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<document ID-DOI="10.5733/afin.056.0209" ID-ISSN="2305-2562" ID-Zenodo-Dep="7914093" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683586032370" checkinUser="felipe" docAuthor="Jäger, Peter &amp; Krehenwinkel, Henrik" docDate="2015" docId="03F987A3ED5FFFFA3511B528FC8A07DD" docLanguage="en" docName="AfrInvertebr.56.2.365-392.pdf" docOrigin="African Invertebrates 56 (2)" docSource="http://www.bioone.org/doi/10.5733/afin.056.0209" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="May Jäger &amp; Krehenwinkel 2015, gen. n." docType="treatment" docVersion="3" lastPageNumber="369" masterDocId="FFC0FFDBED5CFFFE3742B15DFFC30150" masterDocTitle="May gen. n. (Araneae: Sparassidae): a unique lineage from southern Africa supported by morphological and molecular features" masterLastPageNumber="365" masterPageNumber="365" pageNumber="368" updateTime="1683635035534" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>May gen. n. (Araneae: Sparassidae): a unique lineage from southern Africa supported by morphological and molecular features</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jäger, Peter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krehenwinkel, Henrik</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-08-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
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<mods:start>365</mods:start>
<mods:end>365</mods:end>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.056.0209</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.056.0209</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7914093</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7914186" ID-Zenodo-Dep="7914186" LSID="urn:lsid:plazi:treatment:03F987A3ED5FFFFA3511B528FC8A07DD" httpUri="http://treatment.plazi.org/id/03F987A3ED5FFFFA3511B528FC8A07DD" lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368">
<subSubSection box="[595,728,1141,1168]" pageId="3" pageNumber="368" type="nomenclature">
<paragraph blockId="3.[181,1142,1141,1677]" box="[595,728,1141,1168]" pageId="3" pageNumber="368">
<heading box="[595,728,1141,1168]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="368" reason="2">
<taxonomicName ID-CoL="5M7T" authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[595,646,1141,1167]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[595,646,1141,1167]" italics="true" pageId="3" pageNumber="368">May</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[653,728,1142,1168]" pageId="3" pageNumber="368" rank="genus">gen. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368" type="materials_examined">
<paragraph blockId="3.[181,1142,1141,1677]" box="[181,530,1181,1208]" pageId="3" pageNumber="368">
<typeStatus box="[181,237,1182,1208]" pageId="3" pageNumber="368">Type</typeStatus>
species:
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[340,463,1181,1207]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="368" phylum="Arthropoda" rank="species" species="bruno" status="sp. n.">
<emphasis box="[340,463,1181,1207]" italics="true" pageId="3" pageNumber="368">May bruno</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[470,530,1182,1208]" pageId="3" pageNumber="368" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="3.[181,1142,1141,1677]" pageId="3" pageNumber="368">
Etymology: This new genus is named for Bruno
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[765,817,1222,1248]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus" status="gen. n.">May</taxonomicName>
for supporting biodiversity research in Africa through BIOPAT e.V. (Patrons for Biodiversity; www-biopat.de); gender masculine.
</paragraph>
<paragraph blockId="3.[181,1142,1141,1677]" lastBlockId="4.[144,1105,204,1677]" lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368">
Diagnosis: Small to medium-sized spiders belonging to the so-called “African clade” (
<bibRefCitation author="MORADMAND, M. &amp; SCHONHOFER, A. L. &amp; JAGER, P." box="[189,456,1359,1386]" pageId="3" pageNumber="368" pagination="48 - 65" refId="ref12604" refString="MORADMAND, M., SCHONHOFER, A. L. &amp; JAGER, P. 2014. Molecular phylogeny of the huntsman spider family Sparassidae with focus on the genus Eusparassus and notes on the RTA-clade and &quot; Laterigradae &quot;. Molecular Phylogenetics and Evolution 74: 48 - 65." type="journal article" year="2014">
Moradmand
<emphasis box="[335,384,1359,1385]" italics="true" pageId="3" pageNumber="368">et al</emphasis>
. 2014
</bibRefCitation>
) with a body length of
<quantity box="[717,849,1359,1386]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="3" pageNumber="368" unit="mm" value="14.0" valueMax="20.0" valueMin="8.0">8 to 20 mm</quantity>
. Distinguished from other members of this clade by the copulatory organs: males can be recognised by the course of the embolus, arising distally from the tegulum, running prolaterad (
<figureCitation box="[976,1048,1424,1450]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="3" pageNumber="368">Figs 1</figureCitation>
,
<figureCitation box="[1063,1093,1424,1451]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="3" pageNumber="368">50</figureCitation>
,
<figureCitation box="[1106,1137,1424,1450]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="3" pageNumber="368">64</figureCitation>
; retrolaterad in other genera of this clade and almost in all other members of the entire family) and the short and compact RTA(at least one branch elongated in other members). Moreover, the proximo-retrolateral cymbium exhibits a distinct “shoulder” (
<figureCitation box="[1024,1096,1521,1547]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="3" pageNumber="368">Figs 4</figureCitation>
,
<figureCitation box="[1107,1137,1521,1548]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="3" pageNumber="368">50</figureCitation>
,
<figureCitation box="[181,211,1553,1580]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="3" pageNumber="368">59</figureCitation>
,
<figureCitation box="[223,253,1553,1579]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="3" pageNumber="368">67</figureCitation>
,
<figureCitation box="[264,295,1553,1580]" captionStart="Figs 70–77" captionStartId="16.[144,183,1443,1465]" captionTargetBox="[155,1091,208,1422]" captionTargetId="figure-38@16.[155,1091,204,1422]" captionTargetPageId="16" captionText="Figs 70–77.May rudy gen.n. sp. n., holotype ♂ from Copper Valley,Namibia:(70–72) habitus of preserved spider: (70) dorsal, (71) ventral, (72) frontal; (73–75) ♂ palp: (73) ventral, (74) retrolateral, (75) RTA, dorsal); (76–77) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and claw tuft penetrating dorsal setae (white arrow): (76) lateral, (77) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914123" httpUri="https://zenodo.org/record/7914123/files/figure.png" pageId="3" pageNumber="368">75</figureCitation>
), whereas such a structure is missing in other members of the African clade. Females with epigynal furrows not reaching epigastric furrow, i.e. lateral lobes fused, indicated by “fusion bubbles” (
<figureCitation box="[543,722,1618,1644]" captionStart="Figs 11–17" captionStartId="7.[181,220,1399,1421]" captionTargetBox="[192,1121,225,1381]" captionTargetId="figure-62@7.[192,1121,225,1381]" captionTargetPageId="7" captionText="Figs 11–17. May bruno gen. n. sp. n. from South Africa, ♀ paratypes from Witsand, South Africa (11–13) PJ 3524, (14–16) PJ 3525, (17) PJ 3526): (11, 14, 17) epigyne, ventral; (12, 15) vulva, dorsal; (13) vulva, posterior; (16) schematic course of internal duct system, dorsal. Abbreviations: FB – fusion bubbles; FD – fertilisation ducts; G – glands; GP – guiding pockets; SS – slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914101" httpUri="https://zenodo.org/record/7914101/files/figure.png" pageId="3" pageNumber="368">Figs 11, 14, 17</figureCitation>
,
<figureCitation box="[737,859,1618,1644]" captionStart="Figs 78–86" captionStartId="18.[144,183,1534,1556]" captionTargetBox="[144,1092,204,1516]" captionTargetId="figure-21@18.[144,1092,204,1517]" captionTargetPageId="18" captionText="Figs 78–86. May norm gen.n.sp. n.from Messum Crater, Namibia (78–80) holotype ♂ [PJ 1851]; (81–85) ♀ paratype [PJ 3534]; 86 ♀ paratype [PJ 1852]): (78, 81) epigyne, ventral (with mating plugs); (79, 82) vulva, dorsal; (84) vulva,anterior; (80, 83) schematic course of internal duct system, dorsal; (85) schematic course of internal duct system, anterior; (86) sternum, ventral.Abbreviations:FB – fusion bubbles; FD – fertilisationducts; GE – glandular extensions;GP – guiding pockets; SS – slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914127" httpUri="https://zenodo.org/record/7914127/files/figure.png" pageId="3" pageNumber="368">78, 81, 86</figureCitation>
). Serrula is dramatically reduced, i.e. in most cases entirely, in both sexes (
<figureCitation box="[736,823,1650,1676]" captionStart="Figs 20–33" captionStartId="9.[183,222,1469,1491]" captionTargetBox="[182,1141,242,1434]" captionTargetId="figure-15@9.[181,1141,242,1434]" captionTargetPageId="9" captionText="Figs 20–33.May bruno gen.n. sp. n. from SouthAfrica (♀ paratypes from Witsand,♀ from Twee Rivieren): (20–22) tarsus tip: (20) ventral, (21) lateral, (22) dorsal); (23) ♀ palpal claw, lateral; (24–25) Lawrence setae; (26–29) prosoma of ♀♀, showing positions of dorsal marks (white arrows in Fig. 26); (30) metatarsal stopper, dorsal; (31) eye arrangement, dorsal; (32) cheliceral dentition, ventral; (33) gnathocoxae, ventro-distal, showing reduced serrula (black arrow). Abbreviations:AP – plate of Lawrence setae; DP – dorsal plate of elongated setae; IP – plate of indented claw tuft setae; TS – transverse suture with slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914105" httpUri="https://zenodo.org/record/7914105/files/figure.png" pageId="3" pageNumber="368">Figs 33</figureCitation>
,
<figureCitation box="[834,907,1650,1676]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="3" pageNumber="368">60, 63</figureCitation>
; some females of the African clade also show an entire reduction, usually males and females of this group show up to 37 denticles in their serrula). Leg I shortest, not leg III as in all other
<taxonomicName authorityName="Bertkau" authorityYear="1872" box="[963,1099,236,262]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
; femora I with prolateral spines shifted into distal half, i.e. proximal prolateral spine almost reaching medial dorsal spine (
<figureCitation box="[561,634,300,326]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="4" pageNumber="369">Figs 7</figureCitation>
,
<figureCitation box="[646,676,300,326]" captionStart="Figs 34–45" captionStartId="10.[144,183,1494,1516]" captionTargetBox="[157,1089,223,1457]" captionTargetId="figure-20@10.[157,1089,223,1457]" captionTargetPageId="10" captionText="Figs 34–45. May bruno gen. n. sp. n. from Witsand, South Africa (34–36, 41–43) holotype ♂; (37–40, 44–45) ♀ paratype, PJ 3524. (34–39, 43) habitus of preserved spiders: (34, 37) dorsal; (35, 38) ventral; (36, 39, 43) frontal; (40) palp of ♀, lateral, showing rows of elongated setae; (41–42) palp of ♂: (41) ventral, (42) retrolateral; (44–45) tarsal tips, showing light-coloured indented claw tuft setae (white arrow), additional dark claw tufts with Lawrence setae (black arrow), and scopula hairs (grey arrow): (44) ventral, (45) lateral." figureDoi="http://doi.org/10.5281/zenodo.7914109" httpUri="https://zenodo.org/record/7914109/files/figure.png" pageId="4" pageNumber="369">43</figureCitation>
,
<figureCitation box="[688,718,299,326]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="4" pageNumber="369">57</figureCitation>
,
<figureCitation box="[730,761,300,326]" captionStart="Figs 70–77" captionStartId="16.[144,183,1443,1465]" captionTargetBox="[155,1091,208,1422]" captionTargetId="figure-38@16.[155,1091,204,1422]" captionTargetPageId="16" captionText="Figs 70–77.May rudy gen.n. sp. n., holotype ♂ from Copper Valley,Namibia:(70–72) habitus of preserved spider: (70) dorsal, (71) ventral, (72) frontal; (73–75) ♂ palp: (73) ventral, (74) retrolateral, (75) RTA, dorsal); (76–77) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and claw tuft penetrating dorsal setae (white arrow): (76) lateral, (77) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914123" httpUri="https://zenodo.org/record/7914123/files/figure.png" pageId="4" pageNumber="369">72</figureCitation>
,
<figureCitation box="[772,802,300,326]" captionStart="Figs 87–93" captionStartId="19.[181,220,720,742]" captionTargetBox="[181,1122,236,706]" captionTargetId="figure-15@19.[181,1141,236,725]" captionTargetPageId="19" captionText="Figs 87–93. May norm gen. n. sp. n. from Messum Crater, Namibia (87–89, 91–93) ♀ paratype [PJ 1852]; (90) holotype ♂ [PJ 1851]: (87) eye arrangement, dorsal; (88) cheliceral dentition, ventral; (89) metatarsal stopper, dorsal; (90) ♀ palpal claw, lateral; (91) Lawrence setae; (92) Claw tuft setae, showing indented tip; (93) leg I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.7914129" httpUri="https://zenodo.org/record/7914129/files/figure.png" pageId="4" pageNumber="369">93</figureCitation>
,
<figureCitation box="[814,846,300,326]" captionStart="Figs 94–99" captionStartId="19.[182,221,1582,1604]" captionTargetBox="[193,1130,877,1566]" captionTargetId="figure-73@19.[193,1130,877,1566]" captionTargetPageId="19" captionText="Figs 94–99. May norm gen. n. sp. n., paratypes from Messum Crater, Namibia (94–96) PJ 1852; (97–99) PJ 3543: habitus of preserved specimens (94, 97) dorsal; (95, 98) ventral; (96, 99) frontal." figureDoi="http://doi.org/10.5281/zenodo.7914131" httpUri="https://zenodo.org/record/7914131/files/figure.png" pageId="4" pageNumber="369">99</figureCitation>
).
</paragraph>
<paragraph blockId="4.[144,1105,204,1677]" pageId="4" pageNumber="369">
Description: Widest point of DS in posterior half, close to coxa II and III; PL/PW
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males,
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females; fovea mostly above coxa III, may be situated more anteriorly above coxa II. Sternum broadest between posterior part of coxa II to anterior part of coxa III, with 3 pairs of lateral extensions, 3 pairs of double slit sense organs plus additional single slit sense organs posteriorly, long setae on entire sternum, single pair of dense setae brushes at posterior margin (
<figureCitation box="[737,820,500,527]" captionStart="Figs 78–86" captionStartId="18.[144,183,1534,1556]" captionTargetBox="[144,1092,204,1516]" captionTargetId="figure-21@18.[144,1092,204,1517]" captionTargetPageId="18" captionText="Figs 78–86. May norm gen.n.sp. n.from Messum Crater, Namibia (78–80) holotype ♂ [PJ 1851]; (81–85) ♀ paratype [PJ 3534]; 86 ♀ paratype [PJ 1852]): (78, 81) epigyne, ventral (with mating plugs); (79, 82) vulva, dorsal; (84) vulva,anterior; (80, 83) schematic course of internal duct system, dorsal; (85) schematic course of internal duct system, anterior; (86) sternum, ventral.Abbreviations:FB – fusion bubbles; FD – fertilisationducts; GE – glandular extensions;GP – guiding pockets; SS – slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914127" httpUri="https://zenodo.org/record/7914127/files/figure.png" pageId="4" pageNumber="369">Fig. 86</figureCitation>
). Opisthosoma: oval, OL/ OW
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males,
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females. Eyes: in two rows, both rows recurved when viewed from above (
<figureCitation box="[451,525,564,591]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="4" pageNumber="369">Figs 6</figureCitation>
,
<figureCitation box="[540,570,565,591]" captionStart="Figs 20–33" captionStartId="9.[183,222,1469,1491]" captionTargetBox="[182,1141,242,1434]" captionTargetId="figure-15@9.[181,1141,242,1434]" captionTargetPageId="9" captionText="Figs 20–33.May bruno gen.n. sp. n. from SouthAfrica (♀ paratypes from Witsand,♀ from Twee Rivieren): (20–22) tarsus tip: (20) ventral, (21) lateral, (22) dorsal); (23) ♀ palpal claw, lateral; (24–25) Lawrence setae; (26–29) prosoma of ♀♀, showing positions of dorsal marks (white arrows in Fig. 26); (30) metatarsal stopper, dorsal; (31) eye arrangement, dorsal; (32) cheliceral dentition, ventral; (33) gnathocoxae, ventro-distal, showing reduced serrula (black arrow). Abbreviations:AP – plate of Lawrence setae; DP – dorsal plate of elongated setae; IP – plate of indented claw tuft setae; TS – transverse suture with slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914105" httpUri="https://zenodo.org/record/7914105/files/figure.png" pageId="4" pageNumber="369">31</figureCitation>
,
<figureCitation box="[584,614,564,591]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="4" pageNumber="369">54</figureCitation>
,
<figureCitation box="[628,658,564,590]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="4" pageNumber="369">68</figureCitation>
,
<figureCitation box="[672,703,565,591]" captionStart="Figs 87–93" captionStartId="19.[181,220,720,742]" captionTargetBox="[181,1122,236,706]" captionTargetId="figure-15@19.[181,1141,236,725]" captionTargetPageId="19" captionText="Figs 87–93. May norm gen. n. sp. n. from Messum Crater, Namibia (87–89, 91–93) ♀ paratype [PJ 1852]; (90) holotype ♂ [PJ 1851]: (87) eye arrangement, dorsal; (88) cheliceral dentition, ventral; (89) metatarsal stopper, dorsal; (90) ♀ palpal claw, lateral; (91) Lawrence setae; (92) Claw tuft setae, showing indented tip; (93) leg I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.7914129" httpUri="https://zenodo.org/record/7914129/files/figure.png" pageId="4" pageNumber="369">87</figureCitation>
). Chelicerae: with 2 anterior and 3 posterior teeth, without denticles in between them, fang base ventrally with 1 to 9 bristles (
<figureCitation box="[151,238,629,655]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="4" pageNumber="369">Figs 10</figureCitation>
,
<figureCitation box="[250,280,629,655]" captionStart="Figs 20–33" captionStartId="9.[183,222,1469,1491]" captionTargetBox="[182,1141,242,1434]" captionTargetId="figure-15@9.[181,1141,242,1434]" captionTargetPageId="9" captionText="Figs 20–33.May bruno gen.n. sp. n. from SouthAfrica (♀ paratypes from Witsand,♀ from Twee Rivieren): (20–22) tarsus tip: (20) ventral, (21) lateral, (22) dorsal); (23) ♀ palpal claw, lateral; (24–25) Lawrence setae; (26–29) prosoma of ♀♀, showing positions of dorsal marks (white arrows in Fig. 26); (30) metatarsal stopper, dorsal; (31) eye arrangement, dorsal; (32) cheliceral dentition, ventral; (33) gnathocoxae, ventro-distal, showing reduced serrula (black arrow). Abbreviations:AP – plate of Lawrence setae; DP – dorsal plate of elongated setae; IP – plate of indented claw tuft setae; TS – transverse suture with slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914105" httpUri="https://zenodo.org/record/7914105/files/figure.png" pageId="4" pageNumber="369">32</figureCitation>
,
<figureCitation box="[292,322,628,655]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="4" pageNumber="369">53</figureCitation>
,
<figureCitation box="[333,364,629,655]" captionStart="Figs 87–93" captionStartId="19.[181,220,720,742]" captionTargetBox="[181,1122,236,706]" captionTargetId="figure-15@19.[181,1141,236,725]" captionTargetPageId="19" captionText="Figs 87–93. May norm gen. n. sp. n. from Messum Crater, Namibia (87–89, 91–93) ♀ paratype [PJ 1852]; (90) holotype ♂ [PJ 1851]: (87) eye arrangement, dorsal; (88) cheliceral dentition, ventral; (89) metatarsal stopper, dorsal; (90) ♀ palpal claw, lateral; (91) Lawrence setae; (92) Claw tuft setae, showing indented tip; (93) leg I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.7914129" httpUri="https://zenodo.org/record/7914129/files/figure.png" pageId="4" pageNumber="369">88</figureCitation>
; insertion sockets only partly illustrated). Palps in both sexes with two rows of elongated setae forming functional basket (
<figureCitation box="[758,844,661,687]" captionStart="Figs 34–45" captionStartId="10.[144,183,1494,1516]" captionTargetBox="[157,1089,223,1457]" captionTargetId="figure-20@10.[157,1089,223,1457]" captionTargetPageId="10" captionText="Figs 34–45. May bruno gen. n. sp. n. from Witsand, South Africa (34–36, 41–43) holotype ♂; (37–40, 44–45) ♀ paratype, PJ 3524. (34–39, 43) habitus of preserved spiders: (34, 37) dorsal; (35, 38) ventral; (36, 39, 43) frontal; (40) palp of ♀, lateral, showing rows of elongated setae; (41–42) palp of ♂: (41) ventral, (42) retrolateral; (44–45) tarsal tips, showing light-coloured indented claw tuft setae (white arrow), additional dark claw tufts with Lawrence setae (black arrow), and scopula hairs (grey arrow): (44) ventral, (45) lateral." figureDoi="http://doi.org/10.5281/zenodo.7914109" httpUri="https://zenodo.org/record/7914109/files/figure.png" pageId="4" pageNumber="369">Figs 40</figureCitation>
,
<figureCitation box="[854,884,660,687]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="4" pageNumber="369">50</figureCitation>
,
<figureCitation box="[895,925,660,687]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="4" pageNumber="369">58</figureCitation>
,
<figureCitation box="[936,1012,660,687]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="4" pageNumber="369">64–65</figureCitation>
,
<figureCitation box="[1022,1099,661,687]" captionStart="Figs 70–77" captionStartId="16.[144,183,1443,1465]" captionTargetBox="[155,1091,208,1422]" captionTargetId="figure-38@16.[155,1091,204,1422]" captionTargetPageId="16" captionText="Figs 70–77.May rudy gen.n. sp. n., holotype ♂ from Copper Valley,Namibia:(70–72) habitus of preserved spider: (70) dorsal, (71) ventral, (72) frontal; (73–75) ♂ palp: (73) ventral, (74) retrolateral, (75) RTA, dorsal); (76–77) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and claw tuft penetrating dorsal setae (white arrow): (76) lateral, (77) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914123" httpUri="https://zenodo.org/record/7914123/files/figure.png" pageId="4" pageNumber="369">73–74</figureCitation>
; most likely to dig burrows in loose sand/transport sand out of the burrow, as known for
<taxonomicName box="[144,319,725,751]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[144,264,725,751]" italics="true" pageId="4" pageNumber="369">Cebrennus</emphasis>
spp.
</taxonomicName>
; Jäger 2000, 2014
<emphasis box="[521,535,725,751]" italics="true" pageId="4" pageNumber="369">b</emphasis>
). Legs: length in most cases 2143 (n=5), also 2431 (n=3), 3421/4231 (n=1). Legs laterigrade (
<figureCitation box="[614,700,757,783]" captionStart="Figs 34–45" captionStartId="10.[144,183,1494,1516]" captionTargetBox="[157,1089,223,1457]" captionTargetId="figure-20@10.[157,1089,223,1457]" captionTargetPageId="10" captionText="Figs 34–45. May bruno gen. n. sp. n. from Witsand, South Africa (34–36, 41–43) holotype ♂; (37–40, 44–45) ♀ paratype, PJ 3524. (34–39, 43) habitus of preserved spiders: (34, 37) dorsal; (35, 38) ventral; (36, 39, 43) frontal; (40) palp of ♀, lateral, showing rows of elongated setae; (41–42) palp of ♂: (41) ventral, (42) retrolateral; (44–45) tarsal tips, showing light-coloured indented claw tuft setae (white arrow), additional dark claw tufts with Lawrence setae (black arrow), and scopula hairs (grey arrow): (44) ventral, (45) lateral." figureDoi="http://doi.org/10.5281/zenodo.7914109" httpUri="https://zenodo.org/record/7914109/files/figure.png" pageId="4" pageNumber="369">Figs 43</figureCitation>
,
<figureCitation box="[711,741,757,783]" captionStart="Figs 46–49" captionStartId="11.[180,219,802,824]" captionTargetBox="[193,1126,204,787]" captionTargetId="figure-282@11.[193,1126,204,787]" captionTargetPageId="11" captionText="Figs 46–49. May bruno gen. n. sp. n. from Witsand, South Africa: (46) habitat; (47) burrow with opened lid; (48–49) habitus of live spider. (Figs 46–47, 49) by Jon Leroy; (48) by Dirk Kunz." figureDoi="http://doi.org/10.5281/zenodo.7914111" httpUri="https://zenodo.org/record/7914111/files/figure.png" pageId="4" pageNumber="369">48</figureCitation>
,
<figureCitation box="[752,783,757,783]" captionStart="Figs 94–99" captionStartId="19.[182,221,1582,1604]" captionTargetBox="[193,1130,877,1566]" captionTargetId="figure-73@19.[193,1130,877,1566]" captionTargetPageId="19" captionText="Figs 94–99. May norm gen. n. sp. n., paratypes from Messum Crater, Namibia (94–96) PJ 1852; (97–99) PJ 3543: habitus of preserved specimens (94, 97) dorsal; (95, 98) ventral; (96, 99) frontal." figureDoi="http://doi.org/10.5281/zenodo.7914131" httpUri="https://zenodo.org/record/7914131/files/figure.png" pageId="4" pageNumber="369">96</figureCitation>
), spination pattern following general pattern of
<taxonomicName authorityName="Bertkau" authorityYear="1872" box="[345,477,789,815]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
with the following exceptions: Spination (only common patterns for all four species are listed): palp: 131; legs: femur I–III 323, IV 322; patella: variable, as in other
<taxonomicName authorityName="Bertkau" authorityYear="1872" box="[360,493,853,879]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
, posterior patellae generally with more spines; tibia I–IV 22(3)26; metatarsus I–III 3034, IV 3037. Scopulae on tarsi and metatarsi moderately dense to sparse. Palpal claw in females present, elongated, with 8 to 12 long teeth (
<figureCitation captionStart="Figs 20–33" captionStartId="9.[183,222,1469,1491]" captionTargetBox="[182,1141,242,1434]" captionTargetId="figure-15@9.[181,1141,242,1434]" captionTargetPageId="9" captionText="Figs 20–33.May bruno gen.n. sp. n. from SouthAfrica (♀ paratypes from Witsand,♀ from Twee Rivieren): (20–22) tarsus tip: (20) ventral, (21) lateral, (22) dorsal); (23) ♀ palpal claw, lateral; (24–25) Lawrence setae; (26–29) prosoma of ♀♀, showing positions of dorsal marks (white arrows in Fig. 26); (30) metatarsal stopper, dorsal; (31) eye arrangement, dorsal; (32) cheliceral dentition, ventral; (33) gnathocoxae, ventro-distal, showing reduced serrula (black arrow). Abbreviations:AP – plate of Lawrence setae; DP – dorsal plate of elongated setae; IP – plate of indented claw tuft setae; TS – transverse suture with slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914105" httpUri="https://zenodo.org/record/7914105/files/figure.png" pageId="4" pageNumber="369">Figs 23</figureCitation>
,
<figureCitation box="[186,218,949,975]" captionStart="Figs 87–93" captionStartId="19.[181,220,720,742]" captionTargetBox="[181,1122,236,706]" captionTargetId="figure-15@19.[181,1141,236,725]" captionTargetPageId="19" captionText="Figs 87–93. May norm gen. n. sp. n. from Messum Crater, Namibia (87–89, 91–93) ♀ paratype [PJ 1852]; (90) holotype ♂ [PJ 1851]: (87) eye arrangement, dorsal; (88) cheliceral dentition, ventral; (89) metatarsal stopper, dorsal; (90) ♀ palpal claw, lateral; (91) Lawrence setae; (92) Claw tuft setae, showing indented tip; (93) leg I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.7914129" httpUri="https://zenodo.org/record/7914129/files/figure.png" pageId="4" pageNumber="369">90</figureCitation>
). Metatarsal stopper (
<emphasis box="[462,524,949,974]" italics="true" pageId="4" pageNumber="369">sensu</emphasis>
<bibRefCitation author="RAMIREZ, M. J." box="[532,692,949,977]" pageId="4" pageNumber="369" pagination="1 - 374" refId="ref12833" refString="RAMIREZ, M. J. 2014. The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History 390: 1 - 374." type="journal article" year="2014">RamÍrez 2014</bibRefCitation>
) not developed as the usual trilobate membrane, known to be apomorphic for
<taxonomicName authorityName="Bertkau" authorityYear="1872" box="[595,730,981,1007]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
, but with median hook and lateral projections only weakly (
<figureCitation box="[428,509,1013,1039]" captionStart="Figs 20–33" captionStartId="9.[183,222,1469,1491]" captionTargetBox="[182,1141,242,1434]" captionTargetId="figure-15@9.[181,1141,242,1434]" captionTargetPageId="9" captionText="Figs 20–33.May bruno gen.n. sp. n. from SouthAfrica (♀ paratypes from Witsand,♀ from Twee Rivieren): (20–22) tarsus tip: (20) ventral, (21) lateral, (22) dorsal); (23) ♀ palpal claw, lateral; (24–25) Lawrence setae; (26–29) prosoma of ♀♀, showing positions of dorsal marks (white arrows in Fig. 26); (30) metatarsal stopper, dorsal; (31) eye arrangement, dorsal; (32) cheliceral dentition, ventral; (33) gnathocoxae, ventro-distal, showing reduced serrula (black arrow). Abbreviations:AP – plate of Lawrence setae; DP – dorsal plate of elongated setae; IP – plate of indented claw tuft setae; TS – transverse suture with slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914105" httpUri="https://zenodo.org/record/7914105/files/figure.png" pageId="4" pageNumber="369">Fig. 30</figureCitation>
) to well developed (
<figureCitation box="[739,825,1013,1039]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="4" pageNumber="369">Figs 69</figureCitation>
,
<figureCitation box="[837,869,1013,1039]" captionStart="Figs 87–93" captionStartId="19.[181,220,720,742]" captionTargetBox="[181,1122,236,706]" captionTargetId="figure-15@19.[181,1141,236,725]" captionTargetPageId="19" captionText="Figs 87–93. May norm gen. n. sp. n. from Messum Crater, Namibia (87–89, 91–93) ♀ paratype [PJ 1852]; (90) holotype ♂ [PJ 1851]: (87) eye arrangement, dorsal; (88) cheliceral dentition, ventral; (89) metatarsal stopper, dorsal; (90) ♀ palpal claw, lateral; (91) Lawrence setae; (92) Claw tuft setae, showing indented tip; (93) leg I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.7914129" httpUri="https://zenodo.org/record/7914129/files/figure.png" pageId="4" pageNumber="369">89</figureCitation>
).
</paragraph>
<paragraph blockId="4.[144,1105,204,1677]" pageId="4" pageNumber="369">
Coloration (in ethanol;
<figureCitation box="[402,532,1046,1072]" captionStart="Figs 34–45" captionStartId="10.[144,183,1494,1516]" captionTargetBox="[157,1089,223,1457]" captionTargetId="figure-20@10.[157,1089,223,1457]" captionTargetPageId="10" captionText="Figs 34–45. May bruno gen. n. sp. n. from Witsand, South Africa (34–36, 41–43) holotype ♂; (37–40, 44–45) ♀ paratype, PJ 3524. (34–39, 43) habitus of preserved spiders: (34, 37) dorsal; (35, 38) ventral; (36, 39, 43) frontal; (40) palp of ♀, lateral, showing rows of elongated setae; (41–42) palp of ♂: (41) ventral, (42) retrolateral; (44–45) tarsal tips, showing light-coloured indented claw tuft setae (white arrow), additional dark claw tufts with Lawrence setae (black arrow), and scopula hairs (grey arrow): (44) ventral, (45) lateral." figureDoi="http://doi.org/10.5281/zenodo.7914109" httpUri="https://zenodo.org/record/7914109/files/figure.png" pageId="4" pageNumber="369">Figs 34–39</figureCitation>
,
<figureCitation box="[542,618,1045,1072]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="4" pageNumber="369">55–57</figureCitation>
,
<figureCitation box="[628,704,1046,1072]" captionStart="Figs 70–77" captionStartId="16.[144,183,1443,1465]" captionTargetBox="[155,1091,208,1422]" captionTargetId="figure-38@16.[155,1091,204,1422]" captionTargetPageId="16" captionText="Figs 70–77.May rudy gen.n. sp. n., holotype ♂ from Copper Valley,Namibia:(70–72) habitus of preserved spider: (70) dorsal, (71) ventral, (72) frontal; (73–75) ♂ palp: (73) ventral, (74) retrolateral, (75) RTA, dorsal); (76–77) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and claw tuft penetrating dorsal setae (white arrow): (76) lateral, (77) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914123" httpUri="https://zenodo.org/record/7914123/files/figure.png" pageId="4" pageNumber="369">70–72</figureCitation>
,
<figureCitation box="[715,793,1046,1072]" captionStart="Figs 94–99" captionStartId="19.[182,221,1582,1604]" captionTargetBox="[193,1130,877,1566]" captionTargetId="figure-73@19.[193,1130,877,1566]" captionTargetPageId="19" captionText="Figs 94–99. May norm gen. n. sp. n., paratypes from Messum Crater, Namibia (94–96) PJ 1852; (97–99) PJ 3543: habitus of preserved specimens (94, 97) dorsal; (95, 98) ventral; (96, 99) frontal." figureDoi="http://doi.org/10.5281/zenodo.7914131" httpUri="https://zenodo.org/record/7914131/files/figure.png" pageId="4" pageNumber="369">94–99</figureCitation>
): Pale brown to pale yellow, without distinct pattern; conspicuous white setae on prosoma, especially at clypeus and along margin, dark setae in front of and on spinnerets. Coloration (of live specimens;
<figureCitation box="[144,275,1142,1168]" captionStart="Figs 46–49" captionStartId="11.[180,219,802,824]" captionTargetBox="[193,1126,204,787]" captionTargetId="figure-282@11.[193,1126,204,787]" captionTargetPageId="11" captionText="Figs 46–49. May bruno gen. n. sp. n. from Witsand, South Africa: (46) habitat; (47) burrow with opened lid; (48–49) habitus of live spider. (Figs 46–47, 49) by Jon Leroy; (48) by Dirk Kunz." figureDoi="http://doi.org/10.5281/zenodo.7914111" httpUri="https://zenodo.org/record/7914111/files/figure.png" pageId="4" pageNumber="369">Figs 48–49</figureCitation>
, only known from one species): setae white to shimmering pink.
</paragraph>
<paragraph blockId="4.[144,1105,204,1677]" pageId="4" pageNumber="369">
Male palp (
<figureCitation box="[272,375,1175,1201]" captionStart="Figs 1–10" captionStartId="6.[144,183,1212,1234]" captionTargetBox="[145,1104,210,1196]" captionTargetId="figure-177@6.[144,1104,204,1196]" captionTargetPageId="6" captionText="Figs 1–10. May bruno gen. n. sp. n. from Witsand, South Africa, holotype ♂: (1–4) left ♂ palp (1) ventral, (2) retrolateral, (3) distal tegulum, prolateral, (4) RTA, dorsal; (5–6) eye arrangement (5) frontal, (6) dorsal); (7) leg I, prolateral; (8) leg claw; (9) gnathocoxae and labium, ventral; (10) cheliceral dentition, ventral. Abbreviations: C – conductor; E – embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914099" httpUri="https://zenodo.org/record/7914099/files/figure.png" pageId="4" pageNumber="369">Figs 1–4</figureCitation>
,
<figureCitation box="[388,463,1175,1201]" captionStart="Figs 34–45" captionStartId="10.[144,183,1494,1516]" captionTargetBox="[157,1089,223,1457]" captionTargetId="figure-20@10.[157,1089,223,1457]" captionTargetPageId="10" captionText="Figs 34–45. May bruno gen. n. sp. n. from Witsand, South Africa (34–36, 41–43) holotype ♂; (37–40, 44–45) ♀ paratype, PJ 3524. (34–39, 43) habitus of preserved spiders: (34, 37) dorsal; (35, 38) ventral; (36, 39, 43) frontal; (40) palp of ♀, lateral, showing rows of elongated setae; (41–42) palp of ♂: (41) ventral, (42) retrolateral; (44–45) tarsal tips, showing light-coloured indented claw tuft setae (white arrow), additional dark claw tufts with Lawrence setae (black arrow), and scopula hairs (grey arrow): (44) ventral, (45) lateral." figureDoi="http://doi.org/10.5281/zenodo.7914109" httpUri="https://zenodo.org/record/7914109/files/figure.png" pageId="4" pageNumber="369">41–42</figureCitation>
,
<figureCitation box="[475,551,1175,1202]" captionStart="Figs 50–54" captionStartId="12.[144,183,942,964]" captionTargetBox="[144,1104,209,927]" captionTargetId="figure-266@12.[144,1104,204,927]" captionTargetPageId="12" captionText="Figs 50–54. May ansie gen. n. sp. n., holotype ♂ from Kokerboom Forest, Namibia: (50–52) left ♂ palp: (50) ventral, (51) retrolateral, (52) detail of embolus, ventral; (53) chelicerae, ventral; (54) eye arrangement, dorsal. Abbreviations: C – conductor; DE – dorsal extension of proximal embolus; RE – round extension of distal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914115" httpUri="https://zenodo.org/record/7914115/files/figure.png" pageId="4" pageNumber="369">50–52</figureCitation>
,
<figureCitation box="[563,638,1175,1202]" captionStart="Figs 55–63" captionStartId="13.[183,222,1296,1318]" captionTargetBox="[193,1129,204,1277]" captionTargetId="figure-165@13.[193,1129,201,1277]" captionTargetPageId="13" captionText="Figs 55–63.May ansie gen.n. sp. n. from Kokerboom Forest,Namibia (55–57, 61–62) ♂ paratype;(58–60, 63) holotype ♂: (55–57, 60) habitus of preserved spider: (55) dorsal, (56) ventral, (57) frontal, (60) fronto-lateral [left chelicerae removed]; (58–59) ♂ palp: (58) ventral, (59) RTA, dorsal; (61–62) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and dorsal setae (white arrow) penetrating theclaw tuft:(61) lateral, (62) ventral; (63) gnathocoxae, frontal,showing absence of serrula." figureDoi="http://doi.org/10.5281/zenodo.7914117" httpUri="https://zenodo.org/record/7914117/files/figure.png" pageId="4" pageNumber="369">58–59</figureCitation>
,
<figureCitation box="[650,727,1175,1201]" captionStart="Figs 64–69" captionStartId="15.[183,222,1070,1092]" captionTargetBox="[182,1141,245,1049]" captionTargetId="figure-183@15.[181,1141,237,1049]" captionTargetPageId="15" captionText="Figs 64–69.May rudy gen. n. sp. n., holotype ♂ from Copper Valley, Namibia: (64–67) left ♂ palp: (64) ventral, (65) retrolateral, (66) tegulum, ventral, (67) RTA, dorsal; (68) eye arrangement, dorsal; (69) metatarsal stopper, dorsal.Abbreviations:C – conductor;DE – dorsal extension of proximal embolus; S – “shoulder” of basal cymbium." figureDoi="http://doi.org/10.5281/zenodo.7914119" httpUri="https://zenodo.org/record/7914119/files/figure.png" pageId="4" pageNumber="369">64–67</figureCitation>
,
<figureCitation box="[739,817,1175,1202]" captionStart="Figs 70–77" captionStartId="16.[144,183,1443,1465]" captionTargetBox="[155,1091,208,1422]" captionTargetId="figure-38@16.[155,1091,204,1422]" captionTargetPageId="16" captionText="Figs 70–77.May rudy gen.n. sp. n., holotype ♂ from Copper Valley,Namibia:(70–72) habitus of preserved spider: (70) dorsal, (71) ventral, (72) frontal; (73–75) ♂ palp: (73) ventral, (74) retrolateral, (75) RTA, dorsal); (76–77) tarsal tips, showing additional claw tufts with Lawrence setae (black arrows) and claw tuft penetrating dorsal setae (white arrow): (76) lateral, (77) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914123" httpUri="https://zenodo.org/record/7914123/files/figure.png" pageId="4" pageNumber="369">73–75</figureCitation>
): Cymbium with variable shapes. Tegulum extending over the largest part of alveolus, with spermophor running along prolateral margin and S-shaped on retrolateral side; embolus and membranous conductor arising in distal half from tegulum. Embolus relatively short, situated in distal part of alveolus, arising from retrolateral to distal part of tegulum, running prolaterally. Female copulatory organ (
<figureCitation box="[444,576,1336,1362]" captionStart-0="Figs 11–17" captionStart-1="Figs 18–19" captionStartId-0="7.[181,220,1399,1421]" captionStartId-1="8.[144,183,729,751]" captionTargetBox-0="[192,1121,225,1381]" captionTargetBox-1="[185,1059,204,712]" captionTargetId-0="figure-62@7.[192,1121,225,1381]" captionTargetId-1="figure-405@8.[185,1063,204,712]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figs 11–17. May bruno gen. n. sp. n. from South Africa, ♀ paratypes from Witsand, South Africa (11–13) PJ 3524, (14–16) PJ 3525, (17) PJ 3526): (11, 14, 17) epigyne, ventral; (12, 15) vulva, dorsal; (13) vulva, posterior; (16) schematic course of internal duct system, dorsal. Abbreviations: FB – fusion bubbles; FD – fertilisation ducts; G – glands; GP – guiding pockets; SS – slit sensilla." captionText-1="Figs 18–19. May bruno gen. n. sp. n., ♀ from Twee Riviere, South Africa (PJ 3536): (18) epigyne, ventral; (19) vulva, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.7914101" figureDoi-1="http://doi.org/10.5281/zenodo.7914103" httpUri-0="https://zenodo.org/record/7914101/files/figure.png" httpUri-1="https://zenodo.org/record/7914103/files/figure.png" pageId="4" pageNumber="369">Figs 11–19</figureCitation>
,
<figureCitation box="[589,668,1336,1363]" captionStart="Figs 78–86" captionStartId="18.[144,183,1534,1556]" captionTargetBox="[144,1092,204,1516]" captionTargetId="figure-21@18.[144,1092,204,1517]" captionTargetPageId="18" captionText="Figs 78–86. May norm gen.n.sp. n.from Messum Crater, Namibia (78–80) holotype ♂ [PJ 1851]; (81–85) ♀ paratype [PJ 3534]; 86 ♀ paratype [PJ 1852]): (78, 81) epigyne, ventral (with mating plugs); (79, 82) vulva, dorsal; (84) vulva,anterior; (80, 83) schematic course of internal duct system, dorsal; (85) schematic course of internal duct system, anterior; (86) sternum, ventral.Abbreviations:FB – fusion bubbles; FD – fertilisationducts; GE – glandular extensions;GP – guiding pockets; SS – slit sensilla." figureDoi="http://doi.org/10.5281/zenodo.7914127" httpUri="https://zenodo.org/record/7914127/files/figure.png" pageId="4" pageNumber="369">78–85</figureCitation>
): Characterised by fused lateral lobes, i.e. epigynal ledges not reaching posteriorly epigastric furrow. Epigyne with atrium and guiding pockets leading to copulatory ducts. Internal duct system consisting of spherical spermathecae, short copulatory ducts and fertilisation ducts. Epigynal field slightly longer than wide, anteriorly converging, with or without one pair of slit sensilla. Distribution:
<collectingCountry box="[291,433,1505,1531]" name="South Africa" pageId="4" pageNumber="369">South Africa</collectingCountry>
,
<collectingCountry box="[442,538,1505,1531]" name="Namibia" pageId="4" pageNumber="369">Namibia</collectingCountry>
(
<figureCitation box="[552,648,1505,1531]" captionStart="Fig" captionStartId="22.[144,177,1606,1628]" captionTargetBox="[214,1033,1039,1588]" captionTargetId="figure-393@22.[214,1033,1039,1588]" captionTargetPageId="22" captionText="Fig. 120. Distribution records of May gen. n. in Namibia and South Africa: M. bruno sp. n. (green squares; South Africa), M. ansie sp. n. (red circles; Namibia), M. rudy sp. n. (pink diamond; Namibia), M. norm sp. n. (blue triangles; Namibia)." figureDoi="http://doi.org/10.5281/zenodo.7914139" httpUri="https://zenodo.org/record/7914139/files/figure.png" pageId="4" pageNumber="369">Fig. 120</figureCitation>
). Occurring most likely also in
<collectingCountry box="[987,1100,1505,1531]" name="Botswana" pageId="4" pageNumber="369">Botswana</collectingCountry>
. Species included:
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[347,452,1546,1572]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="bruno" status="sp. n.">
<emphasis box="[347,452,1546,1572]" italics="true" pageId="4" pageNumber="369">M. bruno</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[459,520,1546,1572]" pageId="4" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[532,629,1547,1572]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="ansie" status="sp. n.">
<emphasis box="[532,629,1547,1572]" italics="true" pageId="4" pageNumber="369">M. ansie</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[636,697,1546,1572]" pageId="4" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[710,798,1546,1572]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="rudy" status="sp. n.">
<emphasis box="[710,798,1546,1572]" italics="true" pageId="4" pageNumber="369">M. rudy</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[805,867,1546,1572]" pageId="4" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authority="Jäger &amp; Krehenwinkel, 2015" authorityName="Jäger &amp; Krehenwinkel" authorityYear="2015" box="[879,976,1547,1572]" class="Arachnida" family="Sparassidae" genus="May" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="norm" status="sp. n.">
<emphasis box="[879,976,1547,1572]" italics="true" pageId="4" pageNumber="369">M. norm</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[983,1043,1546,1572]" pageId="4" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="4.[144,1105,204,1677]" pageId="4" pageNumber="369">
Biology:
<collectingRegion box="[249,291,1587,1613]" country="Cameroon" name="North" pageId="4" pageNumber="369">Not</collectingRegion>
many observations could be made on live specimens so far. Spiders live in arid environments. One spider was observed in a burrow in a sand dune, while others were active during the night, running on the ground on gravel.
</paragraph>
</subSubSection>
</treatment>
</document>
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