137 lines
13 KiB
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137 lines
13 KiB
XML
<document ID-DOI="10.1111/zoj.12193" ID-ISSN="0024-4082" ID-Zenodo-Dep="5330460" approvalRequired="150" approvalRequired_for_taxonomicNames="66" approvalRequired_for_textStreams="58" approvalRequired_for_treatments="26" checkinTime="1630167834076" checkinUser="felipe" docAuthor="Norman, David B." docDate="2015" docId="03F9879B320FFFCCFF06FD85FB377D9F" docLanguage="en" docName="zoj12193.pdf" docOrigin="Zoological Journal of the Linnean Society 173 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12193" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.1:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="1" docTitle="Probactrosaurus" docType="treatment" docVersion="5" lastPageNumber="158" masterDocId="FFC0FFE3324DFF8EFFBDFFADFFCD7D65" masterDocTitle="On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna)" masterLastPageNumber="189" masterPageNumber="92" pageNumber="158" updateTime="1630625843961" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna)</mods:title>
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</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Norman, David B.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-01-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>173</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>92</mods:start>
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<mods:end>189</mods:end>
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<mods:location>
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<mods:url>https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12193</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/zoj.12193</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5330460</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5398357" ID-Zenodo-Dep="5398357" LSID="urn:lsid:plazi:treatment:03F9879B320FFFCCFF06FD85FB377D9F" httpUri="http://treatment.plazi.org/id/03F9879B320FFFCCFF06FD85FB377D9F" lastPageNumber="158" pageId="66" pageNumber="158">
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<subSubSection pageId="66" pageNumber="158" type="nomenclature">
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<paragraph blockId="66.[187,755,551,605]" pageId="66" pageNumber="158">
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<heading centered="true" fontSize="8" level="2" pageId="66" pageNumber="158" reason="8">
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<emphasis box="[187,537,552,573]" italics="true" pageId="66" pageNumber="158">
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<taxonomicName baseAuthorityName="Norman" baseAuthorityYear="2002" box="[187,409,552,573]" class="Reptilia" family="Iguanodontidae" genus="Probactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="genus">PROBACTROSAURUS</taxonomicName>
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GOBIENSIS
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</emphasis>
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<bibRefCitation author="Rozhdestvensky AK" pageId="66" pageNumber="158" pagination="1243 - 1246" refId="ref64363" refString="Rozhdestvensky AK. 1952. Otkritiye iguanodonta v Mongolii. [Discovery of iguanodonts in Mongolia]. Doklady Akademiya Nauk CCCP LXXXIV: 1243 - 1246." type="book chapter" year="1952">ROZHDESTVENSKY, 1952</bibRefCitation>
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(NORMAN, 2002)
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</heading>
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</paragraph>
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<paragraph blockId="66.[164,779,621,735]" pageId="66" pageNumber="158">
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Remains of this taxon indicate the presence of a medium-sized iguanodontian (attaining
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<emphasis box="[609,622,653,674]" italics="true" pageId="66" pageNumber="158">∼</emphasis>
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<quantity box="[622,663,652,674]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="66" pageNumber="158" unit="m" value="6.0">6 m</quantity>
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in length) collected from the Ulansuhai Formation (Turonian), Maortu,
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<collectingCountry box="[265,334,713,735]" name="China" pageId="66" pageNumber="158">China</collectingCountry>
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(
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<bibRefCitation author="Kobayashi Y & Lu J-C" box="[352,619,713,735]" pageId="66" pageNumber="158" pagination="235 - 259" refId="ref61900" refString="Kobayashi Y, Lu J-C. 2003. A new ornithomimid dinosaurian with gregarious habits from the Late Cretaceous of China. Acta Palaeontologica Polonica 48: 235 - 259." type="journal article" year="2003">Kobayashi & Lu, 2003</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="66" pageNumber="158" type="description">
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<paragraph blockId="66.[164,779,776,1318]" box="[164,337,776,798]" pageId="66" pageNumber="158">
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<emphasis box="[164,337,776,798]" italics="true" pageId="66" pageNumber="158">Teeth and jaws</emphasis>
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</paragraph>
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<paragraph blockId="66.[164,779,776,1318]" pageId="66" pageNumber="158">
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The dentary crowns of this taxon (Norman, 2002: figs 14–16) are narrower and less ornate than those described in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[304,412,868,889]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[304,412,868,889]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[428,505,868,890]" captionStart="Figure 37" captionStartId="42.[164,243,855,874]" captionTargetBox="[323,1284,195,825]" captionTargetId="figure-487@42.[323,1284,195,825]" captionTargetPageId="42" captionText="Figure 37. Hypselospinus cf. fittoni. NHMUK R1831. Teeth. A, dentary tooth in lingual view (isolated specimen on stub of matrix); B, maxillary tooth in labial view. C−E, dentary replacement crowns embedded in the alveolar bone of the dentary. Abbreviations: m, mammillae on the marginal denticles; p, primary ridge; r, minor ridges; st, strand-like ridges." figureDoi="http://doi.org/10.5281/zenodo.5369678" httpUri="https://zenodo.org/record/5369678/files/figure.png" pageId="66" pageNumber="158">Fig. 37</figureCitation>
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). The marginal denticles support mammillations, but these are less numerous than in the case of
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[384,496,929,951]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[384,496,929,951]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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and the denticles do not form a curved shelf as they do in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[546,657,960,981]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[546,657,960,981]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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. The roots of the teeth are also fluted for the compaction of adjacent functional and replacement crowns that form the dental magazine. There are at least two replacement crowns in each alveolus in the deeper portions of the dentary and the occlusal surface is broad because it comprises at least two dentary crowns, unlike
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[164,277,1174,1195]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[164,277,1174,1195]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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. The dentary of
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<emphasis box="[468,638,1174,1195]" italics="true" pageId="66" pageNumber="158">Prob. gobiensis</emphasis>
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is comparatively shorter and deeper than that of
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[607,720,1204,1226]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[607,720,1204,1226]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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with a larger and deeper area devoted to the dental magazine as well as a tall, perpendicular coronoid process (Norman, 2002: fig. 12).
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</paragraph>
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<paragraph blockId="66.[164,779,1359,1595]" box="[164,323,1359,1380]" pageId="66" pageNumber="158">
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<emphasis box="[164,323,1359,1380]" italics="true" pageId="66" pageNumber="158">Axial skeleton</emphasis>
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</paragraph>
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<paragraph blockId="66.[164,779,1359,1595]" pageId="66" pageNumber="158">
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What is known of the dorsal vertebral series (Norman, 2002: fig. 17) shows neither the thickening of the articular margins of the centra, nor any clear indication of the narrow and very tall neural spines that are displayed in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[322,439,1512,1534]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[322,439,1512,1534]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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(this is confirmed by reference to the shape of the anterior caudals – Norman, 2002: fig. 18).
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</paragraph>
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<paragraph blockId="66.[164,779,1636,1903]" box="[164,417,1636,1657]" pageId="66" pageNumber="158">
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<emphasis box="[164,417,1636,1657]" italics="true" pageId="66" pageNumber="158">Appendicular skeleton</emphasis>
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</paragraph>
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<paragraph blockId="66.[164,779,1636,1903]" lastBlockId="66.[826,1442,198,250]" pageId="66" pageNumber="158">
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Most of the postcranial anatomy of
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<emphasis box="[604,779,1666,1687]" italics="true" pageId="66" pageNumber="158">Prob. gobiensis</emphasis>
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(Norman, 2002: figs 20–33) appears to be gracile and generally comparable to that seen in
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<taxonomicName authority="(Norman, 1986)" baseAuthorityName="Norman" baseAuthorityYear="1986" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="atherfieldensis">
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<emphasis box="[582,779,1728,1749]" italics="true" pageId="66" pageNumber="158">M. atherfieldensis</emphasis>
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(
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<bibRefCitation author="Norman DB" box="[172,338,1758,1780]" pageId="66" pageNumber="158" pagination="281 - 372" refId="ref62903" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
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)
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</taxonomicName>
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rather than
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[504,617,1758,1780]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[504,617,1758,1780]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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. The forearm and manus are notably slender and lightly built in
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<emphasis box="[164,329,1819,1840]" italics="true" pageId="66" pageNumber="158">Prob. gobiensis</emphasis>
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, and the pollex ungual is small, narrow, and conical (Norman, 2002: figs 22–26), in marked contrast to these structures in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[470,579,1881,1902]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[470,579,1881,1902]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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. The ischial shaft is heavy, robust, and J-shaped (Norman, 2002: fig. 29) and resembles that seen in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[1154,1267,228,250]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="66" pageNumber="158" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[1154,1267,228,250]" italics="true" pageId="66" pageNumber="158">Hy. fittoni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |