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<document id="F5FD10F906CD8BA387B982148F97ECE1" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno &amp; Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF9AE8150F94886C22FEFB79" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia interior Liew &amp; Schilthuizen &amp; Vermeulen 2009, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="537" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="536" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B16A44BA633F79FEB24E49BBF92CADF7">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:namePart id="29B406C528AE3065685CB9822B297CD0">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="93D87F358C296D8D48D48EC66FA75CF3">Schilthuizen, Menno</mods:namePart>
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<mods:roleTerm id="F5EB1035EC215954FC65544D2ABE4282">Author</mods:roleTerm>
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<mods:namePart id="92C5C989209D54D3FDAF552FE4DD7653">Vermeulen, Jaap Jan</mods:namePart>
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<mods:title id="21EC069A5E595A59A4ACBE735312C07B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="523026F7897E0F5BC2053C2C76843D68">2009</mods:date>
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<mods:number id="9A414A79C4E2556AE27053D6598D9197">2009-10-26</mods:number>
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<mods:number id="1207B5F9F50E9C960BB56BD6699634D1">157</mods:number>
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<mods:number id="603F7F464DC40FE42EA96CA2FC887A7F">3</mods:number>
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<mods:url id="1AE63F2956EF0B7CDD26E8985ACE012C">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
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<mods:identifier id="856D8C5EA6643FD4D969D9621AC3C7CF" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
<mods:identifier id="53D096A2F6CC6EF8482A35901C593FAE" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="35842377598520E605EF89A1FE6660FE" type="Zenodo-Dep">5443023</mods:identifier>
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<treatment id="03F9878AFF9AE8150F94886C22FEFB79" ID-DOI="http://doi.org/10.5281/zenodo.5492193" ID-Zenodo-Dep="5492193" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79" httpUri="http://treatment.plazi.org/id/03F9878AFF9AE8150F94886C22FEFB79" lastPageId="22" lastPageNumber="537" pageId="21" pageNumber="536">
<subSubSection id="C34A6517FF9AE8160F94886C264FFA9E" pageId="21" pageNumber="536" type="nomenclature">
<paragraph id="8BEF369CFF9AE8160F94886C2641FB5E" blockId="21.[951,1316,1151,1172]" box="[951,1316,1151,1172]" pageId="21" pageNumber="536">
<heading id="D0A781F0FF9AE8160F94886C2641FB5E" box="[951,1316,1151,1172]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="536" reason="2">
<taxonomicName id="4C504D1FFF9AE8160F94886C27D0FB59" ID-CoL="6HLVL" authority="Liew &amp; Schilthuizen &amp; Vermeulen, 2009" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[951,1205,1151,1172]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="interior" status="sp. nov.">EVERETTIA INTERIOR</taxonomicName>
<emphasis id="B924EA8EFF9AE816089C88912641FB5E" bold="true" box="[1215,1316,1151,1172]" pageId="21" pageNumber="536">
<taxonomicNameLabel id="A21757F5FF9AE816089C88912641FB5E" box="[1215,1316,1151,1172]" pageId="21" pageNumber="536" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF369CFF9AE8160F1988B5264FFA9E" blockId="21.[826,1441,1189,1364]" pageId="21" pageNumber="536">
<emphasis id="B924EA8EFF9AE8160F1988B520E5FB71" box="[826,896,1190,1211]" italics="true" pageId="21" pageNumber="536">Types:</emphasis>
<materialsCitation id="3B383CC1FF9AE8160FA888B62643FA9E" collectingDate="2005-06-25" collectionCode="T" collectorName="S. Liew &amp; M. Schilthuizen &amp; S. Chiba. Date" country="Malaysia" county="State" latitude="4.7" location="Sapulut area" longLatPrecision="1306" longitude="116.6" municipality="Pensiangan District" pageId="21" pageNumber="536" specimenCode="BMNH 20080627" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="F347760CFF9AE8160FA888B62771FB70" box="[907,1044,1189,1210]" name="Malaysia" pageId="21" pageNumber="536">MALAYSIA</collectingCountry>
:
<collectingCounty id="628E4E10FF9AE816080188B6273AFB70" box="[1058,1119,1189,1210]" pageId="21" pageNumber="536">State</collectingCounty>
of
<collectingRegion id="4994F87EFF9AE81608AA88B627B6FB71" box="[1161,1235,1189,1211]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">Sabah</collectingRegion>
,
<collectingMunicipality id="6B8BACE6FF9AE81608C188B62008FB10" pageId="21" pageNumber="536">Pensiangan District</collectingMunicipality>
,
<location id="8E8F6047FF9AE8160FA488D72748FB13" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160FA488D72748FB13" box="[903,1069,1220,1242]" country="Malaysia" county="State" latitude="4.7" longLatPrecision="1306" longitude="116.6" municipality="Pensiangan District" name="Sapulut area" pageId="21" pageNumber="536" stateProvince="Sabah">Sapulut area</location>
.
<location id="8E8F6047FF9AE816086688D726C4FB13" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816086688D726C4FB13" box="[1093,1441,1220,1242]" country="Malaysia" county="State" latitude="4.7" longLatPrecision="1306" longitude="116.6" municipality="Pensiangan District" name="Limestone - Batu Sanaron" pageId="21" pageNumber="536" stateProvince="Sabah">Limestone Batu Sanaron</location>
(
<geoCoordinate id="EE64505BFF9AE8160F6188F020F4FB32" box="[834,913,1249,1272]" degrees="4" direction="north" minutes="42" orientation="latitude" pageId="21" pageNumber="536" precision="925" value="4.7">
4°42
<emphasis id="B924EA8EFF9AE8160F5688F2201EFB32" box="[885,891,1249,1272]" italics="true" pageId="21" pageNumber="536"></emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE64505BFF9AE8160F8F88F12776FB32" box="[940,1043,1249,1272]" degrees="116" direction="east" minutes="36" orientation="longitude" pageId="21" pageNumber="536" precision="925" value="116.6">
116°36
<emphasis id="B924EA8EFF9AE8160FDA88F2209AFB32" box="[1017,1023,1249,1272]" italics="true" pageId="21" pageNumber="536"></emphasis>
E
</geoCoordinate>
). Collected by T.-
<collectorName id="26A5534AFF9AE816092488F0260DFB32" box="[1287,1384,1250,1272]" pageId="21" pageNumber="536">S. Liew</collectorName>
,
<collectorName id="26A5534AFF9AE81609A088F020A9FADD" pageId="21" pageNumber="536">M. Schilthuizen</collectorName>
, and
<collectorName id="26A5534AFF9AE8160834891227ABFADC" box="[1047,1230,1281,1303]" pageId="21" pageNumber="536">S. Chiba. Date</collectorName>
:
<date id="FFEE105CFF9AE81608C389122632FADD" box="[1248,1367,1281,1303]" pageId="21" pageNumber="536" value="2005-06-25">
<collectingDate id="EFAAE9B4FF9AE81608C389122632FADD" box="[1248,1367,1281,1303]" pageId="21" pageNumber="536" value="2005-06-25">25.vi.2005</collectingDate>
</date>
,
<typeStatus id="54EB883EFF9AE816094A8912200BFAFF" pageId="21" pageNumber="536" type="holotype">holotype</typeStatus>
, BOR/MOL 4277,
<figureCitation id="136B2A19FF9AE8160866893327DCFAFF" box="[1093,1209,1312,1334]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="21" pageNumber="536">Figure 2C</figureCitation>
;
<specimenCount id="9D56FD15FF9AE81608EB8933260EFAFF" box="[1224,1387,1312,1333]" pageId="21" pageNumber="536" type="generic" typeStatus="paratypes">ten paratypes</specimenCount>
, five in
<specimenCode id="DBF69EE7FF9AE8160F78892C2746FA9E" box="[859,1059,1342,1364]" collectionCode="BMNH" pageId="21" pageNumber="536">BMNH 20080627</specimenCode>
and five in SP 13059
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6517FF9AE8150F19896622FEFB79" lastPageId="22" lastPageNumber="537" pageId="21" pageNumber="536" type="etymology">
<paragraph id="8BEF369CFF9AE8160F1989662677FA60" blockId="21.[826,1441,1397,1450]" pageId="21" pageNumber="536">
<emphasis id="B924EA8EFF9AE8160F19896620DDFA40" box="[826,952,1397,1418]" italics="true" pageId="21" pageNumber="536">Etymology:</emphasis>
This species is named
<taxonomicName id="4C504D1FFF9AE81608E98965267AFA41" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[1226,1311,1398,1419]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="interior">
<emphasis id="B924EA8EFF9AE81608E98965267AFA41" box="[1226,1311,1398,1419]" italics="true" pageId="21" pageNumber="536">interior</emphasis>
</taxonomicName>
for its distribution range in the interior of
<collectingRegion id="4994F87EFF9AE81608E08987266BFA60" box="[1219,1294,1428,1450]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">Sabah</collectingRegion>
.
</paragraph>
<paragraph id="8BEF369CFF9AE8160F1989D8275EF8A5" blockId="21.[826,1441,1483,1903]" pageId="21" pageNumber="536">
<emphasis id="B924EA8EFF9AE8160F1989D82745FA2A" box="[826,1056,1483,1504]" italics="true" pageId="21" pageNumber="536">Material examined:</emphasis>
<materialsCitation id="3B383CC1FF9AE816081389D82673F9CA" collectingDate="2000-07" elevation="450" location="Sanaron cave" municipality="Sapulut" pageId="21" pageNumber="536" specimenCount="1" stateProvince="Sabah">
<collectingRegion id="4994F87EFF9AE816081389D827ECFA2A" box="[1072,1161,1483,1504]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">SABAH</collectingRegion>
<collectingMunicipality id="6B8BACE6FF9AE816089489D82670FA2B" box="[1207,1301,1483,1505]" pageId="21" pageNumber="536">Sapulut</collectingMunicipality>
: BOR/MOL 921,
<location id="8E8F6047FF9AE8160F5B89F9277FFA35" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160F5B89F9277FFA35" box="[888,1050,1514,1535]" municipality="Sapulut" name="Sanaron cave" pageId="21" pageNumber="536" stateProvince="Sabah">Sanaron cave</location>
,
<quantity id="4CA89B79FF9AE816080F89FA2717FA35" box="[1068,1138,1513,1535]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">
<elevation id="007DD1AFFF9AE816080F89FA2717FA35" box="[1068,1138,1513,1535]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">450 m</elevation>
</quantity>
alt.,
<date id="FFEE105CFF9AE816089989F92673F9CA" box="[1210,1302,1514,1536]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE816089989F92673F9CA" box="[1210,1302,1514,1536]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE816090A89F927DBF9D4" collectingDate="2001-02" location="Batu Temurung" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 929,
<location id="8E8F6047FF9AE8160FA68A1A2735F9D4" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160FA68A1A2735F9D4" box="[901,1104,1545,1566]" name="Batu Temurung" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Temurung</location>
,
<date id="FFEE105CFF9AE816084C8A1B27DBF9D4" box="[1135,1214,1544,1566]" pageId="21" pageNumber="536" value="2001-02">
<collectingDate id="EFAAE9B4FF9AE816084C8A1B27DBF9D4" box="[1135,1214,1544,1566]" pageId="21" pageNumber="536" value="2001-02">ii.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608FD8A1B270DF9F7" collectingDate="2000-07" location="Sanaron cave" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 939,
<location id="8E8F6047FF9AE8160F198A34208EF9F6" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160F198A34208EF9F6" box="[826,1003,1575,1596]" name="Sanaron cave" pageId="21" pageNumber="536" stateProvince="SABAH">Sanaron cave</location>
,
<date id="FFEE105CFF9AE816082F8A34270DF9F7" box="[1036,1128,1575,1597]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE816082F8A34270DF9F7" box="[1036,1128,1575,1597]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608AA8A3427EBF991" collectingDate="2001-02" elevation="450" location="Batu Sinobang" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 969,
<location id="8E8F6047FF9AE816094A8A3420C2F991" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816094A8A3420C2F991" name="Batu Sinobang" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Sinobang</location>
,
<quantity id="4CA89B79FF9AE8160F958A562099F991" box="[950,1020,1605,1627]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">
<elevation id="007DD1AFFF9AE8160F958A562099F991" box="[950,1020,1605,1627]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">450 m</elevation>
</quantity>
alt.,
<date id="FFEE105CFF9AE816081C8A5627EBF991" box="[1087,1166,1605,1627]" pageId="21" pageNumber="536" value="2001-02">
<collectingDate id="EFAAE9B4FF9AE816081C8A5627EBF991" box="[1087,1166,1605,1627]" pageId="21" pageNumber="536" value="2001-02">ii.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608BD8A5527D0F9B0" collectingDate="2000-07" elevation="450" location="Batu Tinagas" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 1137,
<location id="8E8F6047FF9AE816094A8A5520F2F9B0" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816094A8A5520F2F9B0" name="Batu Tinagas" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Tinagas</location>
,
<quantity id="4CA89B79FF9AE8160F978A77209FF9B3" box="[948,1018,1636,1658]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">
<elevation id="007DD1AFFF9AE8160F978A77209FF9B3" box="[948,1018,1636,1658]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="536" unit="m" value="450.0">450 m</elevation>
</quantity>
alt.,
<date id="FFEE105CFF9AE816087A8A7727D0F9B0" box="[1113,1205,1636,1658]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE816087A8A7727D0F9B0" box="[1113,1205,1636,1658]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608F08A77273BF952" collectingDate="2005-06" location="Sanaron cave" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 4305,
<location id="8E8F6047FF9AE8160F198A90208FF952" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160F198A90208FF952" box="[826,1002,1667,1688]" name="Sanaron cave" pageId="21" pageNumber="536" stateProvince="SABAH">Sanaron cave</location>
,
<date id="FFEE105CFF9AE816082A8A91273BF952" box="[1033,1118,1666,1688]" pageId="21" pageNumber="536" value="2005-06">
<collectingDate id="EFAAE9B4FF9AE816082A8A91273BF952" box="[1033,1118,1666,1688]" pageId="21" pageNumber="536" value="2005-06">vi.2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE816085D8A902760F97D" collectingDate="2005-06" location="Batu Punggul" pageId="21" pageNumber="536" specimenCount="1" stateProvince="SABAH">
BOR/MOL 4332,
<location id="8E8F6047FF9AE816094A8A9020F9F97D" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816094A8A9020F9F97D" name="Batu Punggul" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Punggul</location>
,
<date id="FFEE105CFF9AE8160F938AB22760F97D" box="[944,1029,1697,1719]" pageId="21" pageNumber="536" value="2005-06">
<collectingDate id="EFAAE9B4FF9AE8160F938AB22760F97D" box="[944,1029,1697,1719]" pageId="21" pageNumber="536" value="2005-06">vi.2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE816083A8AB12614F97D" box="[1049,1393,1697,1719]" collectingDate="1986-09" collectionCode="JJ" location="Pun Batu" pageId="21" pageNumber="536" specimenCode="JJ 1304" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE816083A8AB12718F97C" box="[1049,1149,1697,1719]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 1304</specimenCode>
,
<location id="8E8F6047FF9AE81608B28AB1266DF97D" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE81608B28AB1266DF97D" box="[1169,1288,1698,1719]" name="Pun Batu" pageId="21" pageNumber="536" stateProvince="SABAH">Pun Batu</location>
,
<date id="FFEE105CFF9AE816093E8AB22614F97D" box="[1309,1393,1697,1719]" pageId="21" pageNumber="536" value="1986-09">
<collectingDate id="EFAAE9B4FF9AE816093E8AB22614F97D" box="[1309,1393,1697,1719]" pageId="21" pageNumber="536" value="1986-09">ix.1986</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81609A58AB1271FF91F" collectingDate="1987-05" collectionCode="JJ" location="Batu Punggul" pageId="21" pageNumber="536" specimenCode="JJ 1974" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81609A58AB12016F91F" collectionCode="JJ" pageId="21" pageNumber="536">JJ 1974</specimenCode>
,
<location id="8E8F6047FF9AE8160FA38AD32747F91C" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160FA38AD32747F91C" box="[896,1058,1728,1750]" name="Batu Punggul" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Punggul</location>
,
<date id="FFEE105CFF9AE816080C8AD3271FF91F" box="[1071,1146,1728,1749]" pageId="21" pageNumber="536" value="1987-05">
<collectingDate id="EFAAE9B4FF9AE816080C8AD3271FF91F" box="[1071,1146,1728,1749]" pageId="21" pageNumber="536" value="1987-05">v.1987</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608A58AD32016F93E" collectionCode="JJ" location="Batu Sinobang" pageId="21" pageNumber="536" specimenCode="JJ 5627" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81608A58AD32786F91C" box="[1158,1251,1728,1750]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 5627</specimenCode>
,
<location id="8E8F6047FF9AE81608D38AD326F8F91C" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE81608D38AD326F8F91C" box="[1264,1437,1728,1750]" name="Batu Sinobang" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Sinobang</location>
, 1997
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE8160F5C8ACC2792F93E" box="[895,1271,1758,1780]" collectingDate="2000-07" collectionCode="JJ" location="Batu Pungiton" pageId="21" pageNumber="536" specimenCode="JJ 7546" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE8160F5C8ACC20BFF93E" box="[895,986,1758,1780]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 7546</specimenCode>
,
<location id="8E8F6047FF9AE8160FC68ACC27EAF93E" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160FC68ACC27EAF93E" box="[997,1167,1758,1780]" name="Batu Pungiton" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Pungiton</location>
,
<date id="FFEE105CFF9AE81608B98ACD2792F93E" box="[1178,1271,1758,1780]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE81608B98ACD2792F93E" box="[1178,1271,1758,1780]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81609218ACC2765F8D9" collectingDate="2000-07" collectionCode="JJ" location="Batu Tinagas" pageId="21" pageNumber="536" specimenCode="JJ 7626" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81609218ACC2638F93E" box="[1282,1373,1758,1780]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 7626</specimenCode>
,
<location id="8E8F6047FF9AE816094A8ACC20F2F8D9" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816094A8ACC20F2F8D9" name="Batu Tinagas" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Tinagas</location>
,
<date id="FFEE105CFF9AE8160F808AEE2765F8D9" box="[931,1024,1789,1811]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE8160F808AEE2765F8D9" box="[931,1024,1789,1811]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE816082F8AEE261CF8D9" box="[1036,1401,1789,1811]" collectingDate="2000-07" collectionCode="JJ" location="Sanaron cave" pageId="21" pageNumber="536" specimenCode="JJ 7662" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE816082F8AEE270DF8D9" box="[1036,1128,1789,1811]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 7662</specimenCode>
,
<location id="8E8F6047FF9AE81608578AEE2674F8D8" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE81608578AEE2674F8D8" box="[1140,1297,1789,1810]" name="Sanaron cave" pageId="21" pageNumber="536" stateProvince="SABAH">Sanaron cave</location>
,
<date id="FFEE105CFF9AE816093E8AEE261CF8D9" box="[1309,1401,1789,1811]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE816093E8AEE261CF8D9" box="[1309,1401,1789,1811]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81609A58AEE2710F8F8" collectingDate="2000-07" collectionCode="JJ" location="Kg. Labang" pageId="21" pageNumber="536" specimenCode="JJ 7726" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81609A58AEE2016F8F8" collectionCode="JJ" pageId="21" pageNumber="536">JJ 7726</specimenCode>
,
<location id="8E8F6047FF9AE8160FA28B0F276FF8F8" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160FA28B0F276FF8F8" box="[897,1034,1820,1842]" name="Kg. Labang" pageId="21" pageNumber="536" stateProvince="SABAH">Kg. Labang</location>
,
<date id="FFEE105CFF9AE816083B8B0F2710F8F8" box="[1048,1141,1820,1842]" pageId="21" pageNumber="536" value="2000-07">
<collectingDate id="EFAAE9B4FF9AE816083B8B0F2710F8F8" box="[1048,1141,1820,1842]" pageId="21" pageNumber="536" value="2000-07">vii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81608A18B0F20ECF89A" collectingDate="2001-02" collectionCode="JJ" location="Batu Sinobang" pageId="21" pageNumber="536" specimenCode="JJ 7995" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81608A18B0F2785F8FB" box="[1154,1248,1819,1841]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 7995</specimenCode>
,
<location id="8E8F6047FF9AE81608CD8B0F26F8F8F8" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE81608CD8B0F26F8F8F8" box="[1262,1437,1820,1842]" name="Batu Sinobang" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Sinobang</location>
,
<date id="FFEE105CFF9AE8160F198B2920ECF89A" box="[826,905,1850,1872]" pageId="21" pageNumber="536" value="2001-02">
<collectingDate id="EFAAE9B4FF9AE8160F198B2920ECF89A" box="[826,905,1850,1872]" pageId="21" pageNumber="536" value="2001-02">ii.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE8160FB88B28264FF89A" box="[923,1322,1850,1872]" collectingDate="2001-02" collectionCode="JJ" location="Batu Temurung" pageId="21" pageNumber="536" specimenCode="JJ 8038" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE8160FB88B282099F885" box="[923,1020,1850,1872]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 8038</specimenCode>
,
<location id="8E8F6047FF9AE816082E8B2827AFF89A" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE816082E8B2827AFF89A" box="[1037,1226,1851,1872]" name="Batu Temurung" pageId="21" pageNumber="536" stateProvince="SABAH">Batu Temurung</location>
,
<date id="FFEE105CFF9AE81608F88B29264FF89A" box="[1243,1322,1850,1872]" pageId="21" pageNumber="536" value="2001-02">
<collectingDate id="EFAAE9B4FF9AE81608F88B29264FF89A" box="[1243,1322,1850,1872]" pageId="21" pageNumber="536" value="2001-02">ii.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CC1FF9AE81609188B282752F8A5" collectingDate="2001-02" collectionCode="JJ" location="Sanaron cave" pageId="21" pageNumber="536" specimenCode="JJ 8069" specimenCount="1" stateProvince="SABAH">
<specimenCode id="DBF69EE7FF9AE81609188B2826F9F89A" box="[1339,1436,1850,1872]" collectionCode="JJ" pageId="21" pageNumber="536">JJ 8069</specimenCode>
,
<location id="8E8F6047FF9AE8160F198B4A20BCF8A4" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8150F94886C22FEFB79:8E8F6047FF9AE8160F198B4A20BCF8A4" box="[826,985,1881,1902]" name="Sanaron cave" pageId="21" pageNumber="536" stateProvince="SABAH">Sanaron cave</location>
,
<date id="FFEE105CFF9AE8160FC48B4A2752F8A5" box="[999,1079,1881,1903]" pageId="21" pageNumber="536" value="2001-02">
<collectingDate id="EFAAE9B4FF9AE8160FC48B4A2752F8A5" box="[999,1079,1881,1903]" pageId="21" pageNumber="536" value="2001-02">ii.2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF369CFF99E8150CB38CD52178FF30" blockId="22.[144,758,198,250]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB38CD522A2FF11" box="[144,455,198,219]" italics="true" pageId="22" pageNumber="537">Diagnostic characteristics:</emphasis>
Shell suture depressed with widely placed spiral grooves.
</paragraph>
<paragraph id="8BEF369CFF99E8150CB38D372110FB9F" blockId="22.[144,759,291,1109]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB38D37227DFEF3" box="[144,280,292,313]" italics="true" pageId="22" pageNumber="537">Description:</emphasis>
Shell (
<figureCitation id="136B2A19FF99E8150D448D3022A4FEF3" box="[359,449,291,313]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="22" pageNumber="537">Figs 2C</figureCitation>
,
<figureCitation id="136B2A19FF99E8150DEE8D37228AFEF0" box="[461,495,292,314]" captionStart="Figure 3" captionStartId="8.[144,223,1547,1566]" captionTargetBox="[305,1263,196,1518]" captionTargetId="figure-68@8.[303,1263,196,1518]" captionTargetPageId="8" captionText="Figure 3. Shell surface sculptures (scanning electron micrograph). A, upper surface of Everettia interior sp. nov.; B, upper surface of Everettia jucunda; C, upper surface of Everettia layanglayang sp. nov.; D, protoconch of E. layanglayang sp. nov.; E, underside of E. layanglayang sp. nov.; F, upper surface of Everettia dominiki sp. nov.; G, upper surface of Everettia monticola sp. nov.; H, upper surface of Everettia planispira sp. nov. Scale bar = 200 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443029" httpUri="https://zenodo.org/record/5443029/files/figure.png" pageId="22" pageNumber="537">3A</figureCitation>
): medium-sized, rather thin, brownish, moderately elevated, outer whorls rounded. Periphery rounded, suture depressed. Above the periphery, shell shiny with widely placed spiral grooves,
<emphasis id="B924EA8EFF99E8150CD08D8C2260FE7E" box="[243,261,415,436]" italics="true" pageId="22" pageNumber="537">c.</emphasis>
45 per mm, numerous fine spiral striae in between grooves. Below the periphery, shell has fine, densely placed spiral grooves. Height up to
<quantity id="4CA89B79FF99E8150EAD8DC82197FE3B" box="[654,754,475,497]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="22" pageNumber="537" unit="mm" value="11.4">11.4 mm</quantity>
; width up to
<quantity id="4CA89B79FF99E8150D108DE922FDFDC5" box="[307,408,506,527]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="22" pageNumber="537" unit="mm" value="19.7">19.7 mm</quantity>
; diameter of the first three whorls 1.01.1, 0.91.0, and
<quantity id="4CA89B79FF99E8150DF68E0B2139FDE4" box="[469,604,536,558]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="22" pageNumber="537" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
, respectively; number of whorls up to five; height aperture up to
<quantity id="4CA89B79FF99E8150CB38E452382FDA1" box="[144,231,598,619]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="22" pageNumber="537" unit="mm" value="8.2">8.2 mm</quantity>
; width aperture up to
<quantity id="4CA89B79FF99E8150E318E45211DFDA1" box="[530,632,598,619]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="22" pageNumber="537" unit="mm" value="10.7">10.7 mm</quantity>
. Genitalia (
<figureCitation id="136B2A19FF99E8150CBB8E672395FD40" box="[152,240,628,650]" captionStart="Figure 7" captionStartId="13.[164,243,1755,1774]" captionTargetBox="[167,1442,198,1722]" captionTargetId="figure-0@13.[164,1444,194,1725]" captionTargetPageId="13" captionText="Figure 7. Genitalia. A, Everettia klemmantanica; B, Everettia lapidini sp. nov.; C, Everettia paulbasintali sp. nov.; D, Everettia subconsul; E, Everettia themis; F, Everettia layanglayang sp. nov.; G, Everettia jucundior sp. nov.; H, Everettia interior sp. nov.; I, Everettia corrugata corrugata." figureDoi="http://doi.org/10.5281/zenodo.5443037" httpUri="https://zenodo.org/record/5443037/files/figure.png" pageId="22" pageNumber="537">Fig. 7H</figureCitation>
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
<quantity id="4CA89B79FF99E8150CF88EA22249FD0D" box="[219,300,689,711]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="537" unit="mm" value="13.0">13 mm</quantity>
. The penis, dart-sac, and vagina with thick muscular walls. P, V, GO, BC, and DS arranged around the short genital atrium, limited space between the openings of each of the genital structures. The BC is short, about one-third of total DS length. ANIMAL (
<figureCitation id="136B2A19FF99E8150D7C8F5922DFFC95" box="[351,442,842,864]" captionStart="Figure 4" captionStartId="9.[164,243,1177,1196]" captionTargetBox="[167,1442,200,1146]" captionTargetId="figure-273@9.[164,1444,197,1148]" captionTargetPageId="9" captionText="Figure 4. Animal head and body pigmentation. A, Everettia klemmantanica; B, Everettia interior sp. nov.; C, Everettia jucunda; D, Everettia themis; E, Everettia layanglayang sp. nov.; F, Everettia subconsul; GI, possible hybrid of E. themis and E. subconsul." figureDoi="http://doi.org/10.5281/zenodo.5443031" httpUri="https://zenodo.org/record/5443031/files/figure.png" pageId="22" pageNumber="537">Figs 4B</figureCitation>
,
<figureCitation id="136B2A19FF99E8150DE58F59228DFCAA" box="[454,488,842,864]" captionStart="Figure 8" captionStartId="14.[144,223,1760,1779]" captionTargetBox="[147,1419,200,1728]" captionTargetId="figure-0@14.[144,1424,195,1730]" captionTargetPageId="14" captionText="Figure 8. Animal and mantle pigmentation. A, Everettia subconsul west coast lowland; B, Everettia layanglayang sp. nov.; C, Everettia monticola sp. nov.; D, Everettia safriei sp. nov.; E, Everettia interior sp. nov.; F, Everettia subconsul east coast lowland; G, Everettia corrugata corrugata; H, Everettia jucundior sp. nov.; I, Everettia klemmantanica; J, Everettia lapidini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443039" httpUri="https://zenodo.org/record/5443039/files/figure.png" pageId="22" pageNumber="537">8E</figureCitation>
): black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles. Black bands extend downward to just below the eye tentacles and backward from the umbilicus region to the end of the tail, downward to the edge of the foot. The mantle is black and covered by crowded tiny brighter spots and followed by regularly spaced radial black bands at the last half whorl.
</paragraph>
<paragraph id="8BEF369CFF99E8150CB3886D22FEFB79" blockId="22.[144,759,1150,1203]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB3886D22A2FB59" box="[144,455,1150,1171]" italics="true" pageId="22" pageNumber="537">Distribution and habitat:</emphasis>
Primary forest.
<collectingRegion id="4994F87EFF99E8150E8B886D2196FB5E" box="[680,755,1150,1172]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">Sabah</collectingRegion>
: Sapulut area (
<figureCitation id="136B2A19FF99E8150D1A888E22EBFB79" box="[313,398,1181,1203]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="22" pageNumber="537">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>