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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.15.3084" ID-PMC="PMC3492929" ID-Pensoft-Pub="1314-2003-15-1" ID-Pensoft-UUID="FF9AC356FFDAFB26FF95FFE3FFA1520C" ID-PubMed="23185125" ID-Zenodo-Dep="576134" ModsDocID="1314-2003-15-1" checkinTime="1451251993896" checkinUser="pensoft" docAuthor="Soreng, Robert J. &amp; Peterson, Paul M." docDate="2012" docId="13C109C19C905203B6BE95C5E848A7F3" docLanguage="en" docName="PhytoKeys 15: 1-104" docOrigin="PhytoKeys 15" docPubDate="2012-08-06" docSource="http://dx.doi.org/10.3897/phytokeys.15.3084" docTitle="Poa secunda subsp. secunda J. Presl. Reliq. Haenk. 1 (4 5): 271 1830. 1830" docType="treatment" docVersion="3" id="FF9AC356FFDAFB26FF95FFE3FFA1520C" lastPageNumber="60" masterDocId="FF9AC356FFDAFB26FF95FFE3FFA1520C" masterDocTitle="Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: new records, re-evaluation of P. ruprechtii, and two new species, P. palmeri and P. wendtii" masterLastPageNumber="104" masterPageNumber="1" pageNumber="58" updateTime="1668139833991" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: new records, re-evaluation of P. ruprechtii, and two new species, P. palmeri and P. wendtii</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soreng, Robert J.</mods:namePart>
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peterson, Paul M.</mods:namePart>
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-08-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>15</mods:number>
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<mods:start>1</mods:start>
<mods:end>104</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.15.3084</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.15.3084</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-15-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF9AC356FFDAFB26FF95FFE3FFA1520C</mods:identifier>
<mods:identifier type="Zenodo-Dep">576134</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182229308" LSID="urn:lsid:plazi:treatment:13C109C19C905203B6BE95C5E848A7F3" httpUri="http://treatment.plazi.org/id/13C109C19C905203B6BE95C5E848A7F3" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
<subSubSection pageId="57" pageNumber="58" type="nomenclature">
<paragraph pageId="57" pageNumber="58">
19.
<taxonomicName LSID="13C109C1-9C90-5203-B6BE-95C5E848A7F3" authority="secunda J. Presl. Reliq. Haenk. 1 (4 - 5): 271 1830." authorityName="J. Presl. Reliq. Haenk. 1 (4 5): 271 1830." authorityYear="1830" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa secunda subsp. secunda" order="Poales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="subSpecies" species="secunda" subSpecies="secunda">Poa secunda secunda J. Presl. Reliq. Haenk. 1(4-5): 271 1830.</taxonomicName>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. A-G Poa secunda J. Presl. subsp. secunda A habit B sheath, ligule, blade C inflorescence D spikelet E floret F palea G anthers H-L Poa strictiramea Hitchc. H basal tuft I sheath, ligule, blade, ventral view J inflorescence K spikelet L floret. Drawings A-L from Soreng (2007) H, J, L originally drawn from Swallen 1110 as Poa involuta in Hitchcock (1935)." httpUri="https://binary.pensoft.net/fig/10155" pageId="57" pageNumber="58">Fig. 20 A-G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="type">
<paragraph pageId="57" pageNumber="58">Type:</paragraph>
<paragraph pageId="57" pageNumber="58">
Chile, Cordilleris Chilensibus, 1790,
<emphasis italics="true" pageId="57" pageNumber="58">T.Haenke</emphasis>
(holotype: PR!; isotypes: B, BAA-2691! fragm. ex B, GH! fragm., LE!, LE-TRIN-2625.01a!, MO-209304!, US-88729! fragm. ex PR).
<taxonomicName authorityName="Vasey" authorityYear="1887" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="orcuttiana">
<emphasis italics="true" pageId="57" pageNumber="58">Poa orcuttiana</emphasis>
</taxonomicName>
Vasey, W. Amer. Sci. 3: 165-166. 1887. Type: USA, California, near San Diego, 26 May 1884,
<emphasis italics="true" pageId="57" pageNumber="58">C.R.Orcutt 1070</emphasis>
(holotype: US-556833!).
<taxonomicName class="Insecta" family="Delphacidae" genus="Atropis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="scabrella">
<emphasis italics="true" pageId="57" pageNumber="58">Atropis scabrella</emphasis>
</taxonomicName>
Thurb., Bot. California 2: 310-311. 1880.).
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="57" pageNumber="58">Poa scabrella</emphasis>
</taxonomicName>
(Thurb.) Benth. ex Vasey, Grass. U.S. 42. 1883.
<taxonomicName authorityName="Kuntze" authorityYear="1891" baseAuthorityName="Thurb." class="Monocots" family="Poaceae" genus="Panicularia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="57" pageNumber="58" phylum="Magnoliophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="57" pageNumber="58">Panicularia scabrella</emphasis>
</taxonomicName>
(Thurb.) Kuntze, Revis. Gen. Pl. 2: 783. 1891.
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Puccinellia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="57" pageNumber="58" phylum="Angiospermae" rank="species" species="scabrella">
<emphasis italics="true" pageId="57" pageNumber="58">Puccinellia scabrella</emphasis>
</taxonomicName>
(Thurb.) Ponert, Feddes Repert. 84(9-10): 740. 1974. Type: USA, California, Oakland,
<emphasis italics="true" pageId="57" pageNumber="58">H.N.Bolander</emphasis>
(holotype: NY! ex herb. Nash ex herb. Thurber; isotype: US-556836! fragm. ex herb. Nash ex herb. Thurber &amp; photo).
</paragraph>
</subSubSection>
<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph pageId="57" pageNumber="58">Description.</paragraph>
<paragraph lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
Hermaphroditic.
<emphasis bold="true" pageId="57" pageNumber="58">Perennials</emphasis>
; tufted, tufts dense, narrow to medium girth, tiny to medium height (2 to over 20 cm tall), usually narrow based, green or bluish-grey-green; tillers intravaginal (each subtended by a single elongated, 2-keeled, longitudinally split prophyll), without cataphyllous shoots, sterile shoots more numerous than flowering shoots.
<emphasis bold="true" pageId="57" pageNumber="58">Culms</emphasis>
(10-)15-80(-100) cm tall, erect or bases slightly decumbent, slender to stout, leaves mostly basal, terete or weakly compressed; nodes terete, 0-2 exerted.
<emphasis bold="true" pageId="57" pageNumber="58">Leaf</emphasis>
sheaths terete, scabrous, glabrous; butt sheaths papery, smooth, glabrous; flag leaf sheaths (7-)10-20(-25) cm long, margins fused 10-25% the length, (0.95-)1.5-7(-15)
<normalizedToken originalValue="×">x</normalizedToken>
long as its blade; collars smooth or scabrous, glabrous; ligules of culm leaves 2-6(-10) mm long, of sterile tillers mostly 2-6 mm long, abaxially smooth or scabrous, apices truncate to acuminate, sterile shoot ligules similar to those of the culm leaves, blades of cauline leaves 0.4-3 mm wide, flat, folded, thin, soft, and soon withering, lax. smooth, or scabrous mainly over veins, glabrous, narrowly prow-tipped; blades gradually reduced distally or the
<pageBreakToken pageId="58" pageNumber="59" start="start">middle</pageBreakToken>
blades, flag leaf blades 0.8-10(-17) cm long; sterile shoot blades similar in form to cauline blades.
<emphasis bold="true" pageId="58" pageNumber="59">Panicles</emphasis>
2-15(-20) cm long, erect, contracted at maturity, narrowly lanceoloid to ovoid, usually congested or moderately congested (except in flowering), with 10 to over 100 spikelets; rachis usually with 1-3 branches per node; primary branches erect or ascending, infrequently widely spreading at maturity. terete to weakly angled, on and between angles usually sparsely to densely scabrous; lateral pedicels 1/6-1/3 the spikelet length, moderately to somewhat densely scabrous, prickles fairly coarse; longest branches (0.5-)1-8(-10) cm, with (1-)2-20(-60+) spikelets in distal 2/3-1/2.
<emphasis bold="true" pageId="58" pageNumber="59">Spikelets</emphasis>
, (4-)5-8 mm long, (3.8-)4-5
<normalizedToken originalValue="×">x</normalizedToken>
long as wide, usually narrowly lanceolate, sub- terete to weakly laterally compressed, not bulbiferous, drab, green or strongly anthocyanic, sometimes glaucous; florets (2-)3-5(-10), hermaphroditic; rachilla internodes terete or slightly dorsaventrally compressed, usually 1-2 mm long, smooth or muriculate to scabrous or hirtellous; glumes lanceolate to broadly lanceolate, slightly unequal, keels indistinct, keel and upper sides scabrous, apex acute to acuminate, lower glumes (2.5-)3-3.5 mm long, 3-veined; upper glumes 3.5-4 mm long, 3-veined; calluses glabrous, or with a crown of hairs around the base of the lemma, hairs 0.1-0.5(-2) mm long, crisp or slightly sinuous; lemmas 3.5-6 mm long, lanceolate to narrowly lanceolate or slightly oblanceolate, usually weakly keeled, lemma keels and marginal veins short villous to crisply puberulent or softly puberulent over proximal 2/3, between veins usually at least sparsely crisply puberulent or softly puberulent, hairs usually less than 0.5 mm long; intermediate veins obscure, margins strongly inrolled below, broadly scarious above, apices obtuse to broadly acute, blunt or sometimes pointed; paleas scabrous along the length or medially often short villous to softly puberulent over the keels, intercostal region often softly puberulent.
<emphasis bold="true" pageId="58" pageNumber="59">Flowers</emphasis>
chasmogamous; lodicules 0.55-0.8 mm long, broadly lanceolate, with a well developed lateral lobe nearly equaling the main lobe; anthers 1.5-3 mm long.
<emphasis bold="true" pageId="58" pageNumber="59">Caryopses</emphasis>
2.2 mm long, fusiform in side-view, round on back, olivaceous, weakly sulcate, hilum 0.25 mm long, oval, grain free from the palea. 2
<emphasis italics="true" pageId="58" pageNumber="59">n</emphasis>
= 42, 44, 56, 63, ca. 68, 70, ca. 72, ca. 74, ca. 78, ca. 80, 82, ca. 83, 84, 85, 86, ca. 87, ca. 88, ca. 90, ca. 91, 93, ca. 94, ca. 98, ca. 99.
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="distribution">
<paragraph pageId="58" pageNumber="59">Distribution.</paragraph>
<paragraph pageId="58" pageNumber="59">This species is widespread in North America: Canada (all Provinces except New Brunswick, Nova Scotia, Nunavut, PEI); USA (Alaska, and all western states); Mexico (Baja California); South America (Argentina and Chile).</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="ecology">
<paragraph pageId="58" pageNumber="59">Ecology.</paragraph>
<paragraph pageId="58" pageNumber="59">
The species occurs in coastal chaparral communities to coniferous forests at over 1000 m elevation in Baja California. It is high polyploid (
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Phytologia" pageId="69" pageNumber="70" pagination="390 - 413" refId="B44" refString="Soreng, RJ, 1991a. Notes on new subspecific taxa and hybrids in North American Poa (Poaceae). Phytologia 71: 390 - 413" title="Notes on new subspecific taxa and hybrids in North American Poa (Poaceae)." volume="71" year="1991 a">Soreng 1991a</bibRefCitation>
) and predominantly apomictic over much of its geographic range (
<bibRefCitation author="Kellogg, EA" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" publicationUrl="10.2307/2444319" refId="B28" refString="Kellogg, EA, 1987. Apomixis in the Poa secunda complex. American Journal of Botany 74: 1431-1437. ., 10.2307/2444319" title="Apomixis in the Poa secunda complex. American Journal of Botany 74: 1431 - 1437." url="10.2307/2444319" year="1987">Kellogg 1987</bibRefCitation>
). Flowering March to May in B.C.
</paragraph>
</subSubSection>
<subSubSection lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="specimens examined">
<paragraph pageId="58" pageNumber="59">Specimens examined.</paragraph>
<paragraph lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
Mexico.
<emphasis bold="true" pageId="58" pageNumber="59">Baja California:</emphasis>
Municipio Ensenada, Arroyo de Agua Caliente, al E de Ojos Negros, 1 May 1981, R.
<normalizedToken originalValue="Guzmán-M">Guzman-M</normalizedToken>
. 1341 (MEXU). Camino entre rancho
<normalizedToken originalValue="Mikes">Mike's</normalizedToken>
y San Jose Mellin, R.
<normalizedToken originalValue="Guzmán-M">Guzman-M</normalizedToken>
. 1414 (MEXU). Tia Juana Valley, 6 Apr 1882, C.G.Pringle 37 (LL, US, US). Todos Santos Bay, Apr 1882, F.E.Fish 28 (US); ditto, 30 (US); ditto, 33 (US). Canutillas Mts., 1884, C.R.Orcutt 1148 (US). Guadalupe Canyon, 10 Apr 1885, C.R.Orcutt 1269b (US). Carisito
<pageBreakToken pageId="59" pageNumber="60" start="start">,</pageBreakToken>
8 Apr 1885, C.R.Orcutt s.n. (US). near Vallecito, 5 Apr 1886, C.R.Orcutt 1440 (MO, US); South slope 8 miles from Rosario on road to El Marmol, 4 Mar 1930, I.L.Wiggins 4338 (LL, US). 11 mi E of Tecate on road to Mexicali, 3600 ft [1095 m], 23 Dec 1971, A.A.Beetle M-1403 (TAES); Humorosa, 4270 ft [m], 15 Apr 1973, A.A.Beetle M-2710 (TAES). 5 mi E of Humorosa, 3000 ft [910 m], 15 Apr 1973, A.A.Beetle M-2699 (TAES). Just east of Rancho Cuevas, 5 mi S of Rosarito,
<geoCoordinate degrees="32" direction="north" minutes="17" orientation="latitude" precision="925" value="32.283333">32°17'N</geoCoordinate>
,
<geoCoordinate degrees="117" direction="west" minutes="01" orientation="longitude" precision="925" value="-117.01667">117°01'W</geoCoordinate>
, 60 m, 30 Mar 1975, R.Moran 21704 (TAES). Sierra Blanca, east of main summit,
<geoCoordinate degrees="32" direction="north" minutes="03.5" orientation="latitude" precision="92" value="32.058334">32°03.5'N</geoCoordinate>
,
<geoCoordinate degrees="116" direction="west" minutes="30.5" orientation="longitude" precision="92" value="-116.50833">116°30.5'W</geoCoordinate>
, 1175 m, 16 May 1976, R.Moran 23237 (TAES). Sierra
<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
, 12 May 1997, A.Carrillo-S. 96 (MEXU). Laguna Hansen, 1 May 1981, R.
<normalizedToken originalValue="Guzmán-M">Guzman-M</normalizedToken>
. 1351 (MEXU). Sierra
<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
, 5 km W of La Rumorosa,
<geoCoordinate degrees="32" direction="north" minutes="33" orientation="latitude" precision="925" value="32.55">32°33'N</geoCoordinate>
,
<geoCoordinate degrees="116" direction="west" minutes="06" orientation="longitude" precision="925" value="-116.1">116°06'W</geoCoordinate>
, 1325m, 15 May 1977, R.Moran 24100 (TAES). Rancho El Potrereo, 40 km al SW de el Observatorio de Sierra San Pedro
<normalizedToken originalValue="Mártir">Martir</normalizedToken>
, 1050 m, P.Tenorio-L. 13218 &amp; C.Romero-deT. (MEXU). Sierra San Pedro
<normalizedToken originalValue="Mártir">Martir</normalizedToken>
, arroyo 3 km e of El Soccoro,
<geoCoordinate degrees="30" direction="north" minutes="58.4" orientation="latitude" precision="92" value="30.973333">30°58.4'N</geoCoordinate>
,
<geoCoordinate degrees="115" direction="west" minutes="37.8" orientation="longitude" precision="92" value="-115.63">115°37.8'W</geoCoordinate>
, 1450 m, 8 May 1978, R.Moran 25702 (TAES).
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="discussion">
<paragraph pageId="59" pageNumber="60">Discussion.</paragraph>
<paragraph pageId="59" pageNumber="60">
<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" refId="B25" refString="Hitchcock, AS, 1913. Mexican grasses in the United States National Herbarium. Contributions from the United States National Herbarium 17(3): 181-389." title="Mexican grasses in the United States National Herbarium. Contributions from the United States National Herbarium 17 (3): 181 - 389." year="1913">Hitchcock (1913</bibRefCitation>
,
<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" publicationUrl="10.5962/bhl.title.5731" refId="B26" refString="Hitchcock, AS, 1935. Manual of the Grasses of the United States. United States Government Printing Office, Washington, DC. ., 10.5962/bhl.title.5731" title="Manual of the Grasses of the United States. United States Government Printing Office, Washington, DC." url="10.5962/bhl.title.5731" year="1935">1935</bibRefCitation>
) accepted
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="59" pageNumber="60">Poa scabrella</emphasis>
</taxonomicName>
with
<taxonomicName authorityName="Vasey" authorityYear="1887" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="orcuttiana">
<emphasis italics="true" pageId="59" pageNumber="60">Poa orcuttiana</emphasis>
</taxonomicName>
as a synonym.
<bibRefCitation author="Espejo Serna, A" journalOrPublisher="Euphytica" pageId="69" pageNumber="70" refId="B12" refString="Espejo Serna, A, Lopez-Ferrari, AR, Valdes-Reyna, J, 2000. Poaceae.In: A. Espejo Serna A, Lopez-Ferrari AR (Eds) Las Monocotiledoneas Mexicanas: una Sinopsis Floristica, Partes IX-XI. Consejo Nacional de la Flora de Mexico, A.C., Universidad Autonoma Metropolitana-Iztapalapa, and Comision Nacional para el Conocimiento y uso de la Biodiversidad, Mexico, D.F. 10: 8-236." title="Poaceae. In: A. Espejo Serna A, Lopez-Ferrari AR (Eds) Las Monocotiledoneas Mexicanas: una Sinopsis Floristica, Partes IX-XI. Consejo Nacional de la Flora de Mexico, A. C., Universidad Autonoma Metropolitana-Iztapalapa, and Comision Nacional para el Conocimiento y uso de la Biodiversidad, Mexico, D. F. 10: 8 - 236." year="2000">Espejo Serna et al. (2000)</bibRefCitation>
accepted
<taxonomicName authorityName="Vasey" authorityYear="1887" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="orcuttiana">
<emphasis italics="true" pageId="59" pageNumber="60">Poa orcuttiana</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="59" pageNumber="60">Poa scabrella</emphasis>
</taxonomicName>
; and
<bibRefCitation author="Davila Aranda, P" journalOrPublisher="Euphytica" pageId="69" pageNumber="70" refId="B11" refString="Davila Aranda, P, Mejia-Saules, MT, Gomez-Sanchez, M, Valdes-Reyna, J, Ortiz, JJ, Morin, C, Castrejon, J, Ocampo, A, 2006. Catalogo de las Gramineas de Mexico. i-xii, 1-671 + errata." title="Catalogo de las Gramineas de Mexico. i-xii, 1 - 671 + errata." year="2006">
<normalizedToken originalValue="Dávila">Davila</normalizedToken>
Aranda et al. (2006)
</bibRefCitation>
accepted
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="59" pageNumber="60">Poa scabrella</emphasis>
</taxonomicName>
as synonym of
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
subsp.
<emphasis italics="true" pageId="59" pageNumber="60">secunda</emphasis>
but
<taxonomicName authorityName="Vasey" authorityYear="1887" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="orcuttiana">
<emphasis italics="true" pageId="59" pageNumber="60">Poa orcuttiana</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
subsp.
<emphasis italics="true" pageId="59" pageNumber="60">juncifolia</emphasis>
. In our opinion
<taxonomicName authorityName="Vasey" authorityYear="1887" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="orcuttiana">
<emphasis italics="true" pageId="59" pageNumber="60">Poa orcuttiana</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="59" pageNumber="60">Poa scabrella</emphasis>
</taxonomicName>
should be placed as synonyms of
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
subsp.
<emphasis italics="true" pageId="59" pageNumber="60">secunda</emphasis>
(
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Phytologia" pageId="69" pageNumber="70" pagination="390 - 413" refId="B44" refString="Soreng, RJ, 1991a. Notes on new subspecific taxa and hybrids in North American Poa (Poaceae). Phytologia 71: 390 - 413" title="Notes on new subspecific taxa and hybrids in North American Poa (Poaceae)." volume="71" year="1991 a">Soreng 1991a</bibRefCitation>
,
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Madrono" pageId="69" pageNumber="70" refId="B50" refString="Soreng, RJ, 2007. Poa L. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Flora of North America North of Mexico, vol. 24. Oxford University Press, New York, 486-601." title="Poa L. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Flora of North America North of Mexico, vol. 24. Oxford University Press, New York, 486 - 601." year="2007">2007</bibRefCitation>
). There is some question regarding the geographic origin of the type of
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
. It could be from California, and it seems unlikely that Haenke would have missed it there, whereas it is uncommon, at least today, in the southern Andes. There is no doubt in our minds that the North and South American plants are
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
subsp.
<emphasis italics="true" pageId="59" pageNumber="60">secunda</emphasis>
(DNA sequences from South American specimens align with those from North America; Gillespie pers. comm.). If
<taxonomicName authorityName="Benth ex Vasey" authorityYear="1883" baseAuthorityName="Thurb." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="scabrella">
<emphasis italics="true" pageId="59" pageNumber="60">Poa scabrella</emphasis>
</taxonomicName>
formswere to be recognized, it would be best to treat them as a varietyof
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
subsp.
<emphasis italics="true" pageId="59" pageNumber="60">secunda</emphasis>
, but this is not proposed here because it would only cause confusion selecting the correct name at the varietal rank among the many synonyms for this taxon. For this species
<bibRefCitation author="Gould, FW" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" refId="B20" refString="Gould, FW, Moran, R, 1981. The grasses of Baja California, Mexico. San Diego Society of Natural History, Memoir 12:1-140." title="The grasses of Baja California, Mexico. San Diego Society of Natural History, Memoir 12: 1 - 140." year="1981">Gould and Moran (1981)</bibRefCitation>
cite Isla Guadalupe as a location, possibly based on
<emphasis italics="true" pageId="59" pageNumber="60">R.Moran 13816</emphasis>
(TAES) but this specimen is
<taxonomicName authorityName="P. Beauv" authorityYear="1812" baseAuthorityName="Lam." class="Monocotyledoneae" family="Poaceae" genus="Koeleria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="59" pageNumber="60" phylum="Angiospermae" rank="species" species="pyramidata">
<emphasis italics="true" pageId="59" pageNumber="60">Koeleria pyramidata</emphasis>
</taxonomicName>
Lam. [incl.
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Koeleria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="59" pageNumber="60" phylum="Angiospermae" rank="species" species="cristata">
<emphasis italics="true" pageId="59" pageNumber="60">Koeleria cristata</emphasis>
</taxonomicName>
s. auct.,
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Koeleria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="59" pageNumber="60" phylum="Angiospermae" rank="species" species="macrantha">
<emphasis italics="true" pageId="59" pageNumber="60">Koeleria macrantha</emphasis>
</taxonomicName>
(Ledeb.) Schult.].
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10155" pageId="59" pageNumber="60" start="Figure 20" startId="F20">
<paragraph pageId="59" pageNumber="60">
<emphasis bold="true" pageId="59" pageNumber="60">Figure 20.</emphasis>
<emphasis bold="true" pageId="59" pageNumber="60">A-G</emphasis>
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="secunda">
<emphasis italics="true" pageId="59" pageNumber="60">Poa secunda</emphasis>
</taxonomicName>
J.Presl. subsp.
<emphasis italics="true" pageId="59" pageNumber="60">secunda</emphasis>
<emphasis bold="true" pageId="59" pageNumber="60">A</emphasis>
habit
<emphasis bold="true" pageId="59" pageNumber="60">B</emphasis>
sheath, ligule, blade
<emphasis bold="true" pageId="59" pageNumber="60">C</emphasis>
inflorescence
<emphasis bold="true" pageId="59" pageNumber="60">D</emphasis>
spikelet
<emphasis bold="true" pageId="59" pageNumber="60">E</emphasis>
floret
<emphasis bold="true" pageId="59" pageNumber="60">F</emphasis>
palea
<emphasis bold="true" pageId="59" pageNumber="60">G</emphasis>
anthers
<emphasis bold="true" pageId="59" pageNumber="60">H-L</emphasis>
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="strictiramea">
<emphasis italics="true" pageId="59" pageNumber="60">Poa strictiramea</emphasis>
</taxonomicName>
Hitchc.
<emphasis bold="true" pageId="59" pageNumber="60">H</emphasis>
basal tuft
<emphasis bold="true" pageId="59" pageNumber="60">I</emphasis>
sheath,ligule, blade, ventral view
<emphasis bold="true" pageId="59" pageNumber="60">J</emphasis>
inflorescence
<emphasis bold="true" pageId="59" pageNumber="60">K</emphasis>
spikelet
<emphasis bold="true" pageId="59" pageNumber="60">L</emphasis>
floret. Drawings
<emphasis bold="true" pageId="59" pageNumber="60">A-L</emphasis>
from
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Madrono" pageId="69" pageNumber="70" refId="B50" refString="Soreng, RJ, 2007. Poa L. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Flora of North America North of Mexico, vol. 24. Oxford University Press, New York, 486-601." title="Poa L. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Flora of North America North of Mexico, vol. 24. Oxford University Press, New York, 486 - 601." year="2007">Soreng (2007)</bibRefCitation>
<emphasis bold="true" pageId="59" pageNumber="60">H, J, L</emphasis>
originally drawn from
<emphasis italics="true" pageId="59" pageNumber="60">Swallen 1110</emphasis>
as
<taxonomicName authorityName="Hitchc" authorityYear="1928" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="involuta">
<emphasis italics="true" pageId="59" pageNumber="60">Poa involuta</emphasis>
</taxonomicName>
in
<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" publicationUrl="10.5962/bhl.title.5731" refId="B26" refString="Hitchcock, AS, 1935. Manual of the Grasses of the United States. United States Government Printing Office, Washington, DC. ., 10.5962/bhl.title.5731" title="Manual of the Grasses of the United States. United States Government Printing Office, Washington, DC." url="10.5962/bhl.title.5731" year="1935">Hitchcock (1935)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>