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<document id="1514113393D4A855E1D2831BA7974B1E" ID-ISSN="0024-4082" ID-ZooBank="611CD23B-912B-4737-AE4C-82218A2B0216" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630097274136" checkinUser="felipe" docAuthor="Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc &amp; Nel, André" docDate="2021" docId="6B39351BFFB8FFD5FCE451E113A0BF96" docLanguage="en" docName="zlaa078.pdf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Cretapristocera longiscapa Jouault &amp; Ngô-Muller &amp; Pouillon &amp; Nel 2021, SP. NOV." docType="treatment" docVersion="2" masterDocId="97004D63FFBBFFD0FF9357011630BF0D" masterDocTitle="New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea)" updateTime="1699143998499" updateUser="plazi">
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<mods:title id="7E691B090510DA856E1DA4E6083085D2">New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea)</mods:title>
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<mods:namePart id="016C491469A4CB4ED26B38D7ACD50E9E">Jouault, Corentin</mods:namePart>
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<mods:namePart id="AC14434A415CAD565FB3277B804FD950">Ngô-Muller, Valérie</mods:namePart>
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<mods:namePart id="F5D5EF39D18A89510F2D171AA32E7187">Pouillon, Jean-Marc</mods:namePart>
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<mods:namePart id="D0287031277AC4F71A028C685BFC4C3A">Nel, André</mods:namePart>
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<mods:title id="04992D820E89481AEDFAD2E514F2EBEC">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="A9FE40A16D4097ACF5A2A9DD90990C34">2021</mods:date>
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<subSubSection id="AB8AD786FFB8FFD3FCE451E11374B9F5" box="[887,1348,1759,1784]" pageId="3" type="nomenclature">
<paragraph id="E32F840DFFB8FFD3FCE451E11374B9F5" blockId="3.[887,1348,1759,1784]" box="[887,1348,1759,1784]" pageId="3">
<heading id="B8673361FFB8FFD3FCE451E11374B9F5" box="[887,1348,1759,1784]" centered="true" fontSize="9" level="2" pageId="3" reason="2">
<taxonomicName id="2490FF8EFFB8FFD3FCE451E112D0B9FA" authority="Jouault &amp; Ngô-Muller &amp; Pouillon &amp; Nel, 2021" authorityName="Jouault &amp; Ngô-Muller &amp; Pouillon &amp; Nel" authorityYear="2021" box="[887,1248,1760,1783]" class="Insecta" family="Bethylidae" genus="Cretapristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" phylum="Arthropoda" rank="species" species="longiscapa" status="sp. nov.">
<emphasis id="D1E4581FFFB8FFD3FCE451E112D0B9FA" box="[887,1248,1760,1783]" italics="true" pageId="3">CRETAPRISTOCERA LONGISCAPA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAD7E564FFB8FFD3FB7451E21374B9F5" box="[1255,1348,1759,1784]" pageId="3" rank="species">SP. NOV.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AB8AD786FFB8FFD5FBB2501013A0BF96" lastPageId="5" pageId="3" type="description">
<paragraph id="E32F840DFFB8FFD3FBB2501012A9B827" blockId="3.[1057,1177,1809,1834]" box="[1057,1177,1809,1834]" pageId="3">
(
<figureCitation id="7BAB9888FFB8FFD3FBB9501012A1B827" box="[1066,1169,1809,1834]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="4.[827,909,1262,1284]" captionStartId-1="5.[145,226,1200,1222]" captionStartId-2="6.[163,240,956,978]" captionTargetBox-0="[830,1445,198,1220]" captionTargetBox-1="[145,1428,197,1160]" captionTargetBox-2="[323,1283,197,914]" captionTargetId-0="figure-573@4.[827,1448,195,1223]" captionTargetId-1="figure-269@5.[145,1428,197,1160]" captionTargetId-2="figure-25@6.[320,1286,194,917]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 2. Cretapristocera longiscapa, sp. nov. holotype IGR.BU-009. A, habitus, in dorsal view. B, habitus, in ventral view. Scale bars: 0.5 mm." captionText-1="Figure 3. Cretapristocera longiscapa, sp. nov. holotype IGR.BU-009. A, detailed ventral head view. B, detailed dorsal mesosomal view. C, wings. D, detailed view of leg preserved muscles. Scale bars: 0.5 mm." captionText-2="Figure 4. Cretapristocera longiscapa, sp. nov.holotype IGR.BU-009. Line drawing of habitus, in dorsal view. Scale bar: 0.5 mm." pageId="3">FIGS 24</figureCitation>
)
</paragraph>
<paragraph id="E32F840DFFB8FFD3FCBA503B1243B863" blockId="3.[809,1425,1849,1902]" pageId="3">
u r n: l s i d: z o o b a n k. o r g: a c t:
<uuid id="9736BED8FFB8FFD3FB095038125FB863" pageId="3">2 1 A 7 C A E C - 4 2 5 0 - 45A9-A3B3-28F099EAEC25</uuid>
.
</paragraph>
<paragraph id="E32F840DFFBFFFD4FF3057C417C5BE9B" blockId="4.[163,779,197,407]" pageId="4">
<materialsCitation id="53F88E50FFBFFFD4FF3057C417C5BE9B" collectionCode="IGR" country="France" county="University of Rennes" location="Chalcidoidea" municipality="Museum" pageId="4" specimenCount="1" typeStatus="holotype">
<emphasis id="D1E4581FFFBFFFD4FF3057C41724BFD7" box="[163,276,197,218]" italics="true" pageId="4">
<typeStatus id="3C2B3AAFFFBFFFD4FF3057C4173EBFD7" box="[163,270,197,218]" pageId="4" type="holotype">Holotype</typeStatus>
:
</emphasis>
Female, accession number
<collectionCode id="85811CC8FFBFFFD4FDFD57C41493BFD7" box="[622,675,197,218]" pageId="4">IGR</collectionCode>
.BU-009, nearly complete and well-preserved female specimen in a rectangular piece of amber measuring 9 × 7 ×
<quantity id="246829E8FFBFFFD4FF6F56221765BE35" box="[252,341,291,313]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" unit="mm" value="2.5">2.5 mm</quantity>
, together with a small
<taxonomicName id="2490FF8EFFBFFFD4FDF656221537BE35" authorityName="LINNAEUS" authorityYear="1758" box="[613,775,291,312]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<location id="E64FD2D6FFBFFFD4FF3056431770BE55" LSID="urn:lsid:plazi:treatment:6B39351BFFB8FFD5FCE451E113A0BF96:E64FD2D6FFBFFFD4FF3056431770BE55" box="[163,320,322,344]" country="France" county="University of Rennes" municipality="Museum" name="Chalcidoidea" pageId="4">Chalcidoidea</location>
; housed in the amber collection of the
<location id="E64FD2D6FFBFFFD4FF30566317F0BE7A" LSID="urn:lsid:plazi:treatment:6B39351BFFB8FFD5FCE451E113A0BF96:E64FD2D6FFBFFFD4FF30566317F0BE7A" box="[163,448,354,376]" country="France" county="University of Rennes" municipality="Museum" name="Geological Department" pageId="4">Geological Department</location>
and
<collectingMunicipality id="034B1E77FFBFFFD4FD905663145BBE7A" box="[515,619,354,375]" pageId="4">Museum</collectingMunicipality>
(
<collectionCode id="85811CC8FFBFFFD4FDEE56631481BE75" box="[637,689,354,376]" pageId="4">IGR</collectionCode>
) of the
<collectingCounty id="0A4EFC81FFBFFFD4FF30568017A3BE9B" box="[163,403,385,407]" pageId="4">University of Rennes</collectingCounty>
,
<collectingCountry id="9B87C49DFFBFFFD4FE0C568017C1BE9B" box="[415,497,385,406]" name="France" pageId="4">France</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="E32F840DFFBFFFD4FF3056C01747BD19" blockId="4.[163,778,448,532]" pageId="4">
<emphasis id="D1E4581FFFBFFFD4FF3056C017E1BED8" box="[163,465,448,470]" italics="true" pageId="4">Type locality and horizon:</emphasis>
Noije Bum Hill, Hukawng Valley,
<collectingRegion id="21544AEFFFBFFFD4FF6456DE17A2BEF8" box="[247,402,479,501]" country="Myanmar" name="Kachin" pageId="4">Kachin State</collectingRegion>
,
<collectingCountry id="9B87C49DFFBFFFD4FE0C56E11423BEF8" box="[415,531,480,501]" name="Myanmar" pageId="4">Myanmar</collectingCountry>
; Lower Cenomanian, Upper Cretaceous.
</paragraph>
<paragraph id="E32F840DFFBFFFD4FF30553F1778BD9F" blockId="4.[163,779,574,658]" pageId="4">
<emphasis id="D1E4581FFFBFFFD4FF30553F1711BD5E" box="[163,289,574,595]" italics="true" pageId="4">Etymology:</emphasis>
The specific epithet is derived from Latin
<emphasis id="D1E4581FFFBFFFD4FF30555C16DEBD7F" box="[163,238,605,626]" italics="true" pageId="4">longus</emphasis>
, long, and
<emphasis id="D1E4581FFFBFFFD4FEF4555F1783BD7E" box="[359,435,606,627]" italics="true" pageId="4">scapus</emphasis>
, shaft, for the elongated scape of this species.
</paragraph>
<paragraph id="E32F840DFFBFFFD4FF3055C31318B948" blockId="4.[163,779,705,1853]" lastBlockId="4.[827,1443,1429,1605]" pageId="4">
<emphasis id="D1E4581FFFBFFFD4FF3055C3171BBDDA" box="[163,299,706,727]" italics="true" pageId="4">Description:</emphasis>
Female; body
<quantity id="246829E8FFBFFFD4FE4455C0141ABDDA" box="[471,554,705,727]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" unit="mm" value="2.1">2.1 mm</quantity>
long (as preserved), not depressed, almost smooth and not strongly foveolate; LFW
<quantity id="246829E8FFBFFFD4FECB55FE1780BC18" box="[344,432,767,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" unit="mm" value="1.5">1.5 mm</quantity>
; castaneous; wings setose. LH
<quantity id="246829E8FFBFFFD4FF30541E173EBC39" box="[163,270,799,820]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" unit="mm" value="0.43">0.43 mm</quantity>
; WH
<quantity id="246829E8FFBFFFD4FEC0541E1780BC39" box="[339,432,799,821]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" unit="mm" value="0.5">0.5 mm</quantity>
; WF
<quantity id="246829E8FFBFFFD4FE63541E146CBC39" box="[496,604,799,821]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="4" unit="mm" value="0.26">0.26 mm</quantity>
; HE
<quantity id="246829E8FFBFFFD4FD08541E1535BC39" box="[667,773,799,820]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="4" unit="mm" value="0.11">0.11 mm</quantity>
; OOL
<quantity id="246829E8FFBFFFD4FF77543F177FBC59" box="[228,335,830,852]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" unit="mm" value="0.25">0.25 mm</quantity>
; WOT
<quantity id="246829E8FFBFFFD4FE31543F143DBC59" box="[418,525,830,852]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" unit="mm" value="0.16">0.16 mm</quantity>
; DAO
<quantity id="246829E8FFBFFFD4FDCE543F14F8BC59" box="[605,712,830,852]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="4" unit="mm" value="0.04">0.04 mm</quantity>
; VOL
<quantity id="246829E8FFBFFFD4FF30545F1734BC7E" box="[163,260,862,883]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" unit="mm" value="0.1">0.1 mm</quantity>
. Head not pubescent, with small, dorsal, median depression, finely punctuated but not sculptured; mandible with one strong and sharp tooth, projecting forward; six maxillary palpomeres, three (?) visible labial palpomeres; clypeus spined; antenna with ten flagellomeres with cylindrical crosssection; scape 0.37 ×
<quantity id="246829E8FFBFFFD4FE00531817C8BB22" box="[403,504,1049,1071]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="4" unit="mm" value="0.07">0.07 mm</quantity>
, pedicel 0.13 ×
<quantity id="246829E8FFBFFFD4FD3B5318153BBB22" box="[680,779,1049,1071]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="4" unit="mm" value="0.04">0.04 mm</quantity>
(all measurements refer to length × width), F1 0.07 ×
<quantity id="246829E8FFBFFFD4FF7E53591761BB60" box="[237,337,1112,1134]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" unit="mm" value="0.05">0.05 mm</quantity>
, F2 0.05 ×
<quantity id="246829E8FFBFFFD4FE5B5359141CBB60" box="[456,556,1112,1134]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" unit="mm" value="0.05">0.05 mm</quantity>
, F3 0.05 ×
<quantity id="246829E8FFBFFFD4FD3053591537BB60" box="[675,775,1112,1134]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" unit="mm" value="0.06">0.06 mm</quantity>
, F 4 0.06 ×
<quantity id="246829E8FFBFFFD4FEAA5376179EBB81" box="[313,430,1143,1164]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="4" unit="mm" value="0.07">0.07 mm</quantity>
, F5 0.06 ×
<quantity id="246829E8FFBFFFD4FDCE537614E2BB81" box="[605,722,1143,1164]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" unit="mm" value="0.08">0.08 mm</quantity>
, F 6 0.08 ×
<quantity id="246829E8FFBFFFD4FF7E53971761BBA1" box="[237,337,1174,1196]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" unit="mm" value="0.08">0.08 mm</quantity>
, F7 0.08 ×
<quantity id="246829E8FFBFFFD4FE5B5397141CBBA1" box="[456,556,1174,1196]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" unit="mm" value="0.08">0.08 mm</quantity>
, F8 0.08 ×
<quantity id="246829E8FFBFFFD4FD3053971537BBA1" box="[675,775,1174,1196]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="4" unit="mm" value="0.07">0.07 mm</quantity>
, F9 0.09 ×
<quantity id="246829E8FFBFFFD4FEB153B417BEBBC6" box="[290,398,1205,1227]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" unit="mm" value="0.06">0.06 mm</quantity>
, F10 0.17 ×
<quantity id="246829E8FFBFFFD4FDB953B414A6BBC6" box="[554,662,1205,1227]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" unit="mm" value="0.06">0.06 mm</quantity>
; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not equilateral, with two posterior ocelli more distant from each other than from anterior. Mesosoma with dorsum smooth, not roughly sculptured; femora strongly thickened, tibia thin, tarsal claws simple; tibial spur formula 1-2-2; notauli well marked, posteriorly convergent; parapsidal furrows weak; metanotum developed medially and overlapping mesoscutellum posteriorly; metapectal propodeal complex rectangular, not foveolate, with marginal carinae and no posterior spines; posterior margin of propodeum straight. Forewing covered with microsetae and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; costal vein present; only veins C, Sc + R, Rs + M, 2r-rs&amp;Rs, M + Cu, Cu and cells C (hardly visible), R and 2R1 present; vein cu-a absent and cell Cu opened; cell 2R1 distally opened; no basal stub of Rs + M; pterostigma long and narrow; poststigmal abscissa of R1 short, spectral veins not visible. Hindwing with only veins C and Sc + R and with three hamuli. Metasoma with short petiole; six tergites present (including three preserved detached from the body), T2 enlarged, convex, partly covering sternites; sting not preserved; length of clearly visible tergites: T1
<quantity id="246829E8FFBFFFD4FCA8512E1590B948" box="[827,928,1583,1605]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" unit="mm" value="0.23">0.23 mm</quantity>
, T2
<quantity id="246829E8FFBFFFD4FC5D512E1203B948" box="[974,1075,1583,1605]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="4" unit="mm" value="0.29">0.29 mm</quantity>
, T3 at least
<quantity id="246829E8FFBFFFD4FB2D512E1314B948" box="[1214,1316,1583,1605]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="4" unit="mm" value="0.19">0.19 mm</quantity>
.
</paragraph>
<caption id="B7EFD485FFBFFFD4FCA853EF12A2BA32" pageId="4" startId="4.[827,909,1262,1284]" targetBox="[830,1445,198,1220]" targetPageId="4">
<paragraph id="E32F840DFFBFFFD4FCA853EF12A2BA32" blockId="4.[827,1442,1262,1343]" pageId="4">
<emphasis id="D1E4581FFFBFFFD4FCA853EF1599BA0E" bold="true" box="[827,937,1262,1284]" pageId="4">Figure 2.</emphasis>
<taxonomicName id="2490FF8EFFBFFFD4FC2553EF12EDBA09" authority="Jouault &amp; Ngô-Muller &amp; Pouillon &amp; Nel, 2021" authorityName="Jouault &amp; Ngô-Muller &amp; Pouillon &amp; Nel" authorityYear="2021" box="[950,1245,1262,1284]" class="Insecta" family="Bethylidae" genus="Cretapristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" phylum="Arthropoda" rank="species" species="longiscapa" status="sp. nov.">
<emphasis id="D1E4581FFFBFFFD4FC2553EF12EDBA09" box="[950,1245,1262,1284]" italics="true" pageId="4">Cretapristocera longiscapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="CAD7E564FFBFFFD4FB7F53EF130DBA09" box="[1260,1341,1262,1284]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
holotype IGR.BU-009. A, habitus, in dorsal view. B, habitus, in ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="E32F840DFFBFFFD4FCA8516F12F4B830" blockId="4.[827,1443,1646,1853]" pageId="4">
<emphasis id="D1E4581FFFBFFFD4FCA8516F159BB98E" box="[827,939,1646,1667]" italics="true" pageId="4">Remarks:</emphasis>
It seems that the internal musculature of the fossil was preserved during the fossilization process, especially those of the legs (
<figureCitation id="7BAB9888FFBFFFD4FA9951AD1344B9CC" box="[1290,1396,1708,1730]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStartId-0="4.[827,909,1262,1284]" captionStartId-1="5.[145,226,1200,1222]" captionTargetBox-0="[830,1445,198,1220]" captionTargetBox-1="[145,1428,197,1160]" captionTargetId-0="figure-573@4.[827,1448,195,1223]" captionTargetId-1="figure-269@5.[145,1428,197,1160]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figure 2. Cretapristocera longiscapa, sp. nov. holotype IGR.BU-009. A, habitus, in dorsal view. B, habitus, in ventral view. Scale bars: 0.5 mm." captionText-1="Figure 3. Cretapristocera longiscapa, sp. nov. holotype IGR.BU-009. A, detailed ventral head view. B, detailed dorsal mesosomal view. C, wings. D, detailed view of leg preserved muscles. Scale bars: 0.5 mm." pageId="4">Figs 23</figureCitation>
). It could be interesting to compare the evolution of the musculature and the insertion of the muscles between the Cretaceous and extant
<taxonomicName id="2490FF8EFFBFFFD4FB1D50091322B813" authorityName="HALIDAY" authorityYear="1839" box="[1166,1298,1800,1822]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="4" phylum="Arthropoda" rank="family">Bethylidae</taxonomicName>
in a future study, using 3D X-ray tomography.
</paragraph>
<paragraph id="E32F840DFFBEFFD5FD56578513A0BF96" blockId="5.[709,1424,128,155]" box="[709,1424,128,155]" pageId="5">CRETACEOUS EVOLUTION OF BETHYLID WASPS 1049</paragraph>
</subSubSection>
</treatment>
</document>