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<document id="B9569D60124E67D57DBD02CB1ED7E276" ID-CLB-Dataset="9226" ID-DOI="10.11646/zootaxa.4895.4.4" ID-GBIF-Dataset="5580248e-76a7-4521-9548-0a21b0b198ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="4358854" ID-ZooBank="14968BE9-C348-4AC9-8A1D-90D597EDD849" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608285642406" checkinUser="plazi" docAuthor="Maté Nankervis, Jason F." docDate="2020" docId="03F987E9FFADFFE8FF54F72DFA25FD58" docLanguage="en" docName="zootaxa.4895.4.4.pdf" docOrigin="Zootaxa 4895 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sindosium Johnson 2007" docType="key" docVersion="5" lastPageNumber="539" masterDocId="FFC0FF91FFA7FFE3FFC3F01FFFB9FF82" masterDocTitle="Description of two new species of Sindosium Johnson, 2007 from Australia (Coleoptera, Ptiliidae)" masterLastPageNumber="540" masterPageNumber="528" pageNumber="538" updateTime="1698847739734" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C1AEB114CB8F1601FC6506120E847479" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D5AE6C41E586224B6C96E59EF31F6237">
<mods:title id="E2C69643D725576BF5449418897B9A24">Description of two new species of Sindosium Johnson, 2007 from Australia (Coleoptera, Ptiliidae)</mods:title>
</mods:titleInfo>
<mods:name id="5D3C902EA791EB2C66FBF5EDE57F9165" type="personal">
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<mods:roleTerm id="1ACC02E98C854138A21C91B4D5AE02DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3BC727808B65F379308176DEEFBB9487">Maté Nankervis, Jason F.</mods:namePart>
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<mods:typeOfResource id="CF47D87527CCA15952B4139727B856C1">text</mods:typeOfResource>
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<mods:title id="D74E74F02E68ACC3370FAB52827D07EF">Zootaxa</mods:title>
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<mods:part id="E26494B446E2F724E5B4F7EAD9E503FB">
<mods:date id="0F3BDD60277652B71B16188A9CCCD9AF">2020</mods:date>
<mods:detail id="232DD12A45D8F1513D05C0C231E8C786" type="pubDate">
<mods:number id="945EC5075B408F5CF35B1B728DD9A95B">2020-12-17</mods:number>
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<mods:number id="CF00BD42C99B363ED10E0B607B81C40B">4895</mods:number>
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<mods:number id="A9839EC16DB4EDF9E27497E16D449BB0">4</mods:number>
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<mods:classification id="005FBF6671D8C2865E3827C49AD00771">journal article</mods:classification>
<mods:identifier id="1B9938338100256CC0A70477F5D81D62" type="CLB-Dataset">9226</mods:identifier>
<mods:identifier id="736F1142DCFB293B533C490101AF7D18" type="DOI">10.11646/zootaxa.4895.4.4</mods:identifier>
<mods:identifier id="51C13843F9B422F95CD4131414FB81F9" type="GBIF-Dataset">5580248e-76a7-4521-9548-0a21b0b198ee</mods:identifier>
<mods:identifier id="1E2C1949E8A9BFB7447FF1A6E365736A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3198B0EFB9D38F1FBF034CE36206B114" type="Zenodo-Dep">4358854</mods:identifier>
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<subSection id="E2DF2F14FFADFFE8FF54F72DFA25FD58" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538" type="multiple">
<treatment id="03F987E9FFADFFE8FF54F72DFA25FD58" ID-DOI="http://doi.org/10.5281/zenodo.4386328" ID-GBIF-Taxon="176234731" ID-Zenodo-Dep="4386328" LSID="urn:lsid:plazi:treatment:03F987E9FFADFFE8FF54F72DFA25FD58" httpUri="http://treatment.plazi.org/id/03F987E9FFADFFE8FF54F72DFA25FD58" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538">
<subSubSection id="C34A6574FFADFFE9FF54F72DFD65F8CF" box="[151,732,1842,1869]" pageId="10" pageNumber="538" type="nomenclature">
<paragraph id="8BEF36FFFFADFFE9FF54F72DFD65F8CF" blockId="10.[151,732,1842,1869]" box="[151,732,1842,1869]" pageId="10" pageNumber="538">
<heading id="D0A78193FFADFFE9FF54F72DFD65F8CF" bold="true" box="[151,732,1842,1869]" fontSize="11" level="1" pageId="10" pageNumber="538" reason="1">
<emphasis id="B924EAEDFFADFFE9FF54F72DFD65F8CF" bold="true" box="[151,732,1842,1869]" pageId="10" pageNumber="538">
Revised key to the known species of
<taxonomicName id="4C504D7CFFADFFE9FD98F72DFD65F8CF" ID-CoL="8XPW2" authorityName="Johnson" authorityYear="2007" box="[603,732,1842,1869]" class="Insecta" family="Ptiliidae" genus="Sindosium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAEDFFADFFE9FD98F72DFD65F8CF" bold="true" box="[603,732,1842,1869]" italics="true" pageId="10" pageNumber="538">Sindosium</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6574FFADFFE8FF54F764FA25FD58" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538" type="key">
<paragraph id="8BEF36FFFFADFFE9FF54F764FBF4F83B" blockId="10.[151,1436,1915,1977]" pageId="10" pageNumber="538">
The following key has been generated from the original descriptions in
<bibRefCitation id="EFC14B0EFFADFFE9FC7DF764FBD5F817" author="Johnson, C." box="[958,1132,1915,1941]" pageId="10" pageNumber="538" pagination="213 - 226" refId="ref7496" refString="Johnson, C. (2007) Studies on Ptiliidae (Col.) from the Solomon Islands, 1. Entomologists Monthly Magazine, 143, 213 - 226." type="journal article" year="2007">Johnson (2007)</bibRefCitation>
and
<bibRefCitation id="EFC14B0EFFADFFE9FB60F764FAF8F817" author="Darby, M." box="[1187,1345,1915,1941]" pageId="10" pageNumber="538" pagination="43 - 58" refId="ref6980" refString="Darby, M. (2019 a) Studies of Ptiliidae (Coleoptera) in the Spirit Collection of the Natural History Museum, London 4: new records and species collected by M. E. Bacchus and others in Sabah (Borneo) in 1977. Entomologist's Monthly Magazine, 155 (1), 43 - 58. https: // doi. org / 10.31184 / M 00138908.1551.3964" type="journal article" year="2019">Darby (2019a</bibRefCitation>
,
<bibRefCitation id="EFC14B0EFFADFFE9FA8EF764FA2CF814" author="Darby, M." box="[1357,1429,1915,1942]" pageId="10" pageNumber="538" pagination="1 - 50" refId="ref7052" refString="Darby, M. (2019 b) New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum. London, European Journal of Taxonomy, 512, 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">2019b</bibRefCitation>
) as well as from the available material pertaining to the new species described herein.
</paragraph>
<key id="FFC161CEFFADFFE8FF54F7FBFA25FD58" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538">
<keyStep id="30A42E5AFFADFFE8FF54F7FBFA25FF2F" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538">
<paragraph id="8BEF36FFFFADFFE9FF54F7FBFA25F87B" blockId="10.[151,1436,2020,2042]" box="[151,1436,2020,2042]" pageId="10" pageNumber="538">
<keyLead id="30A195CAFFADFFE9FF54F7FBFA25F87B" box="[151,1436,2020,2042]" pageId="10" pageNumber="538">1 Mesoventral collar with a pair of deep pubescent foveae posteriorly............................................. 2</keyLead>
</paragraph>
<paragraph id="8BEF36FFFFACFFE8FF54F087FA25FF2F" blockId="11.[151,1437,151,731]" box="[151,1436,151,173]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F087FA25FF2F" box="[151,1436,151,173]" pageId="11" pageNumber="539">- Mesoventral collar without deep foveae or at most just impressed............................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A42E5AFFACFFE8FF54F0AAFA25FEA0" pageId="11" pageNumber="539">
<paragraph id="8BEF36FFFFACFFE8FF54F0AAFA25FF65" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F0AAFA25FF65" pageId="11" pageNumber="539">2 Pronotum evenly convex, without any impressions or foveae on the disc, clearly punctate throughout, though puncturation may be sparse on disc or absent in a medial strip................................................................ 3</keyLead>
</paragraph>
<paragraph id="8BEF36FFFFACFFE8FF54F0EFFA25FEA0" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F0EFFA25FEA0" pageId="11" pageNumber="539">
- Pronotum impunctate, with a conspicuous medial fovea. Larger species, body length:
<quantity id="4CA89B1AFFACFFE8FBDAF0F0FB27FE87" box="[1049,1182,239,261]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="11.2" metricValueMin="8.6" pageId="11" pageNumber="539" unit="mm" value="0.99" valueMax="1.12" valueMin="0.86">0.86–1.12mm</quantity>
......................................................................................................
<taxonomicName id="4C504D7CFFACFFE8FB77F112FA25FEA0" ID-CoL="4XJMS" authority="Johnson, 2007" authorityName="Johnson" authorityYear="2007" box="[1204,1436,269,291]" class="Insecta" family="Ptiliidae" genus="Sindosium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="foveatum">
<emphasis id="B924EAEDFFACFFE8FB77F112FAB5FEA0" box="[1204,1292,269,290]" italics="true" pageId="11" pageNumber="539">foveatum</emphasis>
<bibRefCitation id="EFC14B0EFFACFFE8FAD1F112FA25FEA0" author="Johnson, C." box="[1298,1436,269,291]" pageId="11" pageNumber="539" pagination="213 - 226" refId="ref7496" refString="Johnson, C. (2007) Studies on Ptiliidae (Col.) from the Solomon Islands, 1. Entomologists Monthly Magazine, 143, 213 - 226." type="journal article" year="2007">Johnson, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A42E5AFFACFFE8FF54F135FA25FE15" pageId="11" pageNumber="539">
<paragraph id="8BEF36FFFFACFFE8FF54F135FA25FEDF" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F135FA25FEDF" pageId="11" pageNumber="539">
3 Foveolate pronotum with shorter pubescence. Mesocoxae more widely separated, equal to the width of the mesotibia. Smaller species, body length:
<quantity id="4CA89B1AFFACFFE8FE71F157FDB8FEDF" box="[434,513,328,349]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="11" pageNumber="539" unit="mm" value="0.94">0.94mm</quantity>
............................................................
<taxonomicName id="4C504D7CFFACFFE8FB1AF158FA25FEDF" ID-CoL="8Y9MD" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1241,1436,327,349]" class="Insecta" family="Ptiliidae" genus="Sindosium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="collinsi">
<emphasis id="B924EAEDFFACFFE8FB1AF158FA99FEDE" box="[1241,1312,327,348]" italics="true" pageId="11" pageNumber="539">collinsi</emphasis>
Darby, 2019
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEF36FFFFACFFE8FF54F17AFA25FE15" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F17AFA25FE15" pageId="11" pageNumber="539">- Pronotum not foveolate, pubescence long and dense throughout. Mesocoxae close, separation decidedly less than the width of the mesotibia. Larger species............................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A42E5AFFACFFE8FF54F1BFFA24FDCA" pageId="11" pageNumber="539">
<paragraph id="8BEF36FFFFACFFE8FF54F1BFFA24FE72" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F1BFFA24FE72" pageId="11" pageNumber="539">
4 Head narrower; pronotum with hind corners curved outwards, obviously projected; metaepisternum posteriorly divergent (
<figureCitation id="136B2A7AFFACFFE8FAB6F180FF46FE51" captionStart="FIGURE 1" captionStartId="3.[151,250,1959,1983]" captionTargetBox="[209,1378,181,1935]" captionTargetId="figure-26@3.[209,1378,181,1935]" captionTargetPageId="3" captionText="FIGURE 1. A. Dorsal habitus of holotype of Sindosium queenslandicum sp. nov.; B Metaventral metepisternum showing the divergent posterior suture (Mt Macartney paratype). Scale bar: 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4358858" httpUri="https://zenodo.org/record/4358858/files/figure.png" pageId="11" pageNumber="539">Fig. 1B</figureCitation>
); metacoxal bifid projection narrower, sinuate (
<figureCitation id="136B2A7AFFACFFE8FD04F1A2FCABFE51" box="[711,786,445,467]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1935]" captionTargetBox="[193,1400,181,1886]" captionTargetId="figure-15@4.[187,1400,181,1887]" captionTargetPageId="4" captionText="FIGURE 2. A. Ventral view of transparent preparation of Sindosium queenslandicum sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen showing wing folding patches in segment 2–6: D. Spermatheca; E. Aedaegus; F. Extended wing showing the structure of the peduncle and the complex venation of the wing relative to other ptiliids. All material from the Mt Macartney type locality." figureDoi="http://doi.org/10.5281/zenodo.4358862" httpUri="https://zenodo.org/record/4358862/files/figure.png" pageId="11" pageNumber="539">Fig. 2B</figureCitation>
); row of setae on 5th ventrite much denser than rest. Aedaegus (
<figureCitation id="136B2A7AFFACFFE8FAB6F1A2FF47FE72" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1935]" captionTargetBox="[193,1400,181,1886]" captionTargetId="figure-15@4.[187,1400,181,1887]" captionTargetPageId="4" captionText="FIGURE 2. A. Ventral view of transparent preparation of Sindosium queenslandicum sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen showing wing folding patches in segment 2–6: D. Spermatheca; E. Aedaegus; F. Extended wing showing the structure of the peduncle and the complex venation of the wing relative to other ptiliids. All material from the Mt Macartney type locality." figureDoi="http://doi.org/10.5281/zenodo.4358862" httpUri="https://zenodo.org/record/4358862/files/figure.png" pageId="11" pageNumber="539">Fig. 2E</figureCitation>
) &amp; spermatheca (
<figureCitation id="136B2A7AFFACFFE8FE6BF1C5FE4FFE72" box="[424,502,474,496]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1935]" captionTargetBox="[193,1400,181,1886]" captionTargetId="figure-15@4.[187,1400,181,1887]" captionTargetPageId="4" captionText="FIGURE 2. A. Ventral view of transparent preparation of Sindosium queenslandicum sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen showing wing folding patches in segment 2–6: D. Spermatheca; E. Aedaegus; F. Extended wing showing the structure of the peduncle and the complex venation of the wing relative to other ptiliids. All material from the Mt Macartney type locality." figureDoi="http://doi.org/10.5281/zenodo.4358862" httpUri="https://zenodo.org/record/4358862/files/figure.png" pageId="11" pageNumber="539">Fig. 2D</figureCitation>
). Length
<quantity id="4CA89B1AFFACFFE8FD93F1C5FD6BFE6D" box="[592,722,474,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.12" metricValueMin="0.98" pageId="11" pageNumber="539" unit="mm" value="1.05" valueMax="1.12" valueMin="0.98">0.98-1.12mm</quantity>
.........................................
<taxonomicName id="4C504D7CFFACFFE8FB04F1C5FADAFE6D" ID-CoL="8Y9MG" authority="Maté Nankervis, 2020" authorityName="Maté Nankervis" authorityYear="2020" box="[1223,1379,474,495]" class="Insecta" family="Ptiliidae" genus="Sindosium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="queenslandicum" status="sp. nov.">
<emphasis id="B924EAEDFFACFFE8FB04F1C5FADAFE6D" box="[1223,1379,474,495]" italics="true" pageId="11" pageNumber="539">queenslandicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2175796FFACFFE8FAAAF1C4FA24FE72" box="[1385,1437,475,496]" pageId="11" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BEF36FFFFACFFE8FF54F1E7FA24FDCA" blockId="11.[151,1437,151,731]" pageId="11" pageNumber="539">
<keyLead id="30A195CAFFACFFE8FF54F1E7FA24FDCA" pageId="11" pageNumber="539">
- Head wider; pronotum with hind corners straight or bent inwards, never projected; metaepisternum posteriorly narrower, partly subparallel (
<figureCitation id="136B2A7AFFACFFE8FE94F20AFE1BFDA9" box="[343,418,533,555]" captionStart="FIGURE 3" captionStartId="6.[151,250,1948,1972]" captionTargetBox="[179,1409,181,1924]" captionTargetId="figure-15@6.[177,1409,181,1924]" captionTargetPageId="6" captionText="FIGURE 3. A. Dorsal habitus of holotype of Sindosium lamingtoni sp. nov.; B. Metaventral metepisternum showing the divergent posterior suture (paratype from Lamington N.P.). Scale bar: 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4358866" httpUri="https://zenodo.org/record/4358866/files/figure.png" pageId="11" pageNumber="539">Fig. 3B</figureCitation>
); metacoxal bifid projection wider, falcate (
<figureCitation id="136B2A7AFFACFFE8FC82F20AFC35FDA9" box="[833,908,533,555]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[175,1412,181,1920]" captionTargetId="figure-26@7.[175,1412,181,1920]" captionTargetPageId="7" captionText="FIGURE 4. A. Ventral view of transparent preparation of Sindosium lamingtoni sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen; D. Spermatheca; E. Aedaegus. All material from the Lamington National Park type locality." figureDoi="http://doi.org/10.5281/zenodo.4358870" httpUri="https://zenodo.org/record/4358870/files/figure.png" pageId="11" pageNumber="539">Fig. 4B</figureCitation>
); abdominal segments with setation similar on all rows. Aedaegus (
<figureCitation id="136B2A7AFFACFFE8FE8DF22DFE2EFDCA" box="[334,407,562,584]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[175,1412,181,1920]" captionTargetId="figure-26@7.[175,1412,181,1920]" captionTargetPageId="7" captionText="FIGURE 4. A. Ventral view of transparent preparation of Sindosium lamingtoni sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen; D. Spermatheca; E. Aedaegus. All material from the Lamington National Park type locality." figureDoi="http://doi.org/10.5281/zenodo.4358870" httpUri="https://zenodo.org/record/4358870/files/figure.png" pageId="11" pageNumber="539">Fig. 4E</figureCitation>
) and spermatheca (
<figureCitation id="136B2A7AFFACFFE8FD92F22DFD27FDCA" box="[593,670,562,584]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[175,1412,181,1920]" captionTargetId="figure-26@7.[175,1412,181,1920]" captionTargetPageId="7" captionText="FIGURE 4. A. Ventral view of transparent preparation of Sindosium lamingtoni sp. nov.; B. Posterior end of metaventrite showing bifid projection; C. Abdomen; D. Spermatheca; E. Aedaegus. All material from the Lamington National Park type locality." figureDoi="http://doi.org/10.5281/zenodo.4358870" httpUri="https://zenodo.org/record/4358870/files/figure.png" pageId="11" pageNumber="539">Fig. 4D</figureCitation>
). Length
<quantity id="4CA89B1AFFACFFE8FD3AF22DFCC3FDC5" box="[761,890,562,583]" metricMagnitude="-3" metricUnit="m" metricValue="1.155" metricValueMax="1.27" metricValueMin="1.04" pageId="11" pageNumber="539" unit="mm" value="1.155" valueMax="1.27" valueMin="1.04">1.04-1.27mm</quantity>
................................
<taxonomicName id="4C504D7CFFACFFE8FB38F22DFADAFDC5" ID-CoL="8Y9MF" authority="Maté Nankervis, 2020" authorityName="Maté Nankervis" authorityYear="2020" box="[1275,1379,562,583]" class="Insecta" family="Ptiliidae" genus="Sindosium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="lamingtoni" status="sp. nov.">
<emphasis id="B924EAEDFFACFFE8FB38F22DFADAFDC5" box="[1275,1379,562,583]" italics="true" pageId="11" pageNumber="539">lamingtoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2175796FFACFFE8FAAAF22CFA24FDCA" box="[1385,1437,563,584]" pageId="11" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
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5 Pronotum sparsely and regularly punctured, laterally evenly explanate. Elytra opaque, with iridiscent sheen. Smaller species,
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- Pronotum sparsely punctured except for median bare strip, with its sides strongly and unevenly explanate, with posterior lateral areas much widened, its boders serrate. Elytra neither opaque nor iridescent. Larger species,
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Darby, 2019
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