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<document id="EADB68BAF1097D8694F9E58A9E53400C" ID-CLB-Dataset="23655" ID-DOI="10.11646/zootaxa.4748.3.2" ID-GBIF-Dataset="d93b19d0-7f75-4fee-8cd3-6bb668651d09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3701023" ID-ZooBank="58F8C4B2-C336-461B-A7C0-FD9CA547F1AA" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="admin" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1583739254815" checkinUser="plazi" docAuthor="Cruz-López, Jesús A. &amp; Francke, Oscar F." docDate="2020" docId="837AAD42FFA26A5459A4FA6EFCB3FED0" docLanguage="en" docName="zootaxa.4783.3.2.pdf" docOrigin="Zootaxa 4748 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guelaguetzia Cruz-López &amp; Francke 2020, gen. nov." docType="treatment" docVersion="10" lastPageNumber="440" masterDocId="7F43D53AFFA46A5D5933FFBBFF89FFCE" masterDocTitle="Two new genera of epigean harvestmen (Opiliones, Stygnopsidae) from Oaxaca, Mexico, with an identification key for the stygnopsine genera" masterLastPageNumber="454" masterPageNumber="431" pageNumber="437" updateTime="1698726022374" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7068E18A5FD948E4FC72723EE8C9F0B2">Two new genera of epigean harvestmen (Opiliones, Stygnopsidae) from Oaxaca, Mexico, with an identification key for the stygnopsine genera</mods:title>
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<mods:namePart id="824911DA8D98063A8D3C149AAAD371F4">Cruz-López, Jesús A.</mods:namePart>
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<mods:namePart id="BE0F44B7367C1360E1BA49436FE289AC">Francke, Oscar F.</mods:namePart>
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<mods:title id="812110989E879F39FB763E492BBAAB55">Zootaxa</mods:title>
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<mods:date id="93C134D7030CCE84A13F7AF9BF93D7A0">2020</mods:date>
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<mods:number id="0EF67F948869F1F2165DAE97FDC92CB4">2020-03-09</mods:number>
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<treatment id="837AAD42FFA26A5459A4FA6EFCB3FED0" ID-DOI="http://doi.org/10.5281/zenodo.3705606" ID-GBIF-Taxon="162467135" ID-Zenodo-Dep="3705606" LSID="urn:lsid:plazi:treatment:837AAD42FFA26A5459A4FA6EFCB3FED0" httpUri="http://treatment.plazi.org/id/837AAD42FFA26A5459A4FA6EFCB3FED0" lastPageId="9" lastPageNumber="440" pageId="6" pageNumber="437">
<subSubSection id="43C94FDFFFA26A5B59A4FA6EFC47F9DA" pageId="6" pageNumber="437" type="nomenclature">
<paragraph id="0B6C1C54FFA26A5B59A4FA6EFE25FA3E" blockId="6.[151,974,1493,1556]" box="[151,428,1493,1520]" pageId="6" pageNumber="437">
<heading id="5024AB38FFA26A5B59A4FA6EFE25FA3E" bold="true" box="[151,428,1493,1520]" fontSize="11" level="1" pageId="6" pageNumber="437" reason="1">
<emphasis id="39A7C046FFA26A5B59A4FA6EFE25FA3E" bold="true" box="[151,428,1493,1520]" pageId="6" pageNumber="437">
<taxonomicName id="CCD367D7FFA26A5B59A4FA6EFEBEFA3E" ID-CoL="BF59K" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[151,311,1493,1520]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B59A4FA6EFEBEFA3E" bold="true" box="[151,311,1493,1520]" italics="true" pageId="6" pageNumber="437">Guelaguetzia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22947D3DFFA26A5B580DFA6DFE25FA3E" box="[318,428,1494,1520]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<uri id="7F421056FFA26A5B59A4FA42FC47F9DA" box="[151,974,1529,1556]" pageId="6" pageNumber="437">
urn:lsid:zoobank.org:act:
<uuid id="7F752681FFA26A5B5899FA42FC47F9DA" box="[426,974,1529,1556]" pageId="6" pageNumber="437">D68B5C50-6145-4D8F-B620-A505A6A4FEA6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="43C94FDFFFA26A5B59A4F985FC30F998" box="[151,953,1597,1622]" pageId="6" pageNumber="437" type="type_taxon">
<paragraph id="0B6C1C54FFA26A5B59A4F985FC30F998" blockId="6.[151,953,1597,1622]" box="[151,953,1597,1622]" pageId="6" pageNumber="437">
<emphasis id="39A7C046FFA26A5B59A4F985FEAAF998" bold="true" box="[151,291,1597,1622]" pageId="6" pageNumber="437">
<typeStatus id="D468A2F6FFA26A5B59A4F985FF45F998" box="[151,204,1598,1622]" pageId="6" pageNumber="437">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="CCD367D7FFA26A5B5819F985FD91F998" ID-CoL="BFQYD" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[298,536,1598,1622]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="species" species="cuicateca" status="sp. nov.">
<emphasis id="39A7C046FFA26A5B5819F985FD91F998" box="[298,536,1598,1622]" italics="true" pageId="6" pageNumber="437">Guelaguetzia cuicateca</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B5B2CF985FDE6F998" bold="true" box="[543,623,1598,1622]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B5B2CF985FDE6F998" box="[543,623,1598,1622]" pageId="6" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, designated in the present work.
</paragraph>
</subSubSection>
<subSubSection id="43C94FDFFFA26A5B59A4F93EFB43F928" pageId="6" pageNumber="437" type="etymology">
<paragraph id="0B6C1C54FFA26A5B59A4F93EFB43F928" blockId="6.[151,1437,1668,2019]" pageId="6" pageNumber="437">
<emphasis id="39A7C046FFA26A5B59A4F93EFE97F951" bold="true" box="[151,286,1669,1695]" pageId="6" pageNumber="437">Etymology.</emphasis>
The word ‘Guelaguetza’ comes from the Zapotec language ‘
<emphasis id="39A7C046FFA26A5B5AD4F93FFBF9F950" box="[999,1136,1668,1694]" italics="true" pageId="6" pageNumber="437">guendalizaa</emphasis>
’, meaning cooperating or collaborating. In addition, the Guelaguetza is an annual festivity held every July in the state of
<collectingRegion id="C917D2B6FFA26A5B5DF2F913FA91F90C" box="[1217,1304,1704,1730]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="437">Oaxaca</collectingRegion>
,
<collectingCountry id="73C45CC4FFA26A5B5C10F913FAF2F90C" box="[1315,1403,1704,1730]" name="Mexico" pageId="6" pageNumber="437">Mexico</collectingCountry>
, in which they carry out many cultural and gastronomic exhibitions. Grammatical gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="43C94FDFFFA26A5459F4F954FCB3FED0" lastPageId="9" lastPageNumber="440" pageId="6" pageNumber="437" type="diagnosis">
<paragraph id="0B6C1C54FFA26A5559F4F954FE3BFD8E" blockId="6.[151,1437,1668,2019]" lastBlockId="8.[151,1437,152,611]" lastPageId="8" lastPageNumber="439" pageId="6" pageNumber="437">
<emphasis id="39A7C046FFA26A5B59F4F954FECBF8C4" bold="true" box="[199,322,1775,1802]" pageId="6" pageNumber="437">Diagnosis.</emphasis>
<taxonomicName id="CCD367D7FFA26A5B587BF94BFE54F8C4" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[328,477,1776,1802]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B587BF94BFE54F8C4" box="[328,477,1776,1802]" italics="true" pageId="6" pageNumber="437">Guelaguetzia</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B58D6F94AFDC7F8C5" bold="true" box="[485,590,1777,1803]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B58D6F94AFDC7F8C5" box="[485,590,1777,1803]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be recognized from the non-troglobitic stygnopsine genera without lateral clear areas on mid-bulge scutum (
<taxonomicName id="CCD367D7FFA26A5B5B75F8AFFD52F8E0" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[582,731,1812,1838]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B5B75F8AFFD52F8E0" box="[582,731,1812,1838]" italics="true" pageId="6" pageNumber="437">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B5BD1F8AEFCC5F8E1" bold="true" box="[738,844,1813,1839]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B5BD1F8AEFCC5F8E1" box="[738,844,1813,1839]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="CCD367D7FFA26A5B5A69F8AFFC6AF8E0" authorityName="Banks" authorityYear="1900" box="[858,995,1812,1838]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5A69F8AFFC6AF8E0" box="[858,995,1812,1838]" italics="true" pageId="6" pageNumber="437">Hoplobunus</emphasis>
</taxonomicName>
and
<taxonomicName id="CCD367D7FFA26A5B5D29F8AEFBEBF8E0" authorityName="Sorensen" authorityYear="1932" box="[1050,1122,1813,1838]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5D29F8AEFBEBF8E0" box="[1050,1122,1813,1838]" italics="true" pageId="6" pageNumber="437">Isaeus</emphasis>
</taxonomicName>
) by the following combination of characters: (a) ocularium apically rounded, without spine (
<figureCitation id="93E800D1FFA26A5B5A4EF883FC52F89C" box="[893,987,1848,1874]" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1975,2000]" captionTargetBox="[151,1435,273,1950]" captionTargetId="figure@10.[151,1435,273,1950]" captionTargetPageId="10" captionText="FIGURES 19–21. Guelaguetzia cuicateca sp. nov., holotype male (CNAN-T01330) and paratype female (CNAN-T01331). 19, habitus holotype, dorsal view. 20, habitus female, dorsal view. 21, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701037" httpUri="https://zenodo.org/record/3701037/files/figure.png" pageId="6" pageNumber="437">Figs. 21</figureCitation>
,
<figureCitation id="93E800D1FFA26A5B5AD4F883FB8BF89D" box="[999,1026,1848,1875]" captionStart="FIGURES 37–39" captionStartId="15.[151,264,1932,1957]" captionTargetBox="[151,1435,181,1907]" captionTargetId="figure@15.[151,1435,181,1907]" captionTargetPageId="15" captionText="FIGURES 37–39. Guelaguetzia serrana sp. nov., holotype male (CNAN-T01334) and paratype female (CNAN-T01335). 37, habitus holotype, dorsal view. 38, habitus female, dorsal view. 39, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701049" httpUri="https://zenodo.org/record/3701049/files/figure.png" pageId="6" pageNumber="437">39</figureCitation>
)—spine small in
<taxonomicName id="CCD367D7FFA26A5B5DFEF883FADFF89C" authorityName="Banks" authorityYear="1900" box="[1229,1366,1848,1874]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5DFEF883FADFF89C" box="[1229,1366,1848,1874]" italics="true" pageId="6" pageNumber="437">Hoplobunus</emphasis>
</taxonomicName>
, large in
<taxonomicName id="CCD367D7FFA26A5B5980F8E6FF72F8B8" authorityName="Sorensen" authorityYear="1932" box="[179,251,1885,1910]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5980F8E6FF72F8B8" box="[179,251,1885,1910]" italics="true" pageId="6" pageNumber="437">Isaeus</emphasis>
</taxonomicName>
and absent in
<taxonomicName id="CCD367D7FFA26A5B58ABF8E7FDA4F8B8" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[408,557,1884,1910]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B58ABF8E7FDA4F8B8" box="[408,557,1884,1910]" italics="true" pageId="6" pageNumber="437">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B5B00F8E6FD12F8B9" bold="true" box="[563,667,1885,1911]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B5B00F8E6FD12F8B9" box="[563,667,1885,1911]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
; (b) pedipalpal patella dorsally covered by small spiniform tubercles, without dorso-apical apophysis (
<figureCitation id="93E800D1FFA26A5B5B34F83BFDECF854" box="[519,613,1920,1946]" captionStart="FIGURES 24–27" captionStartId="12.[151,264,1944,1969]" captionTargetBox="[270,1317,181,1919]" captionTargetId="figure@12.[270,1317,181,1919]" captionTargetPageId="12" captionText="FIGURES 24–27. Guelaguetzia cuicateca sp. nov., holotype male (CNAN-T01330). 24, chelicera, frontal view. 25, chelicera, mesal view. 26, pedipalp, mesal view. 27, pedipalpal tibia and tarsus, ventral view (major spiniform setiferous tubercles on tibia indicated as T1–T3). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701041" httpUri="https://zenodo.org/record/3701041/files/figure.png" pageId="6" pageNumber="437">Figs. 26</figureCitation>
,
<figureCitation id="93E800D1FFA26A5B5B41F83BFD07F855" box="[626,654,1920,1947]" captionStart="FIGURES 42–45" captionStartId="18.[151,264,1945,1970]" captionTargetBox="[176,1410,181,1920]" captionTargetId="figure@18.[176,1410,181,1920]" captionTargetPageId="18" captionText="FIGURES 42–45. Guelaguetzia serrana sp. nov., holotype male (CNAN-T01334). 42, chelicera, frontal view. 43, chelicera, mesal view. 44, pedipalp, mesal view. 45, pedipalpal tibia and tarsus, ventral view (major spiniform setiferous tubercles on tibia indicated as T1–T3). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701053" httpUri="https://zenodo.org/record/3701053/files/figure.png" pageId="6" pageNumber="437">44</figureCitation>
)—apophysis is present in both
<taxonomicName id="CCD367D7FFA26A5B5AC5F83BFBF6F854" authorityName="Banks" authorityYear="1900" box="[1014,1151,1920,1946]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5AC5F83BFBF6F854" box="[1014,1151,1920,1946]" italics="true" pageId="6" pageNumber="437">Hoplobunus</emphasis>
</taxonomicName>
and
<taxonomicName id="CCD367D7FFA26A5B5D84F83AFB76F854" authorityName="Sorensen" authorityYear="1932" box="[1207,1279,1921,1946]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5D84F83AFB76F854" box="[1207,1279,1921,1946]" italics="true" pageId="6" pageNumber="437">Isaeus</emphasis>
</taxonomicName>
and absent in
<taxonomicName id="CCD367D7FFA26A5B59A4F81FFEA5F870" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[151,300,1956,1982]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B59A4F81FFEA5F870" box="[151,300,1956,1982]" italics="true" pageId="6" pageNumber="437">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B5801F81EFE13F871" bold="true" box="[306,410,1957,1983]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B5801F81EFE13F871" box="[306,410,1957,1983]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
; (c) coxa IV visible in dorsal view, protruding (
<figureCitation id="93E800D1FFA26A5B5A97F81FFB89F870" box="[932,1024,1956,1982]" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1975,2000]" captionTargetBox="[151,1435,273,1950]" captionTargetId="figure@10.[151,1435,273,1950]" captionTargetPageId="10" captionText="FIGURES 19–21. Guelaguetzia cuicateca sp. nov., holotype male (CNAN-T01330) and paratype female (CNAN-T01331). 19, habitus holotype, dorsal view. 20, habitus female, dorsal view. 21, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701037" httpUri="https://zenodo.org/record/3701037/files/figure.png" pageId="6" pageNumber="437">Figs. 19</figureCitation>
,
<figureCitation id="93E800D1FFA26A5B5D38F81FFBA3F870" box="[1035,1066,1956,1982]" captionStart="FIGURES 37–39" captionStartId="15.[151,264,1932,1957]" captionTargetBox="[151,1435,181,1907]" captionTargetId="figure@15.[151,1435,181,1907]" captionTargetPageId="15" captionText="FIGURES 37–39. Guelaguetzia serrana sp. nov., holotype male (CNAN-T01334) and paratype female (CNAN-T01335). 37, habitus holotype, dorsal view. 38, habitus female, dorsal view. 39, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701049" httpUri="https://zenodo.org/record/3701049/files/figure.png" pageId="6" pageNumber="437">37</figureCitation>
), trochanter III not globose (
<figureCitation id="93E800D1FFA26A5B5C57F81FFF3CF82C" captionStart="FIGURES 19–21" captionStartId="10.[151,264,1975,2000]" captionTargetBox="[151,1435,273,1950]" captionTargetId="figure@10.[151,1435,273,1950]" captionTargetPageId="10" captionText="FIGURES 19–21. Guelaguetzia cuicateca sp. nov., holotype male (CNAN-T01330) and paratype female (CNAN-T01331). 19, habitus holotype, dorsal view. 20, habitus female, dorsal view. 21, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701037" httpUri="https://zenodo.org/record/3701037/files/figure.png" pageId="6" pageNumber="437">Figs. 19</figureCitation>
,
<figureCitation id="93E800D1FFA26A5B59F1F873FF54F82D" box="[194,221,1992,2019]" captionStart="FIGURES 37–39" captionStartId="15.[151,264,1932,1957]" captionTargetBox="[151,1435,181,1907]" captionTargetId="figure@15.[151,1435,181,1907]" captionTargetPageId="15" captionText="FIGURES 37–39. Guelaguetzia serrana sp. nov., holotype male (CNAN-T01334) and paratype female (CNAN-T01335). 37, habitus holotype, dorsal view. 38, habitus female, dorsal view. 39, lateral view holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701049" httpUri="https://zenodo.org/record/3701049/files/figure.png" pageId="6" pageNumber="437">37</figureCitation>
)—globose in
<taxonomicName id="CCD367D7FFA26A5B584CF873FD81F82C" authorityName="Banks" authorityYear="1900" box="[383,520,1992,2018]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B584CF873FD81F82C" box="[383,520,1992,2018]" italics="true" pageId="6" pageNumber="437">Hoplobunus</emphasis>
</taxonomicName>
and
<taxonomicName id="CCD367D7FFA26A5B5B0CF872FD0EF82C" authorityName="Sorensen" authorityYear="1932" box="[575,647,1993,2018]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFA26A5B5B0CF872FD0EF82C" box="[575,647,1993,2018]" italics="true" pageId="6" pageNumber="437">Isaeus</emphasis>
</taxonomicName>
, not globose in
<taxonomicName id="CCD367D7FFA26A5B5A0FF873FC58F82C" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[828,977,1992,2018]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="437" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFA26A5B5A0FF873FC58F82C" box="[828,977,1992,2018]" italics="true" pageId="6" pageNumber="437">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA26A5B5AEAF872FBCAF82D" bold="true" box="[985,1091,1993,2019]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="22947D3DFFA26A5B5AEAF872FBCAF82D" box="[985,1091,1993,2019]" pageId="6" pageNumber="437" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
; (d) ventral retrolateral row of tubercles on femur IV spiniform and noticeably larger than those on prolateral row (
<figureCitation id="93E800D1FFAC6A555D02FF22FB05FF7D" box="[1073,1164,153,179]" captionStart="FIGURES 28–33" captionStartId="13.[151,264,1172,1197]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure@13.[151,1435,181,1147]" captionTargetPageId="13" captionText="FIGURES 28–33. Guelaguetzia cuicateca sp. nov., 28–32: holotype male (CNAN-T01330). 28, leg IV, ectal view. 29, leg IV, mesal view. 30, trochanter IV, dorsal view. 31, trochanter IV, ventral view. 32, femur IV, ventral view. 33, paratype female (CNAN-T01331), femur IV, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701043" httpUri="https://zenodo.org/record/3701043/files/figure.png" pageId="8" pageNumber="439">Figs. 32</figureCitation>
,
<figureCitation id="93E800D1FFAC6A555DA4FF23FB3BFF7D" box="[1175,1202,152,179]" captionStart="FIGURES 46–51" captionStartId="19.[151,264,1177,1202]" captionTargetBox="[151,1435,181,1151]" captionTargetId="figure@19.[151,1435,181,1151]" captionTargetPageId="19" captionText="FIGURES 46–51. Guelaguetzia serrana sp. nov., 46–50: holotype male (CNAN-T01334). 46, leg IV, ectal view. 47, leg IV, mesal view. 48, trochanter IV, dorsal view. 49, trochanter IV, ventral view. 50, femur IV, ventral view. 51, paratype female (CNAN-T01335), femur IV ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701055" httpUri="https://zenodo.org/record/3701055/files/figure.png" pageId="8" pageNumber="439">50</figureCitation>
)—both rows formed by small spiniform tubercles in
<taxonomicName id="CCD367D7FFAC6A5558C4FF05FDB6FF19" authorityName="Sorensen" authorityYear="1932" box="[503,575,190,215]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A5558C4FF05FDB6FF19" box="[503,575,190,215]" italics="true" pageId="8" pageNumber="439">Isaeus</emphasis>
</taxonomicName>
, tooth-shaped tubercles in
<taxonomicName id="CCD367D7FFAC6A555A5AFF06FC77FF19" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[873,1022,189,215]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFAC6A555A5AFF06FC77FF19" box="[873,1022,189,215]" italics="true" pageId="8" pageNumber="439">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555D36FF06FBE4FF19" bold="true" box="[1029,1133,189,215]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555D36FF06FBE4FF19" box="[1029,1133,189,215]" pageId="8" pageNumber="439" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and tubercles increasing in size apically and cylindrical shaped in
<taxonomicName id="CCD367D7FFAC6A555B7DFF5AFD5EFF35" authorityName="Banks" authorityYear="1900" box="[590,727,225,251]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555B7DFF5AFD5EFF35" box="[590,727,225,251]" italics="true" pageId="8" pageNumber="439">Hoplobunus</emphasis>
</taxonomicName>
; (e) patella IV with ventral and ventro-apical apophyses (
<figureCitation id="93E800D1FFAC6A555C57FF5AFF3CFED1" captionStart="FIGURES 28–33" captionStartId="13.[151,264,1172,1197]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure@13.[151,1435,181,1147]" captionTargetPageId="13" captionText="FIGURES 28–33. Guelaguetzia cuicateca sp. nov., 28–32: holotype male (CNAN-T01330). 28, leg IV, ectal view. 29, leg IV, mesal view. 30, trochanter IV, dorsal view. 31, trochanter IV, ventral view. 32, femur IV, ventral view. 33, paratype female (CNAN-T01331), femur IV, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701043" httpUri="https://zenodo.org/record/3701043/files/figure.png" pageId="8" pageNumber="439">Figs. 32</figureCitation>
,
<figureCitation id="93E800D1FFAC6A5559F2FEBFFF54FED1" box="[193,221,260,287]" captionStart="FIGURES 46–51" captionStartId="19.[151,264,1177,1202]" captionTargetBox="[151,1435,181,1151]" captionTargetId="figure@19.[151,1435,181,1151]" captionTargetPageId="19" captionText="FIGURES 46–51. Guelaguetzia serrana sp. nov., 46–50: holotype male (CNAN-T01334). 46, leg IV, ectal view. 47, leg IV, mesal view. 48, trochanter IV, dorsal view. 49, trochanter IV, ventral view. 50, femur IV, ventral view. 51, paratype female (CNAN-T01335), femur IV ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3701055" httpUri="https://zenodo.org/record/3701055/files/figure.png" pageId="8" pageNumber="439">50</figureCitation>
)—no apophyses in
<taxonomicName id="CCD367D7FFAC6A55588CFEBEFDDDFED1" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[447,596,261,287]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFAC6A55588CFEBEFDDDFED1" box="[447,596,261,287]" italics="true" pageId="8" pageNumber="439">Ampliphallus</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555B68FEBEFD4DFED1" bold="true" box="[603,708,261,287]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555B68FEBEFD4DFED1" box="[603,708,261,287]" pageId="8" pageNumber="439" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="CCD367D7FFAC6A555BE0FEBEFCD5FED1" authorityName="Banks" authorityYear="1900" box="[723,860,261,287]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555BE0FEBEFCD5FED1" box="[723,860,261,287]" italics="true" pageId="8" pageNumber="439">Hoplobunus</emphasis>
</taxonomicName>
and
<emphasis id="39A7C046FFAC6A555AA1FEBDFC68FED1" box="[914,993,262,287]" italics="true" pageId="8" pageNumber="439">
<taxonomicName id="CCD367D7FFAC6A555AA1FEBDFC55FED1" authorityName="Sorensen" authorityYear="1932" box="[914,988,262,287]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">Isaeus</taxonomicName>
,
</emphasis>
only with one ventro-apical apophysis; and (f) by the male genitalia, pars distalis compressed laterally but not curved dorsally and not embracing more than the second half of penis as in
<taxonomicName id="CCD367D7FFAC6A555B19FEF6FD3AFEA9" authorityName="Banks" authorityYear="1900" box="[554,691,333,359]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555B19FEF6FD3AFEA9" box="[554,691,333,359]" italics="true" pageId="8" pageNumber="439">Hoplobunus</emphasis>
</taxonomicName>
and
<taxonomicName id="CCD367D7FFAC6A555BDEFEF5FCBCFEA9" authorityName="Sorensen" authorityYear="1932" box="[749,821,334,359]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555BDEFEF5FCBCFEA9" box="[749,821,334,359]" italics="true" pageId="8" pageNumber="439">Isaeus</emphasis>
</taxonomicName>
, with the flimsy lamina large (
<figureCitation id="93E800D1FFAC6A555DAAFEF6FAAAFEA9" box="[1177,1315,333,359]" captionStart="FIGURES 34–36" captionStartId="14.[151,264,1182,1207]" captionTargetBox="[151,1435,181,1157]" captionTargetId="figure@14.[151,1435,181,1157]" captionTargetPageId="14" captionText="FIGURES 34–36. Guelaguetzia cuicateca sp. nov., penis of holotype male (CNAN-T01330). 16, dorsal. 17, lateral. 18, ventral. Macrosetae groups: yellow=A, blue=B, pink=C, green=D, white arrows=remnant sockets of macrosetae E. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3701045" httpUri="https://zenodo.org/record/3701045/files/figure.png" pageId="8" pageNumber="439">Figs. 34–36</figureCitation>
,
<figureCitation id="93E800D1FFAC6A555C03FEF7FAF3FEA9" box="[1328,1402,332,359]" captionStart="FIGURES 52–54" captionStartId="20.[151,264,1377,1402]" captionTargetBox="[151,1435,381,1352]" captionTargetId="figure@20.[151,1435,381,1352]" captionTargetPageId="20" captionText="FIGURES 52–54. Guelaguetzia serrana sp. nov., penis of holotype male (CNAN-T01334). 52, dorsal. 53, lateral. 54, ventral. Macrosetae groups: yellow=A, blue=B, pink=C, green=D, white arrows=remnant sockets of macrosetae E. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3701057" httpUri="https://zenodo.org/record/3701057/files/figure.png" pageId="8" pageNumber="439">52–54</figureCitation>
); a pair of imperceptible remnant macrosetae D (two small pairs in
<taxonomicName id="CCD367D7FFAC6A555A40FECAFC75FE45" authorityName="Banks" authorityYear="1900" box="[883,1020,369,395]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555A40FECAFC75FE45" box="[883,1020,369,395]" italics="true" pageId="8" pageNumber="439">Hoplobunus</emphasis>
</taxonomicName>
and
<taxonomicName id="CCD367D7FFAC6A555D05FEC9FBF7FE45" authorityName="Sorensen" authorityYear="1932" box="[1078,1150,370,395]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555D05FEC9FBF7FE45" box="[1078,1150,370,395]" italics="true" pageId="8" pageNumber="439">Isaeus</emphasis>
</taxonomicName>
), and with only a pair of remnant sockets of macrosetae E (a pair of small setae in
<taxonomicName id="CCD367D7FFAC6A555A29FE2EFC2AFE61" authorityName="Banks" authorityYear="1900" box="[794,931,405,431]" class="Arachnida" family="Phalangodidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555A29FE2EFC2AFE61" box="[794,931,405,431]" italics="true" pageId="8" pageNumber="439">Hoplobunus</emphasis>
</taxonomicName>
, completely absent in
<taxonomicName id="CCD367D7FFAC6A555DACFE2DFB6EFE61" authorityName="Sorensen" authorityYear="1932" box="[1183,1255,406,431]" class="Arachnida" family="Phalangodidae" genus="Isaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="39A7C046FFAC6A555DACFE2DFB6EFE61" box="[1183,1255,406,431]" italics="true" pageId="8" pageNumber="439">Isaeus</emphasis>
</taxonomicName>
), macrosetae A, B and C large and filiform; follis with two ventral lobes (
<figureCitation id="93E800D1FFAC6A555A16FE02FC26FE1D" box="[805,943,441,467]" captionStart="FIGURES 34–36" captionStartId="14.[151,264,1182,1207]" captionTargetBox="[151,1435,181,1157]" captionTargetId="figure@14.[151,1435,181,1157]" captionTargetPageId="14" captionText="FIGURES 34–36. Guelaguetzia cuicateca sp. nov., penis of holotype male (CNAN-T01330). 16, dorsal. 17, lateral. 18, ventral. Macrosetae groups: yellow=A, blue=B, pink=C, green=D, white arrows=remnant sockets of macrosetae E. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3701045" httpUri="https://zenodo.org/record/3701045/files/figure.png" pageId="8" pageNumber="439">Figs. 34–36</figureCitation>
,
<figureCitation id="93E800D1FFAC6A555A88FE03FB8CFE1D" box="[955,1029,440,467]" captionStart="FIGURES 52–54" captionStartId="20.[151,264,1377,1402]" captionTargetBox="[151,1435,381,1352]" captionTargetId="figure@20.[151,1435,381,1352]" captionTargetPageId="20" captionText="FIGURES 52–54. Guelaguetzia serrana sp. nov., penis of holotype male (CNAN-T01334). 52, dorsal. 53, lateral. 54, ventral. Macrosetae groups: yellow=A, blue=B, pink=C, green=D, white arrows=remnant sockets of macrosetae E. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3701057" httpUri="https://zenodo.org/record/3701057/files/figure.png" pageId="8" pageNumber="439">52–54</figureCitation>
). Additionally, the male genitalia of
<taxonomicName id="CCD367D7FFAC6A5559A4FE66FEA5FE39" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[151,300,477,503]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39A7C046FFAC6A5559A4FE66FEA5FE39" box="[151,300,477,503]" italics="true" pageId="8" pageNumber="439">Guelaguetzia</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555801FE66FE13FE39" bold="true" box="[306,410,477,503]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555801FE66FE13FE39" box="[306,410,477,503]" pageId="8" pageNumber="439" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is quite similar compared to those on the troglobitic
<emphasis id="39A7C046FFAC6A555AEEFE66FBC3FE39" box="[989,1098,477,503]" italics="true" pageId="8" pageNumber="439">Tonalteca</emphasis>
<bibRefCitation id="6F4261A5FFAC6A555D63FE66FA11FE39" author="Cruz-Lopez, J. A. &amp; Francke, O. F." box="[1104,1432,477,503]" pageId="8" pageNumber="439" pagination="317 - 360" refId="ref11330" refString="Cruz-Lopez, J. A. &amp; Francke, O. F. (2017) Total evidence phylogeny of the North American harvestman family Stygnopsidae (Opiliones: Laniatores: Grassatores) reveals hidden diversity. Invertebrate Systematics, 31, 317 - 360. https: // doi. org / 10.1071 / IS 16053" type="journal article" year="2017">Cruz-López &amp; Francke, 2017</bibRefCitation>
, but these two genera can be differentiated by the external morphology due to the troglomorphisms of
<emphasis id="39A7C046FFAC6A555DCFFDBAFAE0FDD5" box="[1276,1385,513,539]" italics="true" pageId="8" pageNumber="439">Tonalteca</emphasis>
(see dichotomous key above).
</paragraph>
<caption id="5FAC4CDCFFA36A5A59A4F821FDD0F834" ID-DOI="http://doi.org/10.5281/zenodo.3701029" ID-Zenodo-Dep="3701029" httpUri="https://zenodo.org/record/3701029/files/figure.png" pageId="7" pageNumber="438" startId="7.[151,264,1946,1971]" targetBox="[222,1366,181,1920]" targetPageId="7">
<paragraph id="0B6C1C54FFA36A5A59A4F821FDD0F834" blockId="7.[151,1436,1946,2043]" pageId="7" pageNumber="438">
<emphasis id="39A7C046FFA36A5A59A4F821FEC1F87D" bold="true" box="[151,328,1946,1971]" pageId="7" pageNumber="438">FIGURES 7–10.</emphasis>
<taxonomicName id="CCD367D7FFA36A5A587DF821FDF3F87C" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[334,634,1946,1970]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="chimalapaensis" status="sp. nov.">
<emphasis id="39A7C046FFA36A5A587DF821FDF3F87C" box="[334,634,1946,1970]" italics="true" pageId="7" pageNumber="438">Ampliphallus chimalapaensis</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFA36A5A5BB2F821FD5BF87C" bold="true" box="[641,722,1946,1970]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="22947D3DFFA36A5A5BB2F821FD5BF87C" box="[641,722,1946,1970]" pageId="7" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CNAN-T01327). 7, chelicera, frontal view. 8, chelicera, mesal view. 9, pedipalp, mesal view. 10, pedipalpal tibia and tarsus, ventral view (major spiniform setiferous tubercles on tibia indicated as T1–T3). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="0B6C1C54FFAC6A5559F4FDF2FB02FDAD" blockId="8.[151,1437,152,611]" box="[199,1163,584,611]" pageId="8" pageNumber="439">
<emphasis id="39A7C046FFAC6A5559F4FDF2FE1BFDAD" bold="true" box="[199,402,584,611]" pageId="8" pageNumber="439">Included species.</emphasis>
<taxonomicName id="CCD367D7FFAC6A5558ABFDF2FD29FDAD" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[408,672,585,611]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="cuicateca" status="sp. nov.">
<emphasis id="39A7C046FFAC6A5558ABFDF2FD29FDAD" box="[408,672,585,611]" italics="true" pageId="8" pageNumber="439">Guelaguetzia cuicateca</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555B94FDF2FC88FDAD" bold="true" box="[679,769,585,611]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555B94FDF2FC88FDAD" box="[679,769,585,611]" pageId="8" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CCD367D7FFAC6A555A04FDF2FBA3FDAD" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[823,1066,585,611]" class="Arachnida" family="Stygnopsidae" genus="Guelaguetzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="serrana" status="sp. nov.">
<emphasis id="39A7C046FFAC6A555A04FDF2FBA3FDAD" box="[823,1066,585,611]" italics="true" pageId="8" pageNumber="439">Guelaguetzia serrana</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555D02FDF2FB02FDAD" bold="true" box="[1073,1163,585,611]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555D02FDF2FB02FDAD" box="[1073,1163,585,611]" pageId="8" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="5FAC4CDCFFAC6A5559A4F81BFAC4F812" ID-DOI="http://doi.org/10.5281/zenodo.3701033" ID-Zenodo-Dep="3701033" httpUri="https://zenodo.org/record/3701033/files/figure.png" pageId="8" pageNumber="439" startId="8.[151,264,1952,1977]" targetBox="[151,1435,633,1927]" targetPageId="8">
<paragraph id="0B6C1C54FFAC6A5559A4F81BFAC4F812" blockId="8.[151,1436,1952,2013]" pageId="8" pageNumber="439">
<emphasis id="39A7C046FFAC6A5559A4F81BFEDAF877" bold="true" box="[151,339,1952,1977]" pageId="8" pageNumber="439">FIGURES 11–15.</emphasis>
<taxonomicName id="CCD367D7FFAC6A555869F81AFD0EF877" authority="Cruz-López &amp; Francke, 2020" authorityName="Cruz-López &amp; Francke" authorityYear="2020" box="[346,647,1953,1977]" class="Arachnida" family="Stygnopsidae" genus="Ampliphallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="chimalapaensis" status="sp. nov.">
<emphasis id="39A7C046FFAC6A555869F81AFD0EF877" box="[346,647,1953,1977]" italics="true" pageId="8" pageNumber="439">Ampliphallus chimalapaensis</emphasis>
</taxonomicName>
<emphasis id="39A7C046FFAC6A555BBEF81AFD56F877" bold="true" box="[653,735,1953,1977]" pageId="8" pageNumber="439">
<taxonomicNameLabel id="22947D3DFFAC6A555BBEF81AFD56F877" box="[653,735,1953,1977]" pageId="8" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CNAN-T01327). 11, trochanter IV, dorsal view. 12, trochanter IV, ventral view. 13, leg IV, ectal view. 14, leg IV, mesal view. 15, femur IV, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="0B6C1C54FFAD6A5459F4FF23FA14FF18" blockId="9.[151,1437,152,287]" pageId="9" pageNumber="440">
<emphasis id="39A7C046FFAD6A5459F4FF23FD8CFF7D" bold="true" box="[199,517,152,179]" pageId="9" pageNumber="440">Geographical distribution.</emphasis>
The genus is only known from two pine forests in northern
<collectingRegion id="C917D2B6FFAD6A545D95FF22FB73FF7D" box="[1190,1274,153,179]" country="Mexico" name="Oaxaca" pageId="9" pageNumber="440">Oaxaca</collectingRegion>
(
<figureCitation id="93E800D1FFAD6A545C3AFF23FAD7FF7D" box="[1289,1374,152,179]" captionStart="FIGURE 57" captionStartId="21.[151,250,1794,1819]" captionTargetBox="[151,1435,1094,1769]" captionTargetId="figure@21.[151,1435,1094,1769]" captionTargetPageId="21" captionText="FIGURE 57. Distribution map of Ampliphallus chimalapaensis gen. nov., sp. nov. (white circle) and two species of Guelaguetzia gen.nov.: G. cuicateca sp. nov. (black square) and G. serrana sp. nov. (white square)." figureDoi="http://doi.org/10.5281/zenodo.3701061" httpUri="https://zenodo.org/record/3701061/files/figure.png" pageId="9" pageNumber="440">Fig. 57</figureCitation>
).
<emphasis id="39A7C046FFAD6A5459F4FF06FE0DFF19" bold="true" box="[199,388,188,215]" pageId="9" pageNumber="440">Natural history.</emphasis>
Both species of this genus were found inside of decomposing pine trunks, with high humidity.
</paragraph>
<paragraph id="0B6C1C54FFAD6A5459A4FF5AFA14FF34" blockId="9.[151,1437,152,287]" box="[151,1437,224,251]" pageId="9" pageNumber="440">In a few instances, a group of one male and two or three females were found, suggesting mate-guarding behavior.</paragraph>
<paragraph id="0B6C1C54FFAD6A5459A4FEBFFCB3FED0" blockId="9.[151,1437,152,287]" box="[151,826,260,287]" pageId="9" pageNumber="440">When the animals were disturbed, they exhibited thanatosis.</paragraph>
</subSubSection>
</treatment>
</document>
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