treatments-xml/data/03/F9/87/03F98785036CEF771C5779D2FEA36FAE.xml

<document id="9247F51D4F648041D9BAA2A6676059E4" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F98785036CEF771C5779D2FEA36FAE" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplodrassus qashqai Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="0EE16C9F-6CC6-4AF4-8343-750215C9FBD6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="90" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="87" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E99C594E2848D37EFA12A1D55B51F542" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CEBCA69F2DD1055904E92C302EDA905A">
<mods:title id="C264BDD306527536B0C101BEA2D56B58">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
</mods:titleInfo>
<mods:name id="A86CC2B09D8E72B4AD4B606351673E48" type="personal">
<mods:role id="A3E9F36B8EA32A53E4D51583B2912C9B">
<mods:roleTerm id="F39C63F415BA66AFBAAB106166D85C49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4CE2B516782137C6313657E2DD69CCD">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="69CE579735E73F829CD69DBB2B8F1E6B" type="LSID">A21C0B82-E2D9-4402-9408-D6160843DAF4</mods:nameIdentifier>
<mods:affiliation id="40A64B44FA82F32B67771E06C4E3165E">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
<mods:nameIdentifier id="5DD3FCF258BD917ECC866EC7687E5104" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9E00F0AB8AA9A19B75BA2C5D0F101F17" type="personal">
<mods:role id="D915A86154EFCEF994BB74631ABDBF7D">
<mods:roleTerm id="7028B74F6F75A70A94C3B583830B6351">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D70D9A35A9891B102988E21BE7B22E26">Chatzaki, Maria</mods:namePart>
<mods:nameIdentifier id="399DF4C851B3152F921027F03A2A20A7" type="LSID">2FBB48ED-84FE-4386-8F47-C958E9F8A2A1</mods:nameIdentifier>
<mods:nameIdentifier id="93F3DACA928A8020515C79B32ACAB010" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="74355986871FF8FD321769296BD2E930" type="personal">
<mods:role id="23A28CC7C2DB030F25B30B1229CA61E5">
<mods:roleTerm id="24002879E9EBFDA77295E045CDEFE83E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5AB867CC0954C4DB04709923CB4D2C7">Esyunin, Sergei L.</mods:namePart>
<mods:nameIdentifier id="19D0F66CC90A19E843D35E4E4782119C" type="LSID">2B591857-5FAB-4B7D-B589-6C4977A3A493</mods:nameIdentifier>
<mods:affiliation id="ABA14365209ADF9F69AC57B9F63EA43B">Sergei.</mods:affiliation>
<mods:nameIdentifier id="3A8521F5CE5660915DDF2588A245E458" type="email">Sergei.Esyunin@psu.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="26987ED41E851BB1DE63AA4F99DE2A07" type="personal">
<mods:role id="60007511F7874D5916BD909A285DAA0A">
<mods:roleTerm id="70D808BD4DEF7A67FF6323DD6C691323">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8D1A161FF0626F25E821AA15B295BA3">Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier id="88BEA5A56200CAF1E313098BAC4964A8" type="LSID">F215BA2C-5072-4CBF-BA1A-5CCBE1626B08</mods:nameIdentifier>
<mods:nameIdentifier id="4B9B880ED1854F5E466EC91F301B94BC" type="email">yurmar@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D1DF0AB548615EAAFBBCB2683E414FC3">text</mods:typeOfResource>
<mods:relatedItem id="43207E4DED7C9B93B7F738D780AB3E70" type="host">
<mods:titleInfo id="714D46F4147461B3086712135BCDABAC">
<mods:title id="6811FF8CC96B0B5340BECE829A5FDAAF">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9A222AA577EDD857453B7FC6D00E1CA2">
<mods:date id="CD897E3B9DFD7A56188BCC741BE39922">2021</mods:date>
<mods:detail id="3F2F7DC1C5B3FA31A0439AC9905230EC" type="pubDate">
<mods:number id="7537444430F323B3C6294E1B8C369E8A">2021-06-04</mods:number>
</mods:detail>
<mods:detail id="DEEC16A2BC22E418F7AF6D46BD28FCD8" type="volume">
<mods:number id="B04D941C7DEE108808476A6658EF27A4">751</mods:number>
</mods:detail>
<mods:detail id="5E9121C49390D9CDA6882E83590DFECB" type="issue">
<mods:number id="C20E7B825A04CAEEE73EFE14288F7096">1</mods:number>
</mods:detail>
<mods:extent id="296E2BBA0E3A906AE4D7457D41AE5442" unit="page">
<mods:start id="992108D76F2FEE2F14D8DEDFADED89D9">68</mods:start>
<mods:end id="096CBE3386733A9FC0406CEB38B888EA">114</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B48ED4252C8C6D39FF04789069B483AE">
<mods:url id="6DCCA13838334FA8B21119A5094388B0">http://dx.doi.org/10.5852/ejt.2021.751.1381</mods:url>
</mods:location>
<mods:classification id="89740CC58871A8613A7C14F906F2990F">journal article</mods:classification>
<mods:identifier id="1512117F42F6460D17C7846716CC7DD9" type="CLB-Dataset">6279</mods:identifier>
<mods:identifier id="FE717CA5B156AB602A684FCB3FC4432D" type="DOI">10.5852/ejt.2021.751.1381</mods:identifier>
<mods:identifier id="4F287CE720EDBB241302F803E2C5A41D" type="GBIF-Dataset">4e3af683-d8a6-40de-991b-c4584d66b8cc</mods:identifier>
<mods:identifier id="D1BD32390A932C9509EA775977075E6E" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="4CB4961B9083C05795F544F355BA59C7" type="Zenodo-Dep">4906007</mods:identifier>
<mods:identifier id="FEFDC20F11ED2696BCA7DB3A742FC9E7" type="ZooBank">1D27EA41-2396-463A-B1B2-0CCF79C9F3D2</mods:identifier>
</mods:mods>
<treatment id="03F98785036CEF771C5779D2FEA36FAE" ID-DOI="http://doi.org/10.5281/zenodo.4906477" ID-GBIF-Taxon="182597842" ID-Zenodo-Dep="4906477" LSID="urn:lsid:zoobank.org:act:0EE16C9F-6CC6-4AF4-8343-750215C9FBD6" httpUri="http://treatment.plazi.org/id/03F98785036CEF771C5779D2FEA36FAE" lastPageId="22" lastPageNumber="90" pageId="19" pageNumber="87">
<subSubSection id="C34A6518036CEF721C5779D2FC106BFD" box="[611,977,1697,1724]" pageId="19" pageNumber="87" type="nomenclature">
<paragraph id="8BEF3693036CEF721C5779D2FC106BFD" blockId="19.[368,1219,1697,1798]" box="[611,977,1697,1724]" pageId="19" pageNumber="87">
<heading id="D0A781FF036CEF721C5779D2FC106BFD" box="[611,977,1697,1724]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="87" reason="2">
<taxonomicName id="4C504D10036CEF721C5779D2FCAE6BFD" ID-CoL="98YW4" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[611,879,1697,1724]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="qashqai" status="sp. nov.">
<emphasis id="B924EA81036CEF721C5779D2FCAE6BFD" bold="true" box="[611,879,1697,1724]" italics="true" pageId="19" pageNumber="87">Haplodrassus qashqai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036CEF721D4279D1FC106BFD" box="[886,977,1698,1724]" pageId="19" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036CEF721F4479B5FC9A6A47" pageId="19" pageNumber="87" type="description">
<paragraph id="8BEF3693036CEF721F4479B5FB026BA0" blockId="19.[368,1219,1697,1798]" box="[368,1219,1734,1761]" pageId="19" pageNumber="87">
<uri id="FFC13A91036CEF721F4479B5FB026BA0" box="[368,1219,1734,1761]" pageId="19" pageNumber="87">
urn:lsid:zoobank.org:act: 
<uuid id="FFF60C46036CEF721CA479B5FB026BA0" box="[656,1219,1734,1761]" pageId="19" pageNumber="87">0EE16C9F-6CC6-4AF4-8343-750215C9FBD6</uuid>
</uri>
</paragraph>
<paragraph id="8BEF3693036CEF721CEC799FFC9A6A47" blockId="19.[368,1219,1697,1798]" box="[728,859,1772,1798]" pageId="19" pageNumber="87">
<figureCitation id="136B2A16036CEF721CEC799FFC9A6A47" box="[728,859,1772,1798]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="20.[189,232,1929,1955]" captionStartId-1="21.[190,233,1927,1953]" captionStartId-2="22.[189,232,1771,1797]" captionTargetBox-0="[189,1397,839,1891]" captionTargetBox-1="[189,1398,1162,1886]" captionTargetBox-2="[189,1398,843,1721]" captionTargetId-0="figure-131@20.[189,1398,839,1891]" captionTargetId-1="figure-278@21.[189,1398,1162,1886]" captionTargetId-2="figure-174@22.[189,1398,843,1724]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="Fig. 12. Haplodrassus qashqai sp. nov. (A–B = paratype, ♀ from Lorestan Province, MHNG; C–H = holotype, ♂, MHNG). A, C. Habitus, dorsal view. B, D. Habitus, ventral view. E–F. Bulb, ventral and proventral views. G–H. Palp, ventral and retrolateral views. Scale bars: 0.2 mm, unless stated otherwise." captionText-1="Fig. 13. Haplodrassus qashqai sp. nov., paratype, ♂ from Lorestan Province (MHNG), SEM graphs of the male palp. A. Ventral view. B. Embolus, dorsal view. C. Apical portion of bulb. D. Tibial apophysis, retrolateral view. Scale bars: 0.1 mm." captionText-2="Fig. 14. Haplodrassus qashqai sp. nov. (A–C= holotype, ♂, MHNG; D–H = paratype, ♀ from Lorestan Province, MHNG). A–B. Palp, ventral and retrolateral views. C. Tibial apophysis, retrodorsal view. D–E. Epigyne, ventral and dorsal views. F. Intact epigyne, ventral view. G–H. Macerated epigyne, ventral and dorsal views. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4906041" figureDoi-1="http://doi.org/10.5281/zenodo.4906043" figureDoi-2="http://doi.org/10.5281/zenodo.4906047" httpUri-0="https://zenodo.org/record/4906041/files/figure.png" httpUri-1="https://zenodo.org/record/4906043/files/figure.png" httpUri-2="https://zenodo.org/record/4906047/files/figure.png" pageId="19" pageNumber="87">Figs 12–14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036CEF751E89785CFD8A6CF0" lastPageId="20" lastPageNumber="88" pageId="19" pageNumber="87" type="diagnosis">
<paragraph id="8BEF3693036CEF721E89785CFEF76A08" blockId="19.[189,310,1839,1865]" box="[189,310,1839,1865]" pageId="19" pageNumber="87">
<heading id="D0A781FF036CEF721E89785CFEF76A08" bold="true" box="[189,310,1839,1865]" fontSize="11" level="3" pageId="19" pageNumber="87" reason="3">
<emphasis id="B924EA81036CEF721E89785CFEF76A08" bold="true" box="[189,310,1839,1865]" pageId="19" pageNumber="87">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036CEF751E897813FD8A6CF0" blockId="19.[189,1399,1888,2026]" lastBlockId="20.[189,1400,265,433]" lastPageId="20" lastPageNumber="88" pageId="19" pageNumber="87">
The new species differs from other males (
<taxonomicName id="4C504D10036CEF721CF07812FB886A3A" authority="(L. Koch, 1866)" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[708,1097,1888,1915]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="dalmatensis">
<emphasis id="B924EA81036CEF721CF07812FCB96A3B" box="[708,888,1888,1914]" italics="true" pageId="19" pageNumber="87">H. dalmatensis</emphasis>
(L. Koch, 1866)
</taxonomicName>
, 
<taxonomicName id="4C504D10036CEF721A627812FB1B6A3B" baseAuthorityName="O.Pickard-Cambridge" baseAuthorityYear="1872" box="[1110,1242,1889,1914]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="B924EA81036CEF721A627812FB1B6A3B" box="[1110,1242,1889,1914]" italics="true" pageId="19" pageNumber="87">H. omissus</emphasis>
</taxonomicName>
(O. Pickard- Cambridge, 1872), 
<taxonomicName id="4C504D10036CEF721F9D78F5FCCC6AE1" authority="(L. Koch, 1866)" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[425,781,1925,1952]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B924EA81036CEF721F9D78F5FD816ADE" box="[425,576,1925,1951]" italics="true" pageId="19" pageNumber="87">H. orientalis</emphasis>
(L. Koch, 1866)
</taxonomicName>
, 
<taxonomicName id="4C504D10036CEF721D2878F5FB856AE1" authority="(Lucas, 1846)" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[796,1092,1925,1952]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B924EA81036CEF721D2878F5FC526ADE" box="[796,915,1925,1951]" italics="true" pageId="19" pageNumber="87">H. rufipes</emphasis>
(Lucas, 1846)
</taxonomicName>
, 
<taxonomicName id="4C504D10036CEF721A6778F5FEA06A85" authority="Bosmans &amp; Abrous, 2018" authorityName="Bosmans &amp; Abrous" authorityYear="2018" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="87" phylum="Arthropoda" rank="species" species="securifer">
<emphasis id="B924EA81036CEF721A6778F5FB226ADE" box="[1107,1251,1925,1951]" italics="true" pageId="19" pageNumber="87">H. securifer</emphasis>
Bosmans &amp; Abrous, 2018
</taxonomicName>
) that have one or two pointed teeth on the embolic apophysis and/or have the indented tibial apophysis by a combination of a largely serrated inner surface of the embolic apophysis and a tibial apophysis with a single pointed tip. Females of the new species resemble those of 
<taxonomicName id="4C504D10036BEF751A907E79FDA56C07" authority="Marusik, Hippa &amp; Koponen, 1996" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudosignifer">
<emphasis id="B924EA81036BEF751A907E79FAB96C62" box="[1188,1400,265,291]" italics="true" pageId="20" pageNumber="88">H. pseudosignifer</emphasis>
Marusik, Hippa &amp; Koponen, 1996
</taxonomicName>
, 
<taxonomicName id="4C504D10036BEF751C407E5EFC676C07" authority="(Simon, 1878)" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[628,934,300,326]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="88" phylum="Arthropoda" rank="species" species="typhon">
<emphasis id="B924EA81036BEF751C407E5EFD2C6C07" box="[628,749,300,326]" italics="true" pageId="20" pageNumber="88">H. typhon</emphasis>
(Simon, 1878)
</taxonomicName>
, 
<taxonomicName id="4C504D10036BEF751D837E5EFEC56C2B" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="88" phylum="Arthropoda" rank="species" species="invalidus">
<emphasis id="B924EA81036BEF751D837E5EFB8A6C07" box="[951,1099,300,326]" italics="true" pageId="20" pageNumber="88">H. invalidus</emphasis>
(O. Pickard-Cambridge, 1872)
</taxonomicName>
and 
<taxonomicName id="4C504D10036BEF751F747E23FD4D6C2B" authority="(Simon, 1878)" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[320,652,335,362]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="88" phylum="Arthropoda" rank="species" species="concertor">
<emphasis id="B924EA81036BEF751F747E23FE186C28" box="[320,473,336,361]" italics="true" pageId="20" pageNumber="88">H. concertor</emphasis>
(Simon, 1878)
</taxonomicName>
in having the epigynal plate with almost parallel margins and conical posterior ends but differ by the presence of distinct longitudinal folds along the spermathecal sacs (vs bell-shaped in all others).
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036BEF751E897EA4FCFF6F02" pageId="20" pageNumber="88" type="etymology">
<paragraph id="8BEF3693036BEF751E897EA4FE856CB0" blockId="20.[189,324,471,497]" box="[189,324,471,497]" pageId="20" pageNumber="88">
<heading id="D0A781FF036BEF751E897EA4FE856CB0" bold="true" box="[189,324,471,497]" fontSize="11" level="3" pageId="20" pageNumber="88" reason="3">
<emphasis id="B924EA81036BEF751E897EA4FE856CB0" bold="true" box="[189,324,471,497]" pageId="20" pageNumber="88">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036BEF751E897D75FCFF6F02" blockId="20.[189,1398,518,579]" pageId="20" pageNumber="88">
The specific epithet is a noun in apposition, referring to a conglomeration of clans in 
<collectingCountry id="F3477603036BEF751A9C7D75FB166F61" box="[1192,1239,518,544]" name="Iran" pageId="20" pageNumber="88">Iran</collectingCountry>
consisting of mostly Turkic peoples but also Lurs, Kurds and Arabs.
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036BEF741E897D18FC4A6C28" lastPageId="21" lastPageNumber="89" pageId="20" pageNumber="88" type="materials_examined">
<paragraph id="8BEF3693036BEF751E897D18FEAD6FC5" blockId="20.[189,364,618,645]" box="[189,364,618,645]" pageId="20" pageNumber="88">
<heading id="D0A781FF036BEF751E897D18FEAD6FC5" bold="true" box="[189,364,618,645]" fontSize="11" level="3" pageId="20" pageNumber="88" reason="3">
<emphasis id="B924EA81036BEF751E897D18FEAD6FC5" bold="true" box="[189,364,618,645]" pageId="20" pageNumber="88">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036BEF751EDE7DEAFE1F6FB8" blockId="20.[189,1399,665,762]" pageId="20" pageNumber="88">
<materialsCitation id="3B383CCE036BEF751EDE7DEAFE1F6FB8" ID-GBIF-Occurrence="3127434328" collectingDate="2018-02-16" collectionCode="MHNG" collectorName="A. Zamani" country="Iran" latitude="27.55" location="Siahu" longLatPrecision="1236" longitude="55.633335" pageId="20" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Hormozgan Province" typeStatus="holotype">
<emphasis id="B924EA81036BEF751EDE7DEAFE9D6FF2" bold="true" box="[234,348,665,691]" pageId="20" pageNumber="88">
<typeStatus id="54EB8831036BEF751EDE7DEAFE9D6FF2" box="[234,348,665,691]" pageId="20" pageNumber="88" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3477603036BEF751E897DCFFEC96F97" box="[189,264,700,726]" name="Iran" pageId="20" pageNumber="88">IRAN</collectingCountry>
• 
<specimenCount id="9D56FD1A036BEF751F167DCEFEFA6F96" box="[290,315,701,727]" pageId="20" pageNumber="88" type="male">♂</specimenCount>
; 
<collectingRegion id="4994F871036BEF751F7C7DCEFD846F97" box="[328,581,700,727]" country="Iran" name="Hormozgan" pageId="20" pageNumber="88">Hormozgan Province</collectingRegion>
, 
<locationDeviation id="9A5D2FB1036BEF751C657DCFFC496F97" box="[593,904,700,727]" location="Bandar Abbas" pageId="20" pageNumber="88">
<quantity id="4CA89B76036BEF751C657DCFFD5A6F97" box="[593,667,700,727]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="88" unit="km" value="75.0">75 km</quantity>
N of Bandar Abbas
</locationDeviation>
, 
<location id="8E8F6048036BEF751DA77DCFFC186F97" LSID="urn:lsid:plazi:treatment:03F98785036CEF771C5779D2FEA36FAE:8E8F6048036BEF751DA77DCFFC186F97" box="[915,985,700,726]" country="Iran" latitude="27.55" longLatPrecision="1236" longitude="55.633335" name="Siahu" pageId="20" pageNumber="88" stateProvince="Hormozgan Province">Siahu</location>
; 
<geoCoordinate id="EE645054036BEF751DD27DC8FB936F96" box="[998,1106,699,727]" degrees="27" direction="north" minutes="33" orientation="latitude" pageId="20" pageNumber="88" precision="925" value="27.55">27°33′ N</geoCoordinate>
, 
<geoCoordinate id="EE645054036BEF751A6A7DC8FB066F96" box="[1118,1223,699,727]" degrees="55" direction="east" minutes="38" orientation="longitude" pageId="20" pageNumber="88" precision="925" value="55.633335">55°38′ E</geoCoordinate>
; 
<date id="FFEE1053036BEF751AE07DCFFAB36F97" box="[1236,1394,700,726]" pageId="20" pageNumber="88" value="2018-02-16">
<collectingDate id="EFAAE9BB036BEF751AE07DCFFAB36F97" box="[1236,1394,700,726]" pageId="20" pageNumber="88" value="2018-02-16">16 Feb. 2018</collectingDate>
</date>
; 
<collectorName id="26A55345036BEF751E897D93FEFA6FB8" box="[189,315,735,762]" pageId="20" pageNumber="88">A. Zamani</collectorName>
leg.; 
<collectionCode id="ED41AE56036BEF751F487DACFE1B6FB8" box="[380,474,735,761]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="20" pageNumber="88" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF2F661B036BEF751E8978FAFAB66AA8" ID-DOI="http://doi.org/10.5281/zenodo.4906041" ID-Zenodo-Dep="4906041" httpUri="https://zenodo.org/record/4906041/files/figure.png" pageId="20" pageNumber="88" startId="20.[189,232,1929,1955]" targetBox="[189,1397,839,1891]" targetPageId="20">
<paragraph id="8BEF3693036BEF751E8978FAFAB66AA8" blockId="20.[189,1399,1929,2026]" pageId="20" pageNumber="88">
<emphasis id="B924EA81036BEF751E8978FAFEDA6AE2" bold="true" box="[189,283,1929,1955]" pageId="20" pageNumber="88">Fig. 12.</emphasis>
<taxonomicName id="4C504D10036BEF751F1178FAFDEF6AE2" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[293,558,1929,1955]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="88" phylum="Arthropoda" rank="species" species="qashqai" status="sp. nov.">
<emphasis id="B924EA81036BEF751F1178FAFDEF6AE2" box="[293,558,1929,1955]" italics="true" pageId="20" pageNumber="88">Haplodrassus qashqai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036BEF751C0378F9FD506AE5" box="[567,657,1930,1956]" pageId="20" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
(A–B = paratype, ♀ from Lorestan Province, MHNG; C–H = holotype, ♂, MHNG). 
<emphasis id="B924EA81036BEF751FFF78DFFDC46A87" bold="true" box="[459,517,1964,1990]" pageId="20" pageNumber="88">A, C</emphasis>
. Habitus, dorsal view. 
<emphasis id="B924EA81036BEF751D2278DFFC8C6A87" bold="true" box="[790,845,1964,1990]" pageId="20" pageNumber="88">B, D</emphasis>
. Habitus, ventral view. 
<emphasis id="B924EA81036BEF751A5378DFFB5A6A87" bold="true" box="[1127,1179,1964,1990]" pageId="20" pageNumber="88">E–F</emphasis>
. Bulb, ventral and proventral views. 
<emphasis id="B924EA81036BEF751FB878BCFE086AA8" bold="true" box="[396,457,1999,2025]" pageId="20" pageNumber="88">G–H</emphasis>
. Palp, ventral and retrolateral views. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph id="8BEF3693036AEF741EDE7E7AFC4A6C28" blockId="21.[189,1399,265,361]" pageId="21" pageNumber="89">
<materialsCitation id="3B383CCE036AEF741EDE7E7AFB976C07" ID-GBIF-Occurrence="3127434329" collectingDate="2018-03" collectionCode="MHNG" collectorName="A. Zamani" country="Iran" location="Dezful" pageId="21" pageNumber="89" specimenCount="1" specimenCount-female="1" stateProvince="Khuzestan Province" typeStatus="paratype">
<emphasis id="B924EA81036AEF741EDE7E7AFEA66C62" bold="true" box="[234,359,265,291]" pageId="21" pageNumber="89">
<typeStatus id="54EB8831036AEF741EDE7E7AFEA66C62" box="[234,359,265,291]" pageId="21" pageNumber="89" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3477603036AEF741E897E5FFEC96C07" box="[189,264,300,326]" name="Iran" pageId="21" pageNumber="89">IRAN</collectingCountry>
• 
<specimenCount id="9D56FD1A036AEF741F167E5FFE8E6C07" box="[290,335,300,326]" pageId="21" pageNumber="89" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="4994F871036AEF741F697E5FFD8A6C07" box="[349,587,300,326]" country="Iran" name="Khuzestan" pageId="21" pageNumber="89">Khuzestan Province</collectingRegion>
, 
<location id="8E8F6048036AEF741C637E5FFD6A6C07" LSID="urn:lsid:plazi:treatment:03F98785036CEF771C5779D2FEA36FAE:8E8F6048036AEF741C637E5FFD6A6C07" box="[599,683,300,326]" country="Iran" name="Dezful" pageId="21" pageNumber="89" stateProvince="Khuzestan Province">Dezful</location>
; 
<date id="FFEE1053036AEF741C827E5FFCF36C07" box="[694,818,300,326]" pageId="21" pageNumber="89" value="2018-03">
<collectingDate id="EFAAE9BB036AEF741C827E5FFCF36C07" box="[694,818,300,326]" pageId="21" pageNumber="89" value="2018-03">Mar. 2018</collectingDate>
</date>
; 
<collectorName id="26A55345036AEF741D087E5FFC7B6C07" box="[828,954,300,326]" pageId="21" pageNumber="89">A. Zamani</collectorName>
leg.; 
<collectionCode id="ED41AE56036AEF741DC87E5FFB976C07" box="[1020,1110,300,326]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="89" type="Museum">MHNG</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B383CCE036AEF741A5A7E5FFC496C28" ID-GBIF-Occurrence="3127434325" collectingDate="2017-04" collectionCode="MHNG" collectorName="A. Zamani" country="Iran" location="Pol-e Dokhtar" pageId="21" pageNumber="89" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Lorestan Province" typeStatus="paratype">
<specimenCount id="9D56FD1A036AEF741A5A7E5FFB736C07" box="[1134,1202,300,326]" pageId="21" pageNumber="89" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="9D56FD1A036AEF741A8B7E5FFAC06C07" box="[1215,1281,300,326]" pageId="21" pageNumber="89" type="female">6 ♀♀</specimenCount>
; 
<collectingRegion id="4994F871036AEF741B247E5FFEE96C28" country="Iran" name="Lorestan" pageId="21" pageNumber="89">Lorestan Province</collectingRegion>
, 
<location id="8E8F6048036AEF741F007E3CFE1C6C28" LSID="urn:lsid:plazi:treatment:03F98785036CEF771C5779D2FEA36FAE:8E8F6048036AEF741F007E3CFE1C6C28" box="[308,477,335,361]" country="Iran" name="Pol-e Dokhtar" pageId="21" pageNumber="89" stateProvince="Lorestan Province">Pol-e Dokhtar</location>
; 
<date id="FFEE1053036AEF741FD37E3CFDA16C28" box="[487,608,335,361]" pageId="21" pageNumber="89" value="2017-04">
<collectingDate id="EFAAE9BB036AEF741FD37E3CFDA16C28" box="[487,608,335,361]" pageId="21" pageNumber="89" value="2017-04">Apr. 2017</collectingDate>
</date>
; 
<collectorName id="26A55345036AEF741C5E7E3CFD296C28" box="[618,744,335,361]" pageId="21" pageNumber="89">A. Zamani</collectorName>
leg.; 
<collectionCode id="ED41AE56036AEF741D1E7E3CFC496C28" box="[810,904,335,361]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="89" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036AEF771E897EE3FB986F1A" lastPageId="22" lastPageNumber="90" pageId="21" pageNumber="89" type="description">
<paragraph id="8BEF3693036AEF741E897EE3FE8F6CEB" blockId="21.[189,334,400,426]" box="[189,334,400,426]" pageId="21" pageNumber="89">
<heading id="D0A781FF036AEF741E897EE3FE8F6CEB" bold="true" box="[189,334,400,426]" fontSize="11" level="3" pageId="21" pageNumber="89" reason="3">
<emphasis id="B924EA81036AEF741E897EE3FE8F6CEB" bold="true" box="[189,334,400,426]" pageId="21" pageNumber="89">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036AEF741EDE7ECCFE6A6C9B" blockId="21.[188,1401,447,790]" box="[234,427,447,474]" pageId="21" pageNumber="89">
<emphasis id="B924EA81036AEF741EDE7ECCFEEB6C98" bold="true" box="[234,298,447,473]" pageId="21" pageNumber="89">Male</emphasis>
(
<typeStatus id="54EB8831036AEF741F0D7ECCFE626C9B" box="[313,419,447,474]" pageId="21" pageNumber="89" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BEF3693036AEF741E897E91FDD26E57" blockId="21.[188,1401,447,790]" pageId="21" pageNumber="89">
Habitus as in 
<figureCitation id="136B2A16036AEF741F687E91FE2B6CBD" box="[348,490,482,508]" captionStart="Fig" captionStartId="20.[189,232,1929,1955]" captionTargetBox="[189,1397,839,1891]" captionTargetId="figure-131@20.[189,1398,839,1891]" captionTargetPageId="20" captionText="Fig. 12. Haplodrassus qashqai sp. nov. (A–B = paratype, ♀ from Lorestan Province, MHNG; C–H = holotype, ♂, MHNG). A, C. Habitus, dorsal view. B, D. Habitus, ventral view. E–F. Bulb, ventral and proventral views. G–H. Palp, ventral and retrolateral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906041" httpUri="https://zenodo.org/record/4906041/files/figure.png" pageId="21" pageNumber="89">Fig. 12C–D</figureCitation>
. Total length 5.00. Carapace 2.26 long, 1.87 wide. Eye sizes and interdistance of PMEs: AME = 0.13, ALE = 0.15, PME = 0.14, PLE = 0.10, PME–PME = 0.08. Carapace dark brown, lighter in pars thoracica, with faded radiating lines from fovea to submarginal area. Chelicerae, maxillae and labium dark brown. Sternum light brown, slightly darker anteriorly. Chelicera with 2 pro- and 2 retromarginal teeth. Legs light yellowish and without annulations. Abdomen dorsally dark gray, with numerous transverse light bands, forming a patch posteriorly, ventrally with distinct tracheal marks and a V-shaped light pattern surrounding them. Spinnerets light gray and uniform in color. Measurements of legs: I: 6.19 (1.76, 1.00, 1.42, 1.16, 0.85), II missing, III: 4.74 (1.32, 0.68, 0.91, 1.02, 0.81), IV: 6.82 (1.82, 0.96, 1.47, 1.60, 0.97).
</paragraph>
<paragraph id="8BEF3693036AEF741E897C4EFC396EDC" blockId="21.[187,1399,829,926]" pageId="21" pageNumber="89">
Palp as in 
<figureCitation id="136B2A16036AEF741F057C4EFE7E6E16" box="[305,447,829,855]" captionStart="Fig" captionStartId="20.[189,232,1929,1955]" captionTargetBox="[189,1397,839,1891]" captionTargetId="figure-131@20.[189,1398,839,1891]" captionTargetPageId="20" captionText="Fig. 12. Haplodrassus qashqai sp. nov. (A–B = paratype, ♀ from Lorestan Province, MHNG; C–H = holotype, ♂, MHNG). A, C. Habitus, dorsal view. B, D. Habitus, ventral view. E–F. Bulb, ventral and proventral views. G–H. Palp, ventral and retrolateral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906041" httpUri="https://zenodo.org/record/4906041/files/figure.png" pageId="21" pageNumber="89">Figs 12E–H</figureCitation>
, 
<figureCitation id="136B2A16036AEF741FFD7C4EFDE36E16" box="[457,546,829,855]" captionStart="Fig" captionStartId="21.[190,233,1927,1953]" captionTargetBox="[189,1398,1162,1886]" captionTargetId="figure-278@21.[189,1398,1162,1886]" captionTargetPageId="21" captionText="Fig. 13. Haplodrassus qashqai sp. nov., paratype, ♂ from Lorestan Province (MHNG), SEM graphs of the male palp. A. Ventral view. B. Embolus, dorsal view. C. Apical portion of bulb. D. Tibial apophysis, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906043" httpUri="https://zenodo.org/record/4906043/files/figure.png" pageId="21" pageNumber="89">13A–D</figureCitation>
, 
<figureCitation id="136B2A16036AEF741C197C4EFD446E16" box="[557,645,829,855]" captionStart="Fig" captionStartId="22.[189,232,1771,1797]" captionTargetBox="[189,1398,843,1721]" captionTargetId="figure-174@22.[189,1398,843,1724]" captionTargetPageId="22" captionText="Fig. 14. Haplodrassus qashqai sp. nov. (A–C= holotype, ♂, MHNG; D–H = paratype, ♀ from Lorestan Province, MHNG). A–B. Palp, ventral and retrolateral views. C. Tibial apophysis, retrodorsal view. D–E. Epigyne, ventral and dorsal views. F. Intact epigyne, ventral view. G–H. Macerated epigyne, ventral and dorsal views. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906047" httpUri="https://zenodo.org/record/4906047/files/figure.png" pageId="21" pageNumber="89">14A–C</figureCitation>
. Tibial apophysis as long as tibia, paddle-shaped, angled dorsally, with a pointed tip; embolic apophysis with distinctly serrated inner surface; embolus with wide, blunt end and with an additional pointed outgrowth that arises from its base.
</paragraph>
<paragraph id="8BEF3693036AEF741EDE7CB6FD6A6EA1" blockId="21.[188,1399,965,1097]" box="[234,683,965,992]" pageId="21" pageNumber="89">
<emphasis id="B924EA81036AEF741EDE7CB6FE846E9E" bold="true" box="[234,325,965,991]" pageId="21" pageNumber="89">Female</emphasis>
(
<typeStatus id="54EB8831036AEF741F617CB6FE7E6E9E" box="[341,447,965,991]" pageId="21" pageNumber="89" type="paratype">paratype</typeStatus>
, 
<collectingRegion id="4994F871036AEF741FFE7CB6FD636EA1" box="[458,674,965,992]" country="Iran" name="Lorestan" pageId="21" pageNumber="89">Lorestan Province</collectingRegion>
)
</paragraph>
<paragraph id="8BEF3693036AEF771E897C9BFA916CCC" blockId="21.[188,1399,965,1097]" lastBlockId="22.[187,1398,265,397]" lastPageId="22" lastPageNumber="90" pageId="21" pageNumber="89">
Habitus as in 
<figureCitation id="136B2A16036AEF741F6F7C9BFE2C6943" box="[347,493,1000,1026]" captionStart="Fig" captionStartId="20.[189,232,1929,1955]" captionTargetBox="[189,1397,839,1891]" captionTargetId="figure-131@20.[189,1398,839,1891]" captionTargetPageId="20" captionText="Fig. 12. Haplodrassus qashqai sp. nov. (A–B = paratype, ♀ from Lorestan Province, MHNG; C–H = holotype, ♂, MHNG). A, C. Habitus, dorsal view. B, D. Habitus, ventral view. E–F. Bulb, ventral and proventral views. G–H. Palp, ventral and retrolateral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906041" httpUri="https://zenodo.org/record/4906041/files/figure.png" pageId="21" pageNumber="89">Figs 12A–B</figureCitation>
. Total length 5.49. Carapace 2.54 long, 1.93 wide. Eye sizes and interdistance of PMEs: AME = 0.09, ALE = 0.14, PME = 0.15, PLE = 0.09, PME–PME = 0.05. Carapace, chelicerae, maxillae, labium and sternum light yellowish brown. Chelicera with 3 pro- and 2 retromarginal teeth. Legs light gray and without annulations. Abdomen dorsally dark gray, with two longitudinal light gray bands leading to numerous transverse light bands, ventrally with a light gray patch posteriorly. Spinnerets light gray and uniform in color. Measurements of legs: I: 6.74 (1.97, 1.13, 1.55, 1.24, 0.85), II: 5.99 (1.76, 1.01, 1.28, 1.04, 0.90), III: 5.24 (1.55, 0.78, 0.98, 1.07, 0.86), IV: 7.41 (2.05, 1.08, 1.67, 1.79, 0.82).
</paragraph>
<caption id="DF2F661B036AEF741E8A78F4FDB26AA9" ID-DOI="http://doi.org/10.5281/zenodo.4906043" ID-Zenodo-Dep="4906043" httpUri="https://zenodo.org/record/4906043/files/figure.png" pageId="21" pageNumber="89" startId="21.[190,233,1927,1953]" targetBox="[189,1398,1162,1886]" targetPageId="21">
<paragraph id="8BEF3693036AEF741E8A78F4FDB26AA9" blockId="21.[188,1398,1927,2024]" pageId="21" pageNumber="89">
<emphasis id="B924EA81036AEF741E8A78F4FED86AE0" bold="true" box="[190,281,1927,1953]" pageId="21" pageNumber="89">Fig. 13.</emphasis>
<taxonomicName id="4C504D10036AEF741F1578F4FDE96AE0" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[289,552,1927,1953]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="89" phylum="Arthropoda" rank="species" species="qashqai" status="sp. nov.">
<emphasis id="B924EA81036AEF741F1578F4FDE96AE0" box="[289,552,1927,1953]" italics="true" pageId="21" pageNumber="89">Haplodrassus qashqai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036AEF741C0478FBFD4C6AE3" box="[560,653,1928,1954]" pageId="21" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ from Lorestan Province (MHNG), SEM graphs of the male palp. 
<emphasis id="B924EA81036AEF741F5378D9FEBD6A85" bold="true" box="[359,380,1962,1988]" pageId="21" pageNumber="89">A</emphasis>
. Ventral view. 
<emphasis id="B924EA81036AEF741C1E78D8FDFF6A84" bold="true" box="[554,574,1963,1989]" pageId="21" pageNumber="89">B</emphasis>
. Embolus, dorsal view. 
<emphasis id="B924EA81036AEF741D6278D9FCAA6A85" bold="true" box="[854,875,1962,1988]" pageId="21" pageNumber="89">C</emphasis>
. Apical portion of bulb. 
<emphasis id="B924EA81036AEF741ABE78D8FB5E6A84" bold="true" box="[1162,1183,1963,1989]" pageId="21" pageNumber="89">D</emphasis>
. Tibial apophysis, retrolateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEF36930369EF771E897EC7FB986F1A" blockId="22.[187,1399,436,603]" pageId="22" pageNumber="90">
Epigyne as in 
<figureCitation id="136B2A160369EF771F687EC7FE296C8F" box="[348,488,436,462]" captionStart="Fig" captionStartId="22.[189,232,1771,1797]" captionTargetBox="[189,1398,843,1721]" captionTargetId="figure-174@22.[189,1398,843,1724]" captionTargetPageId="22" captionText="Fig. 14. Haplodrassus qashqai sp. nov. (A–C= holotype, ♂, MHNG; D–H = paratype, ♀ from Lorestan Province, MHNG). A–B. Palp, ventral and retrolateral views. C. Tibial apophysis, retrodorsal view. D–E. Epigyne, ventral and dorsal views. F. Intact epigyne, ventral view. G–H. Macerated epigyne, ventral and dorsal views. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906047" httpUri="https://zenodo.org/record/4906047/files/figure.png" pageId="22" pageNumber="90">Fig. 14D–H</figureCitation>
. Epigynal plate about 1.3 times as long as wide, with weakly sclerotized anterior margin; atrium long and thin, almost 2 times as long as wide and ⅓ of epigynal plate width, with parallel lateral margins converging posteriorly and lateral sclerotization at anterior half; receptacles round, as wide as the narrowest part of the atrium; fertilization ducts long; distinct anterior bursae present with longitudinal folds; lateral anterior glandular heads at outer edges of the bursae.
</paragraph>
</subSubSection>
<subSubSection id="C34A65180369EF771E897DF1FEA36FAE" pageId="22" pageNumber="90" type="distribution">
<paragraph id="8BEF36930369EF771E897DF1FE976FDD" blockId="22.[189,342,642,668]" box="[189,342,642,668]" pageId="22" pageNumber="90">
<heading id="D0A781FF0369EF771E897DF1FE976FDD" bold="true" box="[189,342,642,668]" fontSize="11" level="3" pageId="22" pageNumber="90" reason="3">
<emphasis id="B924EA810369EF771E897DF1FE976FDD" bold="true" box="[189,342,642,668]" pageId="22" pageNumber="90">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930369EF771E897DC2FEA36FAE" blockId="22.[189,1398,689,751]" pageId="22" pageNumber="90">
Known from the listed localities in 
<collectingRegion id="4994F8710369EF771C5F7DC2FD3B6F8A" box="[619,762,689,715]" country="Iran" name="Hormozgan" pageId="22" pageNumber="90">Hormozgan</collectingRegion>
, 
<collectingRegion id="4994F8710369EF771D3C7DC2FC446F8A" box="[776,901,689,715]" country="Iran" name="Khuzestan" pageId="22" pageNumber="90">Khuzestan</collectingRegion>
and 
<collectingRegion id="4994F8710369EF771DF67DC2FBEB6F8A" box="[962,1066,689,715]" country="Iran" name="Lorestan" pageId="22" pageNumber="90">Lorestan</collectingRegion>
Provinces, southwestern to southern 
<collectingCountry id="F34776030369EF771F1F7DA6FE9F6FAE" box="[299,350,725,751]" name="Iran" pageId="22" pageNumber="90">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>