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<document ID-DOI="http://doi.org/10.5281/zenodo.3790612" ID-GBIF-Dataset="ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7" ID-GBIF-Taxon="163754305" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790612" ID-ZooBank="BB883B1E-E58F-4074-92EB-7E814E78F678" approvalRequired="19" approvalRequired_for_textStreams="19" checkinTime="1587996592918" checkinUser="plazi" docAuthor="Hopp, Katie &amp; Caterino, Michael" docDate="2009" docId="4157F97AFF9AFFF0FF7EFD6EFC71170F" docLanguage="en" docName="ZK_article_2083.pdf" docOrigin="ZooKeys 24 (24)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5c452e69" docStyleName="zookeys.2008.journal_article" docTitle="Cepehennium celsifrons Hopp &amp; Caterino 2009, sp. n." docType="treatment" docVersion="5" lastPageId="10" lastPageNumber="41" masterDocId="BD6E8102FF9DFFFAFFF2FF92FFBD1451" masterDocTitle="Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species" masterLastPageNumber="54" masterPageNumber="31" pageId="7" pageNumber="38" updateTime="1643563761986" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hopp, Katie</mods:namePart>
<mods:affiliation>,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael</mods:namePart>
<mods:affiliation>Clemson University, Clemson, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-10-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>24</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>24</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.24.247</mods:identifier>
<mods:identifier type="GBIF-Dataset">ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576549</mods:identifier>
<mods:identifier type="ZooBank">BB883B1E-E58F-4074-92EB-7E814E78F678</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790612" ID-GBIF-Taxon="163754305" ID-Zenodo-Dep="3790612" LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F" httpUri="http://treatment.plazi.org/id/4157F97AFF9AFFF0FF7EFD6EFC71170F" lastPageId="10" lastPageNumber="41" pageId="7" pageNumber="38">
<subSubSection pageId="7" pageNumber="38" type="nomenclature">
<paragraph blockId="7.[140,991,764,862]" box="[140,720,764,791]" pageId="7" pageNumber="38">
<heading box="[140,720,764,791]" fontSize="11" level="2" pageId="7" pageNumber="38" reason="3">
<emphasis bold="true" box="[140,720,764,791]" pageId="7" pageNumber="38">
<taxonomicName authority="Hopp &amp; Caterino" authorityName="Hopp &amp; Caterino" authorityYear="2009" box="[140,642,764,791]" class="Insecta" family="Scydmaenidae" genus="Cepehennium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="celsifrons" status="sp. n.">
<emphasis bold="true" box="[140,420,764,791]" italics="true" pageId="7" pageNumber="38">
<emphasis box="[140,304,764,791]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Cepehennium</emphasis>
celsifrons
</emphasis>
Hopp &amp; Caterino
</taxonomicName>
,
<taxonomicNameLabel box="[655,720,765,791]" pageId="7" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[140,991,764,862]" box="[140,991,800,827]" pageId="7" pageNumber="38">
<uri box="[140,991,800,827]" pageId="7" pageNumber="38">
urn:lsid:zoobank.org:act:
<uuid box="[421,991,800,827]" pageId="7" pageNumber="38">FB4E5DDF-D621-4B8E-A233-E2956D8C66EB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,470,835,862]" pageId="7" pageNumber="38" type="description">
<paragraph blockId="7.[140,991,764,862]" box="[140,470,835,862]" pageId="7" pageNumber="38">Figs 1B, 2B, 3B, 4A-B, 5B, 6</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="38" type="materials_examined">
<paragraph blockId="7.[140,1108,905,1672]" pageId="7" pageNumber="38">
<emphasis bold="true" box="[140,318,905,932]" pageId="7" pageNumber="38">
<emphasis box="[140,198,906,932]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Type</emphasis>
Material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608963404" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" county="Calaveras Co." determinerName="A. Newton" elevation="686" latitude="38.416" location="West Point" longLatPrecision="6" longitude="-120.5515" pageId="7" pageNumber="38" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis bold="true" box="[329,527,906,932]" pageId="7" pageNumber="38">
<typeStatus box="[329,445,906,932]" pageId="7" pageNumber="38">Holotype</typeStatus>
. Male.
</emphasis>
“CALIF:
<collectingCounty box="[650,811,906,932]" pageId="7" pageNumber="38">Calaveras Co.</collectingCounty>
; 3.0 mi NW
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFC21FC18FBE917F5" box="[979,1108,906,932]" county="Calaveras Co." latitude="38.416" longLatPrecision="6" longitude="-120.5515" name="West Point" pageId="7" pageNumber="38">West Point</location>
[̴
<geoCoordinate box="[157,294,941,967]" degrees="38.4160" direction="north" orientation="latitude" pageId="7" pageNumber="38" precision="5" value="38.416">38.4160°N</geoCoordinate>
,
<geoCoordinate box="[307,461,940,967]" degrees="120.5515" direction="west" orientation="longitude" pageId="7" pageNumber="38" precision="5" value="-120.5515">120.5515°W</geoCoordinate>
];
<quantity box="[480,575,941,968]" metricMagnitude="2" metricUnit="m" metricValue="6.858" pageId="7" pageNumber="38" unit="ft" value="2250.0">
<elevation box="[480,575,941,968]" metricMagnitude="2" metricUnit="m" metricValue="6.858" pageId="7" pageNumber="38" unit="ft" value="2250.0">2250 ft.</elevation>
</quantity>
, v.20.1976; berl.; litter, mixed hdwd.-
<taxonomicName box="[1021,1082,942,967]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="38" phylum="Tracheophyta" rank="genus">
<emphasis box="[1021,1082,942,967]" italics="true" pageId="7" pageNumber="38">Pinus</emphasis>
</taxonomicName>
-;
<emphasis box="[140,342,976,1003]" italics="true" pageId="7" pageNumber="38">
<taxonomicName authorityName="Endlicher" authorityYear="1847" box="[140,256,976,1002]" class="Pinopsida" family="Cupressaceae" genus="Libocedrus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="38" phylum="Tracheophyta" rank="genus">Libocedrus</taxonomicName>
– Abies
</emphasis>
; for.;
<collectorName box="[410,538,976,1003]" pageId="7" pageNumber="38">A. Newton</collectorName>
,
<collectorName box="[550,671,976,1003]" pageId="7" pageNumber="38">M. Thayer</collectorName>
”/ “
<taxonomicName box="[709,849,976,1002]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis box="[709,849,976,1002]" italics="true" pageId="7" pageNumber="38">Cephennium</emphasis>
</taxonomicName>
sp.
<specimenCount box="[895,910,977,1003]" pageId="7" pageNumber="38" type="female">♀</specimenCount>
;
<determinerName box="[925,1050,976,1003]" pageId="7" pageNumber="38">A. Newton</determinerName>
det. 1978” (
<collectionCode box="[226,321,1012,1038]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="7.[140,1108,905,1672]" pageId="7" pageNumber="38">
<materialsCitation ID-GBIF-Occurrence="2608963399" collectionCode="SBMN" collectorName="A. Newton &amp; M. Thayer" county="Calaveras Co." determinerName="A. Newton" elevation="686" latitude="38.416" location="West Point" longLatPrecision="6" longitude="-120.5515" pageId="7" pageNumber="38" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[185,300,1047,1073]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">
<typeStatus box="[185,300,1047,1073]" pageId="7" pageNumber="38">Paratypes</typeStatus>
</emphasis>
(8):
<specimenCount box="[360,489,1047,1073]" pageId="7" pageNumber="38" type="generic">1 specimen</specimenCount>
with same data as holotype (
<collectionCode box="[830,913,1047,1073]" pageId="7" pageNumber="38">SBMN</collectionCode>
[gold coated for SEM])
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963408" collectionCode="MCZC" collectorName="A. Newton &amp; M. Thayer &amp; M. Th" county="Calaveras Co." determinerName="A. Newton" elevation="686" isTailEnumeration="true" latitude="38.416" location="West" longLatPrecision="6" longitude="-120.5515" pageId="7" pageNumber="38" specimenCount="1" typeStatus="paratype">
“CALIF: Calaveras; Co.,
<quantity box="[522,567,1082,1108]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="7" pageNumber="38" unit="mi" value="3.0">3mi</quantity>
NW
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFD89FBA8FD0B1005" box="[635,694,1082,1108]" name="West" pageId="7" pageNumber="38">West</location>
; Point 2250’; v.20.1976”/ “
<collectorName pageId="7" pageNumber="38">A. Newton</collectorName>
;
<collectorName box="[195,318,1117,1144]" pageId="7" pageNumber="38">M. Thayer</collectorName>
; collectors”/ “
<collectorName box="[477,606,1117,1144]" pageId="7" pageNumber="38">A. Newton</collectorName>
;
<collectorName box="[619,685,1117,1144]" pageId="7" pageNumber="38">M. Th</collectorName>
ayer; collectors” (1
<collectionCode box="[915,1005,1117,1143]" pageId="7" pageNumber="38">MCZC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963391" collectionCode="FMNH" collectorName="A. Newton" county="Calaveras Co." determinerName="A. Newton" location="3 mi NE Glencoe" pageId="7" pageNumber="38" specimenCount="1" typeStatus="paratype">
“CAL.:
<collectingCounty box="[140,299,1152,1178]" pageId="7" pageNumber="38">Calaveras Co.</collectingCounty>
;
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFECBFB12FDBA10CB" box="[313,519,1152,1179]" county="Calaveras Co." name="3 mi NE Glencoe" pageId="7" pageNumber="38">
<quantity box="[313,367,1152,1178]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="7" pageNumber="38" unit="mi" value="3.0">3 mi</quantity>
NE Glencoe
</location>
2000’; VI.25.1975 berl. litter,; oak – conifer forest away; from stream;
<collectorName box="[363,489,1188,1214]" pageId="7" pageNumber="38">A. Newton</collectorName>
”/ “
<taxonomicName box="[527,667,1187,1213]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis box="[527,667,1187,1213]" italics="true" pageId="7" pageNumber="38">Cephennium</emphasis>
</taxonomicName>
;
<determinerName box="[681,806,1188,1214]" pageId="7" pageNumber="38">A. Newton</determinerName>
det. 1975” (2
<collectionCode box="[974,1066,1188,1214]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38" type="Museum">FMNH</collectionCode>
, females)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963402" collectingDate="1971-03-03" collectionCode="FMNH" county="Amador Co." location="Cal." municipality="Bar" pageId="7" pageNumber="38" specimenCount="1" stateProvince="Michigan" typeStatus="paratype">
“3 miW
<collectingRegion box="[323,432,1223,1249]" country="United States of America" name="Michigan" pageId="7" pageNumber="38">Michigan</collectingRegion>
<collectingMunicipality box="[437,479,1223,1249]" pageId="7" pageNumber="38">Bar</collectingMunicipality>
;
<collectingCounty box="[489,631,1223,1249]" pageId="7" pageNumber="38">Amador Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFD70FB55FD0910B0" box="[642,692,1223,1249]" county="Amador Co." municipality="Bar" name="Cal." pageId="7" pageNumber="38" stateProvince="Michigan">Cal.</location>
;
<date box="[702,825,1223,1249]" pageId="7" pageNumber="38" value="1971-03-03">
<collectingDate box="[702,825,1223,1249]" pageId="7" pageNumber="38" value="1971-03-03">III-3-1971</collectingDate>
</date>
”/ “ex Berlese-; oak duff”/ “Fred G.; Andrews; collector” (1
<collectionCode box="[533,625,1258,1284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38" type="Museum">FMNH</collectionCode>
, male [disarticulated])
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963403" collectingDate="1972-04-15" collectionCode="CSCA" county="Amador Co." location="Pine Grove" pageId="7" pageNumber="38" specimenCount="1" stateProvince="Michigan" typeStatus="paratype">
“CALIF:
<collectingCounty pageId="7" pageNumber="38">Amador Co.</collectingCounty>
;
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFF34FA9FFEF41176" box="[198,329,1293,1319]" county="Amador Co." name="Pine Grove" pageId="7" pageNumber="38" stateProvince="Michigan">Pine Grove</location>
;
<date box="[342,480,1293,1320]" pageId="7" pageNumber="38" value="1972-04-15">
<collectingDate box="[342,480,1293,1320]" pageId="7" pageNumber="38" value="1972-04-15">IV-15-1972</collectingDate>
</date>
; Fred G Andrews”/ “Berlesed; Oak; Litter” (1
<collectionCode box="[1017,1095,1293,1319]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="7" pageNumber="38" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963392" collectingDate="1971-02-14" collectionCode="CSCA" collectorName="R. F. Wilkey &amp; Ex. Rotten" county="Amador Co." location="1 mi. W Pine Grove" pageId="7" pageNumber="38" specimenCount="1" stateProvince="Michigan" typeStatus="paratype">
“CALIF:
<collectingCounty box="[250,397,1328,1354]" pageId="7" pageNumber="38">Amador Co.</collectingCounty>
;
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFE6EFAA2FD2F111B" box="[412,658,1328,1355]" county="Amador Co." name="1 mi. W Pine Grove" pageId="7" pageNumber="38" stateProvince="Michigan">
<quantity box="[412,476,1328,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="7" pageNumber="38" unit="mi" value="1.0">1 mi.</quantity>
W Pine Grove
</location>
;
<date box="[672,801,1328,1354]" pageId="7" pageNumber="38" value="1971-02-14">
<collectingDate box="[672,801,1328,1354]" pageId="7" pageNumber="38" value="1971-02-14">II-14-1971</collectingDate>
</date>
;
<collectorName box="[817,968,1328,1355]" pageId="7" pageNumber="38">R. F. Wilkey</collectorName>
;
<collectorName box="[982,1108,1329,1355]" pageId="7" pageNumber="38">Ex. Rotten</collectorName>
wood” (1
<collectionCode box="[251,329,1364,1390]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="7" pageNumber="38" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963409" collectionCode="CSCA" county="Amador Co." location="Cal." municipality="Pine Grove" pageId="7" pageNumber="38" specimenCount="1" stateProvince="Michigan" typeStatus="paratype">
“
<collectingMunicipality box="[357,488,1363,1390]" pageId="7" pageNumber="38">Pine Grove</collectingMunicipality>
;
<collectingCounty box="[499,641,1363,1390]" pageId="7" pageNumber="38">Amador Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9AFFF0FF7EFD6EFC71170F:CC211EB7FF9AFFFDFD7EFAC1FD03113C" box="[652,702,1363,1389]" county="Amador Co." municipality="Pine Grove" name="Cal." pageId="7" pageNumber="38" stateProvince="Michigan">Cal.</location>
; IV.15.1972”/ “Berlesed; from; Oak duff” (1
<collectionCode box="[237,310,1399,1425]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="7" pageNumber="38" type="Museum">CSCA</collectionCode>
[disarticulated])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="38" type="etymology">
<paragraph blockId="7.[140,1108,905,1672]" pageId="7" pageNumber="38">
<emphasis bold="true" box="[185,320,1434,1460]" pageId="7" pageNumber="38">
<emphasis box="[185,315,1434,1460]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Etymology</emphasis>
.
</emphasis>
This species name is derived from the latin
<emphasis box="[836,893,1434,1460]" italics="true" pageId="7" pageNumber="38">celsus</emphasis>
(elevated or lofty) in combination with
<emphasis box="[384,436,1469,1495]" italics="true" pageId="7" pageNumber="38">frons</emphasis>
, in reference to the unique, prominent sexual dimorphism exhibited by the male.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="39" pageId="7" pageNumber="38" type="diagnosis">
<paragraph blockId="7.[140,1108,905,1672]" lastBlockId="8.[140,1108,1187,1672]" lastPageId="8" lastPageNumber="39" pageId="7" pageNumber="38">
<emphasis bold="true" box="[185,312,1539,1565]" pageId="7" pageNumber="38">
<emphasis box="[185,307,1539,1565]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Diagnosis</emphasis>
.
</emphasis>
This species can be distinguished from all
<collectingRegion box="[795,909,1539,1565]" country="United States of America" name="California" pageId="7" pageNumber="38">California</collectingRegion>
congeners by the character combination of the presence of two ommatidia on each side of the head (
<figureCitation captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="7" pageNumber="38">Fig. 4A</figureCitation>
), the mesosternal process truncate at the apex (
<figureCitation box="[711,799,1610,1636]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="7" pageNumber="38">Fig. 2B</figureCitation>
), and the frons of the male with a median longitudinal ridge (
<figureCitation box="[529,617,1645,1671]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="7" pageNumber="38">Fig. 4B</figureCitation>
). Th e male frons structure is unique to this species.
<taxonomicName authorityName="Hopp &amp; Caterino C" authorityYear="2009" box="[232,475,1187,1213]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis box="[232,475,1187,1213]" italics="true" pageId="8" pageNumber="39">Cephennium celsifrons</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[535,678,1188,1214]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[535,678,1188,1214]" italics="true" pageId="8" pageNumber="39">C. mariposae</emphasis>
</taxonomicName>
are otherwise very similar but can be distinguished by the number of ommatidia present (2 vs. 4, respectively,
<figureCitation box="[963,1093,1223,1249]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="8" pageNumber="39">Figs. 4A, C</figureCitation>
).
<taxonomicName box="[140,368,1258,1284]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[140,368,1258,1284]" italics="true" pageId="8" pageNumber="39">Cephennium aridum</emphasis>
</taxonomicName>
has a single ommatidium on each side of the head (
<figureCitation box="[952,1044,1258,1284]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="8" pageNumber="39">Fig. 4D</figureCitation>
), and no other species exhibit any traces of eyes.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770031" ID-Zenodo-Dep="3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="8" pageNumber="39" startId="8.[140,212,1040,1062]" targetBox="[140,1108,165,1021]" targetPageId="8">
<paragraph blockId="8.[140,1108,997,1126]" pageId="8" pageNumber="39">
<emphasis bold="true" box="[140,297,1040,1062]" pageId="8" pageNumber="39">Figure 4. A–D</emphasis>
Characters on the head, including number of ommatidia and male frons
<emphasis bold="true" box="[980,1027,1040,1062]" pageId="8" pageNumber="39">A–B</emphasis>
<taxonomicName authority="Hopp &amp; Caterino C" authorityName="Hopp &amp; Caterino C" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis italics="true" pageId="8" pageNumber="39">Cephennium celsifrons</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[464,483,1072,1094]" pageId="8" pageNumber="39">C</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino D" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[490,910,1072,1094]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[490,702,1072,1094]" italics="true" pageId="8" pageNumber="39">Cephennium mariposae</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[891,910,1072,1094]" pageId="8" pageNumber="39">D</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino." authorityName="Hopp &amp; Caterino." class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[918,1107,1072,1093]" italics="true" pageId="8" pageNumber="39">Cephennium aridum</emphasis>
Hopp &amp; Caterino.
</taxonomicName>
</paragraph>
</caption>
<subSubSection lastPageId="10" lastPageNumber="41" pageId="8" pageNumber="39" type="description">
<paragraph blockId="8.[140,1108,1187,1672]" lastBlockId="10.[140,1108,167,862]" lastPageId="10" lastPageNumber="41" pageId="8" pageNumber="39">
<emphasis bold="true" box="[185,408,1328,1354]" pageId="8" pageNumber="39">
<emphasis box="[185,330,1328,1354]" inLineHeading="true" pageId="8" pageNumber="39" reason="1">Description</emphasis>
. Male.
</emphasis>
Length:
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<geoCoordinate box="[508,574,1328,1354]" degrees="0.817" direction="north" orientation="latitude" pageId="8" pageNumber="39" precision="55" value="0.817">0.817</geoCoordinate>
<geoCoordinate box="[574,654,1328,1354]" degrees="0.893" direction="west" orientation="longitude" pageId="8" pageNumber="39" precision="55" value="-0.893">–0.893</geoCoordinate>
mm
</quantity>
; pronotal width:
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<geoCoordinate box="[901,966,1328,1354]" degrees="0.323" direction="north" orientation="latitude" pageId="8" pageNumber="39" precision="55" value="0.323">0.323</geoCoordinate>
<geoCoordinate box="[966,1048,1328,1354]" degrees="0.380" direction="west" orientation="longitude" pageId="8" pageNumber="39" precision="55" value="-0.38">–0.380</geoCoordinate>
mm
</quantity>
; elytral width:
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<geoCoordinate box="[302,368,1364,1390]" degrees="0.380" direction="north" orientation="latitude" pageId="8" pageNumber="39" precision="55" value="0.38">0.380</geoCoordinate>
<geoCoordinate box="[369,449,1364,1390]" degrees="0.418" direction="west" orientation="longitude" pageId="8" pageNumber="39" precision="55" value="-0.418">–0.418</geoCoordinate>
mm
</quantity>
. Body broad, ovate, slightly convex, rufo-testaceous to amber yellow, evenly densely pubescent, pubescence golden, slender, long, moderately decumbent (Fig. 1B). Head small, deflexed, sparsely pubescent, not narrowing anteriorly from antennal insertions; frons pinched medially forming a medial ridge (
<figureCitation box="[148,236,1504,1530]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="8" pageNumber="39">Fig. 4B</figureCitation>
); two ommatidia present on each side of the head (
<figureCitation box="[820,910,1504,1530]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="8" pageNumber="39">Fig. 4A</figureCitation>
). Antenna setose, antennomere I and II longer than broad, antennomeres III-VI quadrate and smaller than antennomeres II and VII, antennomere VIII smaller than antennomeres VII and IX, antennomeres IX-XI gradually clavate forming loose club. Pronotum densely pubescent, broadest between middle and anterior third, disc very convex medially and weakly flattened near each posterior angle; anterior margin not visible from above; anterior and posterior margin lacking marginal bead; marginal bead complete laterally, gradually widening towards base; lateral edge broadly rounded to posterior third, then evenly curved to base (Fig. 1B). Hypomeron smooth, sparsely setose towards upper quarter and along outside (lateral) edge, hypomeral bead anterolaterad procoxae sinuate. Prosternum without nodules anterolaterad procoxal cavities (
<figureCitation box="[930,1018,237,263]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="10" pageNumber="41">Fig. 2B</figureCitation>
). Elytra impunctate, pubescent as pronotum, covering all abdominal segments, weakly truncate at apex; elytral suture flat; elytral striae absent; basomedial fovea present on each elytron, fovea small, with dense inwardly directed setae (Figs. 1B, 3B). Humeral angles of elytron raised, dorsally flattened (plateau-like), laterally rounded; weakly curved posterad around anterolateral angle (
<figureCitation box="[560,649,413,439]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="10" pageNumber="41">Fig. 3B</figureCitation>
). Scutellum roundly triangular, without setae (
<figureCitation box="[209,291,448,474]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="10" pageNumber="41">Fig 3B</figureCitation>
). Mesosternal keel setose with indentation around each seta; smooth in the posterior quarter portion, apex straight (
<figureCitation box="[659,750,483,510]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="10" pageNumber="41">Fig. 2B</figureCitation>
). Metathoracic wings vestigial. Femora strongly clavate in distal half, tibiae expanded and becoming more densely setose towards distal half. Six visible abdominal sternites (fusion between ventrites V and VI incomplete). Aedeagus with median lobe rather lightly sclerotized, bulbous and deeply emarginate at base; apex with keel-like median dorsal process and dorsally curved, apically convex digiform process extending over part of apical collar (
<figureCitation box="[1007,1094,659,686]" captionStart="Figure 5" captionStartId="9.[140,212,1408,1430]" captionTargetBox="[156,1106,208,1373]" captionTargetId="figure@9.[140,1108,165,1389]" captionTargetPageId="9" captionText="Figure 5. A–F Dorsal and lateral view of the aedeagus A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium grandarboreum Hopp &amp; Caterino D Cephennium canestroi Hopp &amp; Caterino E Cephennium urbanum Hopp &amp; Caterino F Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770035" httpUri="https://zenodo.org/record/3770035/files/figure.png" pageId="10" pageNumber="41">Fig. 5B</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770035" ID-Zenodo-Dep="3770035" httpUri="https://zenodo.org/record/3770035/files/figure.png" pageId="9" pageNumber="40" startId="9.[140,212,1408,1430]" targetBox="[156,1106,208,1373]" targetPageId="9">
<paragraph blockId="9.[140,1108,1408,1526]" pageId="9" pageNumber="40">
<emphasis bold="true" box="[140,287,1408,1430]" pageId="9" pageNumber="40">Figure 5. A–F</emphasis>
Dorsal and lateral view of the aedeagus
<emphasis bold="true" box="[668,687,1408,1430]" pageId="9" pageNumber="40">A</emphasis>
<taxonomicName authority="Brendel B" authorityName="Brendel B" authorityYear="1889" box="[694,1069,1408,1431]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[694,966,1409,1430]" italics="true" pageId="9" pageNumber="40">Cephennium anophthalmicum</emphasis>
Brendel
<emphasis bold="true" box="[1052,1069,1408,1430]" pageId="9" pageNumber="40">B</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino C" authorityName="Hopp &amp; Caterino C" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis italics="true" pageId="9" pageNumber="40">Cephennium celsifrons</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[500,519,1440,1462]" pageId="9" pageNumber="40">C</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino D" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[525,986,1440,1463]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[525,783,1441,1462]" italics="true" pageId="9" pageNumber="40">Cephennium grandarboreum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[967,986,1440,1462]" pageId="9" pageNumber="40">D</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino E" authorityName="Hopp &amp; Caterino E" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="canestroi">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis italics="true" pageId="9" pageNumber="40">Cephennium canestroi</emphasis>
</taxonomicName>
Hopp &amp; Caterino
<emphasis bold="true" box="[400,415,1472,1494]" pageId="9" pageNumber="40">E</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino F" authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[422,825,1472,1494]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="urbanum">
<taxonomicName authority="Hopp &amp; Caterino" authorityName="Hopp &amp; Caterino" box="[422,804,1472,1494]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[422,626,1473,1494]" italics="true" pageId="9" pageNumber="40">Cephennium urbanum</emphasis>
Hopp &amp; Caterino
</taxonomicName>
<emphasis bold="true" box="[811,825,1472,1494]" pageId="9" pageNumber="40">F</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino." authorityName="Hopp &amp; Caterino." class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="aridum">
<taxonomicName authority="Hopp" authorityName="Hopp" box="[832,1081,1473,1494]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="aridum">
<taxonomicName box="[832,1019,1473,1494]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[832,1019,1473,1494]" italics="true" pageId="9" pageNumber="40">Cephennium aridum</emphasis>
</taxonomicName>
Hopp
</taxonomicName>
&amp; Caterino.
</taxonomicName>
</paragraph>
</caption>
<paragraph blockId="10.[140,1108,167,862]" box="[185,670,695,721]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[185,277,695,721]" pageId="10" pageNumber="41">
<emphasis box="[185,272,695,721]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Female</emphasis>
.
</emphasis>
Identical to male except frons flat.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="biology_ecology">
<paragraph blockId="10.[140,1108,167,862]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[185,283,729,755]" pageId="10" pageNumber="41">
<emphasis box="[185,278,729,755]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Biology</emphasis>
.
</emphasis>
This species has been collected from berlesed litters of oak, rotten wood, mixed oak/conifer, and mixed hardwood. This species apparently favors moist habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="distribution">
<paragraph blockId="10.[140,1108,167,862]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[185,344,800,826]" pageId="10" pageNumber="41">
<emphasis box="[185,339,800,826]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Distribution</emphasis>
.
</emphasis>
This species has been collected in the mid-elevations of the central Sierra
<collectingRegion box="[212,300,835,861]" country="United States of America" name="Nevada" pageId="10" pageNumber="41">Nevada</collectingRegion>
, from localities in Amador and Calaveras Counties (Fig. 6).
</paragraph>
</subSubSection>
</treatment>
</document>
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