
Island. The colony was found in decayed wood stump along the hiking trails (ca.
|
||
<quantity id="8C7D9B0AFFCA143DB370D53A6DFEE379" box="[833,919,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="23" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l.). Workers and males were found but no queen was found from the colony.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCA143CB0EAD5F16FFAE414" blockId="4.[165,1225,1599,1837]" lastBlockId="5.[173,1233,175,871]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
|
||
So far, based on
|
||
<bibRefCitation id="2F144B1EFFCA143DB192D5F16C53E3C7" author="Forel, A." box="[419,570,1705,1731]" pageId="4" pageNumber="23" pagination="1 - 148" refId="ref3183" refString="Forel, A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911 - 1912. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 36, 1 - 148." type="journal article" year="1913">Forel (1913)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2F144B1EFFCA143DB277D5F16D62E3C0" author="Karavaiev, V." box="[582,779,1705,1732]" pageId="4" pageNumber="23" pagination="276 - 296" refId="ref3510" refString="Karavaiev, V. 1925. Ponerinen (Fam. Formicidae) aus dem Indo-Australischen Gebiet. (Schluss). Konowia, 4, 276 - 296. https: // doi. org / 10.5281 / zenodo. 25961" type="journal article" year="1925">Karavaiev (1925</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2F144B1EFFCA143DB326D5F16D3BE3C7" author="Karavaiev, V." box="[791,850,1705,1731]" pageId="4" pageNumber="23" pagination="413 - 445" refId="ref3551" refString="Karavaiev, V. 1926. Ameisen aus dem Indo-Australischen Gebiet. Treubia, 8, 413 - 445. https: // doi. org / 10.5281 / zenodo. 25966" type="journal article" year="1926">1926</bibRefCitation>
|
||
) and this paper, the male caste is known for the following four species and one subspecies of the genus
|
||
<taxonomicName id="8C854D6CFFCA143DB434D5956AEEE3E2" authorityName="Roger" authorityYear="1861" box="[1029,1159,1741,1766]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCA143DB434D5956AEEE3E2" box="[1029,1159,1741,1766]" italics="true" pageId="4" pageNumber="23">Leptogenys</emphasis>
|
||
</taxonomicName>
|
||
from the island of
|
||
<collectingRegion id="8941F80DFFCA143DB175D5B76FC5E20D" box="[324,428,1775,1801]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="23">Sumatra</collectingRegion>
|
||
:
|
||
<taxonomicName id="8C854D6CFFCA143DB18BD5A86DE9E20E" authority="(F. Smith, 1857)" baseAuthorityName="F. Smith" baseAuthorityYear="1857" box="[442,896,1775,1802]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="diminuta">
|
||
<emphasis id="79F1EAFDFFCA143DB18BD5A86CDBE20D" box="[442,690,1775,1801]" italics="true" pageId="4" pageNumber="23">Leptogenys diminuta</emphasis>
|
||
(F.
|
||
<bibRefCitation id="2F144B1EFFCA143DB2D2D5B76D1EE20E" author="Smith, F." box="[739,887,1775,1802]" pageId="4" pageNumber="23" pagination="42 - 88" refId="ref4451" refString="Smith, F. 1857. Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace [part]. Journal of the proceedings of the Linnean Society: Zoology, 2, 42 - 88. https: // doi. org / 10.1111 / j. 1096 - 3642.1857. tb 01759. x" type="journal article" year="1857">Smith, 1857</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="8C854D6CFFCA143DB3BCD5A86FF0E228" authority="Emery, 1896" authorityName="Emery" authorityYear="1896" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="diminuta" subSpecies="fruhstorferi">
|
||
<emphasis id="79F1EAFDFFCA143DB3BCD5A86E93E229" italics="true" pageId="4" pageNumber="23">Leptogenys diminuta fruhstorferi</emphasis>
|
||
<bibRefCitation id="2F144B1EFFCA143DB130D44B6FF0E228" author="Emery, C." box="[257,409,1810,1837]" pageId="4" pageNumber="23" pagination="245 - 249" refId="ref3096" refString="Emery, C. 1896. Formicides recoltes a Buitenzorg (Java), par M. Massart. Annales De La Societe Entomologique De Belgique, 40, 245 - 249. https: // doi. org / 10.5281 / zenodo. 25469" type="journal article" year="1896">Emery, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="8C854D6CFFCA143DB195D44B6D5BE229" authority="Forel, 1905" authorityName="Forel" authorityYear="1905" box="[420,818,1810,1837]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="kraepelini">
|
||
<emphasis id="79F1EAFDFFCA143DB195D44B6CCAE228" box="[420,675,1810,1836]" italics="true" pageId="4" pageNumber="23">Leptogenys kraepelini</emphasis>
|
||
<bibRefCitation id="2F144B1EFFCA143DB29BD44A6D5BE229" author="Forel, A." box="[682,818,1810,1837]" pageId="4" pageNumber="23" pagination="1 - 26" refId="ref3141" refString="Forel, A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Jahrbuch Der Hamburgischen Wissenschaftlichen Anstalten, 22, 1 - 26. https: // doi. org / 10.5281 / zenodo. 14415" type="journal article" year="1905">Forel, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="8C854D6CFFCA143DB30CD44B6AADE229" authority="(Emery, 1887)" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[829,1220,1811,1837]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="myops">
|
||
<emphasis id="79F1EAFDFFCA143DB30CD44B6A79E228" box="[829,1040,1811,1836]" italics="true" pageId="4" pageNumber="23">Leptogenys myops</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCA143DB42ED44B6AD2E229" author="Emery, C." box="[1055,1211,1811,1837]" pageId="4" pageNumber="23" pagination="427 - 473" refId="ref3029" refString="Emery, C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova), 5 (25), 427 - 473. https: // doi. org / 10.5281 / zenodo. 25420" type="journal article" year="1887">Emery, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="8C854D6CFFCB143CB11BD3E86C58E5CD" authorityName="Wheeler" authorityYear="1919" box="[298,561,175,201]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="79F1EAFDFFCB143CB11BD3E86C58E5CD" box="[298,561,175,201]" italics="true" pageId="5" pageNumber="24">Leptogenys borneensis</emphasis>
|
||
</taxonomicName>
|
||
in the present study. However, the morphology of the male genitalia of these species remains unknown, except for
|
||
<taxonomicName id="8C854D6CFFCB143CB3B9D38B6A4BE5E8" authorityName="Wheeler" authorityYear="1919" box="[904,1058,210,236]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="79F1EAFDFFCB143CB3B9D38B6A4BE5E8" box="[904,1058,210,236]" italics="true" pageId="5" pageNumber="24">L. borneensis</emphasis>
|
||
</taxonomicName>
|
||
, which is provided in this study.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCB143CB0D2D2416C47E647" blockId="5.[173,1233,175,871]" pageId="5" pageNumber="24">
|
||
The morphology of male genitalia provides useful characteristics to confirm and delimitate the species of ants, and to predict evolutionary trends in behavior and reproductive biology (
|
||
<bibRefCitation id="2F144B1EFFCB143CB125D2076FB9E47D" author="Boudinot, B. E." box="[276,464,351,377]" pageId="5" pageNumber="24" pagination="29 - 49" refId="ref2876" refString="Boudinot, B. E. 2013. The male genitalia of ants: musculature, homology, and functional morphology (Hymenoptera, Aculeata, Formicidae). Journal of Hymenoptera Research, 30, 29 - 49. https: // doi. org / 10.3897 / jhr. 30.3535" type="journal article" year="2013">Boudinot, 2013</bibRefCitation>
|
||
). The important characteristic of male morphology and genitalia that used for species delimitation were highlighted in the following previous studies: in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB0C9D2FE6CE2E4C4" authority="(Satria et al., 2015)" baseAuthorityName="Satria" baseAuthorityYear="2015" box="[248,651,421,448]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB0C9D2FE6FC0E4C4" box="[248,425,422,448]" italics="true" pageId="5" pageNumber="24">Odontomachus</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB186D2FE6CEAE4C4" author="Satria, R. & Kurushima, H. & Herwina, H. & Yamane, S. & Eguchi, K." box="[439,643,421,448]" pageId="5" pageNumber="24" pagination="1 - 36" refId="ref4087" refString="Satria, R., Kurushima, H., Herwina, H., Yamane, S. & Eguchi, K. 2015. The trap-jaw ant genus Odontomachus Latreille from Sumatra, with a new species description. Zootaxa, 4048, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 4048.1.1" type="journal article" year="2015">Satria et al., 2015</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB318D2FE6AC7E4C4" authority="(Yamada et al., 2020)" baseAuthorityName="Yamada" baseAuthorityYear="2020" box="[809,1198,422,448]" class="Insecta" family="Formicidae" genus="Opamyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB318D2FE6DC6E4C4" box="[809,943,422,448]" italics="true" pageId="5" pageNumber="24">Opamyrma</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB38CD2FE6ACFE4C4" author="Yamada, A. & Nguyen, D. D. & Eguchi, K." box="[957,1190,422,448]" pageId="5" pageNumber="24" pagination="27 - 52" refId="ref4913" refString="Yamada, A., Nguyen, D. D. & Eguchi, K. 2020. Unveiling the morphology of the Oriental rare monotypic ant genus Opamyrma Yamane, Bui & Eguchi, 2008 (Hymenoptera: Formicidae: Leptanillinae) and its evolutionary implications, with first descriptions of the male, larva, tentorium, and sting apparatus. Myrmecological News, 30, 27 - 52. https: // doi. org / 10.25849 / myrmecol. news _ 030: 027" type="journal article" year="2020">Yamada et al., 2020</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB110D2926C10E4E7" authority="(Wang et al., 2019)" baseAuthorityName="Wang" baseAuthorityYear="2019" box="[289,633,457,483]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB110D2926FFCE4E7" box="[289,405,458,483]" italics="true" pageId="5" pageNumber="24">Metapone</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB195D2916C18E4E7" author="Wang, W. Y. & Yamada, A. & Eguchi, K." box="[420,625,457,483]" pageId="5" pageNumber="24" pagination="125 - 141" refId="ref4814" refString="Wang, W. Y., Yamada, A. & Eguchi, K. 2019. Discovery of a new ant species of the elusive termitophilous genus Metapone in Singapore (Hymenoptera, Formicidae, Myrmicinae), with the first detailed description of male genitalia of the genus. ZooKeys, 876, 125 - 141. https: // doi. org / 10.3897 / zookeys. 876.35739" type="journal article" year="2019">Wang et al., 2019</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB322D2916A33E4E7" authority="(Wang et al., 2018)" baseAuthorityName="Wang" baseAuthorityYear="2018" box="[787,1114,457,483]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB322D2916D1FE4E7" box="[787,886,457,483]" italics="true" pageId="5" pageNumber="24">Pheidole</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB3B4D2916A3BE4E7" author="Wang, W. Y. & Yamada, A. & Eguchi, K." box="[901,1106,457,483]" pageId="5" pageNumber="24" pagination="652 - 663" refId="ref4742" refString="Wang, W. Y., Yamada, A. & Eguchi, K. 2018. First discovery of the mangrove ant Pheidole sexspinosa Mayr, 1870 (Formicidae: Myrmicinae) from the Oriental region, with redescriptions of the worker, queen and male. Raffles Bulletin of Zoology, 66, 652 - 663. https: // doi. org / 10.5281 / zenodo. 5360652" type="journal article" year="2018">Wang et al., 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB0EED2B56F03E702" authorityName="Roger" authorityYear="1861" box="[223,362,493,518]" class="Insecta" family="Formicidae" genus="Dinoponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB0EED2B56F03E702" box="[223,362,493,518]" italics="true" pageId="5" pageNumber="24">Dinoponera</emphasis>
|
||
</taxonomicName>
|
||
(Lenhart et al., 2013;
|
||
<bibRefCitation id="2F144B1EFFCB143CB25BD2B56D14E702" author="Tozetto, L. & Lattke, J. E." box="[618,893,492,519]" pageId="5" pageNumber="24" pagination="1 - 12" refId="ref4523" refString="Tozetto, L. & Lattke, J. E. 2020. Revealing male genital morphology in the giant ant genus Dinoponera with geometric morphometrics. Arthropod Structure & Development, 57, 1 - 12. https: // doi. org / 10.1016 / j. asd. 2020.100943" type="journal article" year="2020">Tozetto & Lattke, 2020</bibRefCitation>
|
||
): in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB417D2B46F80E72D" authority="(Schmidt & Heinze, 2017)" baseAuthorityName="Schmidt & Heinze" baseAuthorityYear="2017" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB417D2B46AB8E702" box="[1062,1233,492,518]" italics="true" pageId="5" pageNumber="24">Cardiocondyla</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB087D1576F88E72D" author="Schmidt, C. V. & Heinze, J." box="[182,481,527,553]" pageId="5" pageNumber="24" pagination="59 - 80" refId="ref4394" refString="Schmidt, C. V. & Heinze, J. 2017. Genital Morphology of Winged and Wingless Males in the Ant Genus Cardiocondyla (Formicidae, Myrmicinae). Insect Systematics & Evolution, 49 (1), 59 - 80. https: // doi. org / 10.1163 / 1876312 X- 48022163" type="journal article" year="2017">Schmidt & Heinze, 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In the case of the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB32CD1576DA5E72D" box="[797,972,527,553]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB32CD1576DA5E72D" box="[797,972,527,553]" italics="true" pageId="5" pageNumber="24">Odontomachus</emphasis>
|
||
</taxonomicName>
|
||
, where the taxonomy problems of
|
||
<taxonomicName id="8C854D6CFFCB143CB10ED16A6C21E748" authorityName="Smith" authorityYear="1857" box="[319,584,562,588]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="rixosus">
|
||
<emphasis id="79F1EAFDFFCB143CB10ED16A6C21E748" box="[319,584,562,588]" italics="true" pageId="5" pageNumber="24">Odontomachus rixosus</emphasis>
|
||
</taxonomicName>
|
||
complex confused the myrmecologist for decades, then it can be resolved by using the information of the morphology and genitalia of the male caste (
|
||
<bibRefCitation id="2F144B1EFFCB143CB0C8D1216FAEE790" author="Satria, R. & Kurushima, H. & Herwina, H. & Yamane, S. & Eguchi, K." box="[249,455,633,660]" pageId="5" pageNumber="24" pagination="1 - 36" refId="ref4087" refString="Satria, R., Kurushima, H., Herwina, H., Yamane, S. & Eguchi, K. 2015. The trap-jaw ant genus Odontomachus Latreille from Sumatra, with a new species description. Zootaxa, 4048, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 4048.1.1" type="journal article" year="2015">Satria et al., 2015</bibRefCitation>
|
||
). However, the study of the male genitalia in the genus
|
||
<taxonomicName id="8C854D6CFFCB143CB463D1226AB8E797" authorityName="Roger" authorityYear="1861" box="[1106,1233,634,659]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB463D1226AB8E797" box="[1106,1233,634,659]" italics="true" pageId="5" pageNumber="24">Leptogenys</emphasis>
|
||
</taxonomicName>
|
||
is still limited,
|
||
<bibRefCitation id="2F144B1EFFCB143CB166D1C56C13E7B2" author="Trager, J. C. & Johnson, C." box="[343,634,668,695]" pageId="5" pageNumber="24" pagination="29 - 34" refId="ref4575" refString="Trager, J. C. & Johnson, C. 1988. The ant genus Leptogenys (Hymenoptera: Formicidae, Ponerinae) in the United States. In: Trager, J. C., ed. Advances in myrmecology. E. J. Brill, Leiden, 29 - 34. https: // doi. org / 10.1163 / 9789004630765 _ 007" type="book chapter" year="1988">Trager & Johnson (1988)</bibRefCitation>
|
||
confirmed the status of
|
||
<taxonomicName id="8C854D6CFFCB143CB3A3D1C56F76E7DD" authority="(Buckley, 1866)" baseAuthorityName="Buckley" baseAuthorityYear="1866" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="79F1EAFDFFCB143CB3A3D1C56A13E7B2" box="[914,1146,668,694]" italics="true" pageId="5" pageNumber="24">Leptogenys elongata</emphasis>
|
||
(Buckley, 1866)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8C854D6CFFCB143CB168D1986CEAE7DD" authority="(Buckley, 1866)" baseAuthorityName="Buckley" baseAuthorityYear="1866" box="[345,643,703,729]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="manni">
|
||
<emphasis id="79F1EAFDFFCB143CB168D1986FAAE7DD" box="[345,451,704,729]" italics="true" pageId="5" pageNumber="24">L. manni</emphasis>
|
||
(Buckley, 1866)
|
||
</taxonomicName>
|
||
. They stated that the study of male genitalia is important in future revisionary work on
|
||
<taxonomicName id="8C854D6CFFCB143CB244D1BB6A4EE7F9" authority="(Trager & Johnson 1988)" baseAuthorityName="Trager & Johnson" baseAuthorityYear="1988" box="[629,1063,738,765]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="79F1EAFDFFCB143CB244D1BB6C9DE7F8" box="[629,756,739,764]" italics="true" pageId="5" pageNumber="24">Leptogenys</emphasis>
|
||
(
|
||
<bibRefCitation id="2F144B1EFFCB143CB333D1BB6A77E7F9" author="Trager, J. C. & Johnson, C." box="[770,1054,738,765]" pageId="5" pageNumber="24" pagination="29 - 34" refId="ref4575" refString="Trager, J. C. & Johnson, C. 1988. The ant genus Leptogenys (Hymenoptera: Formicidae, Ponerinae) in the United States. In: Trager, J. C., ed. Advances in myrmecology. E. J. Brill, Leiden, 29 - 34. https: // doi. org / 10.1163 / 9789004630765 _ 007" type="book chapter" year="1988">Trager & Johnson 1988</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="2F144B1EFFCB143CB403D1BA6AB9E7F9" author="Lattke, E. J." box="[1074,1232,738,765]" pageId="5" pageNumber="24" pagination="127 - 264" refId="ref3669" refString="Lattke, E. J. 2011. Revision of the New World species of the genus Leptogenys Roger (Insecta: Hymenoptera: Formicidae: Ponerinae). Arthropod Systematics & Phylogeny, 69 (3), 127 - 264. https: // doi. org / 10.3897 / asp. 69. e 31744" type="journal article" year="2011">Lattke (2011)</bibRefCitation>
|
||
used male morphology as part of his argument for synonymising
|
||
<taxonomicName id="8C854D6CFFCB143CB395D05E6AB9E624" authority="(Mayr, 1870)" baseAuthorityName="Mayr" baseAuthorityYear="1870" box="[932,1232,774,800]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="79F1EAFDFFCB143CB395D05E6A58E61B" box="[932,1073,774,799]" italics="true" pageId="5" pageNumber="24">L. mexicana</emphasis>
|
||
(Mayr, 1870)
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="8C854D6CFFCB143CB0D8D0726C40E647" authority="(Buckley, 1866)" baseAuthorityName="Buckley" baseAuthorityYear="1866" box="[233,553,809,835]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="79F1EAFDFFCB143CB0D8D0726F00E647" box="[233,361,809,835]" italics="true" pageId="5" pageNumber="24">L. elongata</emphasis>
|
||
(Buckley, 1866)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCB143CB0D2D0156AFBE662" blockId="5.[173,1233,175,871]" box="[227,1170,844,871]" pageId="5" pageNumber="24">
|
||
D i s t r i b u t i o n.
|
||
<collectingCountry id="3392767FFFCB143CB194D0146C74E662" box="[421,541,844,870]" name="Indonesia" pageId="5" pageNumber="24">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="8941F80DFFCB143CB217D0146CE0E662" box="[550,649,844,870]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="24">Sumatra</collectingRegion>
|
||
(new record);
|
||
<collectingCountry id="3392767FFFCB143CB307D0146DC8E662" box="[822,929,844,870]" name="Malaysia" pageId="5" pageNumber="24">Malaysia</collectingCountry>
|
||
;
|
||
<collectingCountry id="3392767FFFCB143CB39DD0146A70E662" box="[940,1049,844,870]" name="Thailand" pageId="5" pageNumber="24">Thailand</collectingCountry>
|
||
;
|
||
<collectingCountry id="3392767FFFCB143CB415D0146AE7E662" box="[1060,1166,844,870]" name="Vietnam" pageId="5" pageNumber="24">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCB143CB09FD0D76F1BE6A1" blockId="5.[174,370,911,933]" box="[174,370,911,933]" pageId="5" pageNumber="24">
|
||
<emphasis id="79F1EAFDFFCB143CB09FD0D76F1BE6A1" bold="true" box="[174,370,911,933]" pageId="5" pageNumber="24">Acknowledgements</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4B3A36EFFFCB143CB0D2D0926C3AE156" blockId="5.[173,1234,969,1106]" pageId="5" pageNumber="24">
|
||
We wish to thank Mr. Rudi Yuliandri, Ms. Nur Aqsha, Ms. Putri Ratna Desi, Mr. Niko, and Mr. Rezha for their helps in field research. This research was financed mostly by PNBP Research Grants of Universitas Negeri Padang for Biodiversity of
|
||
<collectingRegion id="8941F80DFFCB143CB1C4D75B6C2FE11C" box="[501,582,1027,1048]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="24">Sumatra</collectingRegion>
|
||
Research Group under contract No. 1750/UN35.13/LT/2022 (Team Leader: Rijal Satria), and partly by
|
||
<collectingRegion id="8941F80DFFCB143CB1CCD7786C52E132" box="[509,571,1056,1078]" country="Japan" name="Tokyo" pageId="5" pageNumber="24">Tokyo</collectingRegion>
|
||
Metropolitan University Fund for TMU Strategic Research (Leader: Prof. Noriaki Murakami; FY2020–FY2022).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |