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<mods:title id="3FFC9D63A386752F47DF2B300ABE55D9">On the blue butterfly taxa established by L. G. Courvoisier with the type locality “ Narün ” (Lepidoptera, Lycaenidae)</mods:title>
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<treatment id="03F987C3A64E8A0AFF63428B7D56F9B3" ID-DOI="http://doi.org/10.5281/zenodo.5974748" ID-GBIF-Taxon="145927819" ID-Zenodo-Dep="5974748" LSID="urn:lsid:plazi:treatment:03F987C3A64E8A0AFF63428B7D56F9B3" httpUri="http://treatment.plazi.org/id/03F987C3A64E8A0AFF63428B7D56F9B3" lastPageNumber="590" pageId="1" pageNumber="590">
<subSubSection id="C34A655EA64E8A0AFF63428B7E96FB53" pageId="1" pageNumber="590" type="nomenclature">
<paragraph id="8BEF36D5A64E8A0AFF63428B7858FB73" blockId="1.[151,1436,1123,1631]" box="[151,1037,1156,1183]" pageId="1" pageNumber="590">
<heading id="D0A781B9A64E8A0AFF63428B7858FB73" bold="true" box="[151,1037,1156,1183]" fontSize="11" level="1" pageId="1" pageNumber="590" reason="1">
<emphasis id="B924EAC7A64E8A0AFF63428B7858FB73" bold="true" box="[151,1037,1156,1183]" pageId="1" pageNumber="590">
[
<taxonomicName id="4C504D56A64E8A0AFF55428A7DEDFB72" ID-CoL="9F5Z9" authority="Phlaeas" authorityName="Phlaeas" box="[161,440,1156,1182]" class="Insecta" family="Lycaenidae" genus="Chrysophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAC7A64E8A0AFF55428A7DEDFB72" bold="true" box="[161,440,1156,1182]" italics="true" pageId="1" pageNumber="590">Chrysophanus Phlaeas</emphasis>
</taxonomicName>
] F.
<taxonomicName id="4C504D56A64E8A0AFE13428A7F7BFB73" authority="Courvoisier, 1911" authorityName="in Courvoisier" authorityYear="1911" box="[487,814,1156,1183]" class="Insecta" family="Lycaenidae" genus="Phengaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="subSpecies" species="cyanecula" subSpecies="naruena">
<emphasis id="B924EAC7A64E8A0AFE13428A7E1BFB72" bold="true" box="[487,590,1156,1182]" italics="true" pageId="1" pageNumber="590">naruena</emphasis>
Courvoisier, 1911
</taxonomicName>
(
<figureCitation id="136B2A50A64E8A0AFCC9428B7F91FB73" box="[829,964,1157,1183]" captionStart="FIGURES 115" captionStartId="3.[151,258,1626,1648]" captionTargetBox="[151,1436,194,1602]" captionTargetId="figure@3.[151,1436,194,1602]" captionTargetPageId="3" captionText="FIGURES 115. Polyommatus, Plebejus, Phengaris and Lycaena specimens. 1, 2: Polyommatus venus (Staudinger, 1886), lectotype male, upperside (1), underside (2). 3: Phengaris cyanecula (Eversmann, 1848), male, Russia, Bashkortostan, Yantysh village, upperside (right side) and underside (left side). 4, 5: Polyommatus icarus wiskotti (Courvoisier, 1911), lectotype male, upperside (4), underside (5). 6: Phengaris cyanecula (Eversmann, 1848), male, Kyrgyzstan, Suusamyr valley, Sary-Kaiky gorge, upperside (right side) and underside (left side). 7, 8: P. cyanecula naruena (Courvoisier, 1911), paralectotype male, upperside (7), underside (8). 9, 12: P. cyanecula naruena (Courvoisier, 1911), lectotype female, upperside (9), underside (12). 10, 11: Plebejus idas naruena (Courvoisier, 1913), lectotype male, upperside (10), underside (11). 13: P. idas (Linnaeus, 1761), male, Kyrgyzstan, South shore of Issyk-Kul lake, Kichi-Dzhargylchak environs, upperside (left side), underside (right side). 14, 15: Lycaena phlaeas naruena (Courvoisier, 1911), lectotype male, upperside (14), underside (15)." httpUri="https://zenodo.org/record/1285006/files/figure.png" pageId="1" pageNumber="590">figs. 14, 15</figureCitation>
,
<figureCitation id="136B2A50A64E8A0AFC25428B7850FB73" box="[977,1029,1157,1183]" captionStart="FIGURE 16" captionStartId="4.[151,244,988,1010]" captionTargetBox="[151,1436,191,967]" captionTargetId="figure@4.[151,1436,191,967]" captionTargetPageId="4" captionText="FIGURE 16. The designated lecotype labels. A: Lycaena phlaeas naruena (Courvoisier, 1911); B: Polyommatus icarus wiskotti (Courvoisier, 1911); C: Phengaris cyanecula naruena (Courvoisier, 1911); D: Plebejus idas naruena (Courvoisier, 1913)." httpUri="https://zenodo.org/record/1285008/files/figure.png" pageId="1" pageNumber="590">16A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36D5A64E8A0AFF6342A67E96FB53" blockId="1.[151,1436,1123,1631]" box="[151,707,1192,1215]" pageId="1" pageNumber="590">
<bibRefCitation id="EFC14B24A64E8A0AFF6342A67D30FB53" box="[151,357,1192,1215]" pageId="1" pageNumber="590" refId="ref3423" refString="Courvoisier, L. G. (1911 a) Entdeckungsreisen und kritische Spaziergange ins Gebiet der Lycaeniden. Entomologische Zeitschrift Internationalen Vereins, 24 (37), 59 - 60 + 65 - 67 + 70 - 71 + 77 - 79 + 81 - 82 + 88 - 89 + 92 - 94 + 99 - 101 + 106 - 108 + 112 + 125 - 127 + 131 - 132 + 135 - 137 + 141 - 142 + 147 - 148 + 156 + 167 - 170 + 176 - 177 + 181 - 182 + 185 - 188 + 190 - 192 + 196 - 200 + 201 - 204 + 205 - 209 + 210 - 214 + 227 - 228 + 230 - 231 + 233 - 236 + 237 - 238 + 241 - 242 + 245 - 247 + 249 - 250 + 253 - 255 + 257 - 259 + 261 - 263 + 265 - 266 + 269 - 270." type="journal article">Courvoisier, 1911a</bibRefCitation>
: 262, type locality: “aus Narün”.
</paragraph>
</subSubSection>
<subSubSection id="C34A655EA64E8A0AFF3342C47D56F9B3" pageId="1" pageNumber="590" type="description">
<paragraph id="8BEF36D5A64E8A0AFF3342C47E84FACB" blockId="1.[151,1436,1123,1631]" pageId="1" pageNumber="590">
The populations of
<taxonomicName id="4C504D56A64E8A0AFE6E42C47F58FB0D" authority="(Linnaeus, 1761)" authorityName="Linnaeus" authorityYear="1761" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[410,781,1225,1249]" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="phlaeas">
<emphasis id="B924EAC7A64E8A0AFE6E42C47E1BFB0D" box="[410,590,1225,1249]" italics="true" pageId="1" pageNumber="590">Lycaena phlaeas</emphasis>
(Linnaeus, 1761)
</taxonomicName>
from the low-mountainous and middle-mountainous Central Asia were considered by some authors as nominate subspecies (
<bibRefCitation id="EFC14B24A64E8A0AFC9F42E37815FAE8" box="[875,1088,1261,1284]" pageId="1" pageNumber="590" refId="ref3983" refString="Tshikolovets, V. V. (2005) The butterflies of Kyrgyzstan Tshikolovets Press, Kyev-Brno, 511 pp." type="book">Tshikolovets, 2005</bibRefCitation>
), and by others as
<taxonomicName id="4C504D56A64E8A0AFAEB42E37DF0FACB" authority="Groum-Grshimailo, 1890" authorityName="Groum-Grshimailo" authorityYear="1890" pageId="1" pageNumber="590" rank="subSpecies" subSpecies="oxiana">
ssp.
<emphasis id="B924EAC7A64E8A0AFAA742E379C9FAE8" box="[1363,1436,1261,1284]" italics="true" pageId="1" pageNumber="590">oxiana</emphasis>
Groum-Grshimaïlo, 1890
</taxonomicName>
(
<bibRefCitation id="EFC14B24A64E8A0AFE40431E7E93FACB" box="[436,710,1296,1319]" pageId="1" pageNumber="590" refId="ref3803" refString="Korb, S. K. &amp; Bolshakov, L. V. (2016) A systematic catalogue of butterflies of the former Soviet Union (Armenia, Azerbaijan, Belarus, Estonia, Georgia, Kyrgyzstan, Kazakhstan, Latvia, Lituania, Moldova, Russia, Tajikistan, Turkmenistan, Ukraine, Uzbekistan) with special account to their type specimens (Lepidoptera: Hesperioidea, Papilionoidea). Zootaxa, 4160 (1), 1 - 324. https: // doi. org / 10.11646 / zootaxa. 4160.1.1" type="journal article">Korb &amp; Bolshakov, 2016</bibRefCitation>
).
</paragraph>
<paragraph id="8BEF36D5A64E8A0AFF33433C7F20FA63" blockId="1.[151,1436,1123,1631]" pageId="1" pageNumber="590">
The type series contain 2 syntypes. To preserve the nomenclatural stability, I designate here the lectotype of [
<taxonomicName id="4C504D56A64E8A0AFF54435A7DC3FA80" authority="Phlaeas" authorityName="Phlaeas" box="[160,406,1364,1388]" class="Insecta" family="Lycaenidae" genus="Chrysophanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAC7A64E8A0AFF54435A7DC3FA80" box="[160,406,1364,1388]" italics="true" pageId="1" pageNumber="590">Chrysophanus Phlaeas</emphasis>
</taxonomicName>
] F.
<taxonomicName id="4C504D56A64E8A0AFE4A435B7E88FA80" authority="Courvoisier, 1911" authorityName="in Courvoisier" authorityYear="1911" box="[446,733,1365,1388]" class="Insecta" family="Lycaenidae" genus="Phengaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="subSpecies" species="cyanecula" subSpecies="naruena">
<emphasis id="B924EAC7A64E8A0AFE4A435B7E42FA80" box="[446,535,1365,1388]" italics="true" pageId="1" pageNumber="590">naruena</emphasis>
Courvoisier, 1911
</taxonomicName>
, a male specimen (
<figureCitation id="136B2A50A64E8A0AFC51435B784AFA80" box="[933,1055,1365,1388]" captionStart="FIGURES 115" captionStartId="3.[151,258,1626,1648]" captionTargetBox="[151,1436,194,1602]" captionTargetId="figure@3.[151,1436,194,1602]" captionTargetPageId="3" captionText="FIGURES 115. Polyommatus, Plebejus, Phengaris and Lycaena specimens. 1, 2: Polyommatus venus (Staudinger, 1886), lectotype male, upperside (1), underside (2). 3: Phengaris cyanecula (Eversmann, 1848), male, Russia, Bashkortostan, Yantysh village, upperside (right side) and underside (left side). 4, 5: Polyommatus icarus wiskotti (Courvoisier, 1911), lectotype male, upperside (4), underside (5). 6: Phengaris cyanecula (Eversmann, 1848), male, Kyrgyzstan, Suusamyr valley, Sary-Kaiky gorge, upperside (right side) and underside (left side). 7, 8: P. cyanecula naruena (Courvoisier, 1911), paralectotype male, upperside (7), underside (8). 9, 12: P. cyanecula naruena (Courvoisier, 1911), lectotype female, upperside (9), underside (12). 10, 11: Plebejus idas naruena (Courvoisier, 1913), lectotype male, upperside (10), underside (11). 13: P. idas (Linnaeus, 1761), male, Kyrgyzstan, South shore of Issyk-Kul lake, Kichi-Dzhargylchak environs, upperside (left side), underside (right side). 14, 15: Lycaena phlaeas naruena (Courvoisier, 1911), lectotype male, upperside (14), underside (15)." httpUri="https://zenodo.org/record/1285006/files/figure.png" pageId="1" pageNumber="590">figs. 14, 15</figureCitation>
), labelled: white paper, handwritten “F.
<taxonomicName id="4C504D56A64E8A0AFF4843767D41FA63" authorityName="in Courvoisier" authorityYear="1911" box="[188,276,1400,1423]" class="Insecta" family="Lycaenidae" genus="Phengaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="subSpecies" species="cyanecula" subSpecies="naruena">naruena</taxonomicName>
. m.
<date id="FFEE1015A64E8A0AFEB443767D26FA63" box="[320,371,1400,1423]" pageId="1" pageNumber="590" value="1919-02">19ii.</date>
”; pink paper, handwritten “Narün | 1909 Wisk”.
</paragraph>
<paragraph id="8BEF36D5A64E8A0AFF3343947FB4FA5D" blockId="1.[151,1436,1123,1631]" box="[199,993,1434,1457]" pageId="1" pageNumber="590">
The other
<typeStatus id="54EB8877A64E8A0AFEC043947DD2FA5D" box="[308,391,1434,1457]" pageId="1" pageNumber="590" type="syntype">syntype</typeStatus>
specimen, a female, is designated here as
<typeStatus id="54EB8877A64E8A0AFCBF43947F8BFA5D" box="[843,990,1434,1457]" pageId="1" pageNumber="590" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
<paragraph id="8BEF36D5A64E8A0AFF3343B37D56F9B3" blockId="1.[151,1436,1123,1631]" pageId="1" pageNumber="590">
The
<typeStatus id="54EB8877A64E8A0AFF0F43B37D08FA38" box="[251,349,1469,1492]" pageId="1" pageNumber="590" type="lectotype">lectotype</typeStatus>
has no differences from European specimens (it has been figured in many sources, for example: www.lepiforum.de, www.eurobutterflies.com etc., so we do not publish the images of
<taxonomicName id="4C504D56A64E8A0AFBDD43EE78CDFA1B" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[1065,1176,1503,1527]" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="phlaeas">
<emphasis id="B924EAC7A64E8A0AFBDD43EE78CDFA1B" box="[1065,1176,1503,1527]" italics="true" pageId="1" pageNumber="590">L. phlaeas</emphasis>
</taxonomicName>
from Europe here), thus it must be synonymized with the nominate subspecies:
<taxonomicName id="4C504D56A64E8A0AFCF3400C78CAF9F5" authority="phlaeas (Linnaeus, 1761)" authorityName="phlaeas (Linnaeus" authorityYear="1761" box="[775,1183,1537,1561]" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="subSpecies" species="phlaeas" subSpecies="phlaeas">
<emphasis id="B924EAC7A64E8A0AFCF3400C7F8CF9F5" box="[775,985,1537,1561]" italics="true" pageId="1" pageNumber="590">L. phlaeas phlaeas</emphasis>
(Linnaeus, 1761)
</taxonomicName>
=
<taxonomicName id="4C504D56A64E8A0AFB30400C7D3FF9D0" authority="Courvoisier, 1911" authorityName="Courvoisier" authorityYear="1911" baseAuthorityName="Courvoisier" baseAuthorityYear="1911" class="Insecta" family="Lycaenidae" genus="Lycaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="subSpecies" species="phlaeas" subSpecies="naruena">
<emphasis id="B924EAC7A64E8A0AFB30400C79C9F9F5" box="[1220,1436,1537,1561]" italics="true" pageId="1" pageNumber="590">L. phlaeas naruena</emphasis>
(Courvoisier, 1911)
</taxonomicName>
,
<taxonomicNameLabel id="A21757BCA64E8A0AFE82402A7DECF9D0" box="[374,441,1572,1596]" pageId="1" pageNumber="590">
<emphasis id="B924EAC7A64E8A0AFE82402A7DECF9D0" bold="true" box="[374,441,1572,1596]" pageId="1" pageNumber="590">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="54EB8877A64E8A0AFE3C402B7DA2F9D0" box="[456,503,1573,1596]" pageId="1" pageNumber="590">type</typeStatus>
locality by the
<typeStatus id="54EB8877A64E8A0AFD54402B7F57F9D0" box="[672,770,1573,1596]" pageId="1" pageNumber="590" type="lectotype">lectotype</typeStatus>
designation: Narün NE from
<collectingRegion id="4994F837A64E8A0AFBB0402B78E4F9D0" box="[1092,1201,1573,1596]" country="Russia" name="Astrakhan'" pageId="1" pageNumber="590">Astrakhan</collectingRegion>
between rivers Volga and Ural).
</paragraph>
</subSubSection>
</treatment>
</document>