treatments-xml/data/03/F6/C7/03F6C753FFF2FFBBFCA9DD86FEF1F824.xml
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<document id="76411D8ED3364880DE0752BFF789B63C" ID-DOI="10.21248/contrib.entomol.67.2.275-318" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638268338673" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2017" docId="03F6C753FFF2FFBBFCA9DD86FEF1F824" docLanguage="en" docName="ContribEntomol.67.2.275-318.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Eleusis panamaensis Irmler 2017, spec. nov." docType="treatment" docVersion="4" lastPageNumber="299" masterDocId="FFCFBF2BFFE5FFA3FFFFD92CFFDAFF9C" masterDocTitle="The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="318" masterPageNumber="275" pageNumber="298" updateTime="1699437037312" updateUser="plazi">
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<mods:title id="D227244B7CFC0A1BF16E3F7EC0862BCC">The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:name id="6238BBF87A3F370A131F3B6CD4510794" type="personal">
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<mods:namePart id="ABCF002CBEDCE12461F12D7D99308F8F">Irmler, Ulrich</mods:namePart>
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<mods:title id="EFCA8E88DC2CF025E65B49A9C1353A6A">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="C4494AD473803006718B5F2B152FE8FC">
<mods:date id="F8E92CADC84608F557073655C98607CC">2017</mods:date>
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<mods:number id="2B1A801E6411CD07E87D161685ED4ED7">67</mods:number>
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<mods:identifier id="0479080D05D5FF872EB80452062FCB61" type="ISSN">0005-805X</mods:identifier>
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<treatment id="03F6C753FFF2FFBBFCA9DD86FEF1F824" ID-DOI="http://doi.org/10.5281/zenodo.5742696" ID-Zenodo-Dep="5742696" LSID="urn:lsid:plazi:treatment:03F6C753FFF2FFBBFCA9DD86FEF1F824" httpUri="http://treatment.plazi.org/id/03F6C753FFF2FFBBFCA9DD86FEF1F824" lastPageId="24" lastPageNumber="299" pageId="23" pageNumber="298">
<subSubSection id="C34525CEFFF2FFB4FCA9DD86FA81FB7A" pageId="23" pageNumber="298" type="nomenclature">
<paragraph id="8BE07645FFF2FFB4FCA9DD86FB46FB59" blockId="23.[854,1371,1194,1286]" box="[854,1180,1194,1221]" pageId="23" pageNumber="298">
<heading id="D0A8C129FFF2FFB4FCA9DD86FB46FB59" box="[854,1180,1194,1221]" fontSize="12" level="2" pageId="23" pageNumber="298" reason="2">
<taxonomicName id="4C5F0DC6FFF2FFB4FCA9DD86FBF6FB59" ID-CoL="398QW" authority="Irmler, 2017" authorityName="Irmler" authorityYear="2017" box="[854,1068,1194,1221]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="panamaensis" status="sp. nov.">
<emphasis id="B92BAA57FFF2FFB4FCA9DD86FBF6FB59" box="[854,1068,1194,1221]" italics="true" pageId="23" pageNumber="298">Eleusis panamaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218172CFFF2FFB4FBCDDD86FB46FB59" box="[1074,1180,1194,1221]" pageId="23" pageNumber="298" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BE07645FFF2FFB4FCA9DDFEFA81FB7A" blockId="23.[854,1371,1194,1286]" box="[854,1371,1234,1254]" pageId="23" pageNumber="298">
<uri id="FFCE7A47FFF2FFB4FCA9DDFEFA81FB7A" box="[854,1371,1234,1254]" pageId="23" pageNumber="298">
urn:lsid:zoobank.org:act:
<uuid id="FFF94C90FFF2FFB4FBFCDDFEFA81FB7A" box="[1027,1371,1234,1254]" pageId="23" pageNumber="298">7078364E-083E-4BC7-ADE2-045789D19FFA</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFF2FFB4FCA9DDDEFC0AFA9A" box="[854,976,1266,1286]" pageId="23" pageNumber="298" type="description">
<paragraph id="8BE07645FFF2FFB4FCA9DDDEFC0AFA9A" blockId="23.[854,1371,1194,1286]" box="[854,976,1266,1286]" pageId="23" pageNumber="298">
<figureCitation id="13646AC0FFF2FFB4FCA9DDDEFC56FA9A" box="[854,908,1266,1286]" captionStart="Fig" captionStartId="35.[174,207,2051,2073]" captionTargetBox="[184,1398,974,2044]" captionTargetId="figure-101@35.[174,1421,950,2050]" captionTargetPageId="35" captionText="Fig. 2427: E. peruviana (24), E. panamaensis (25); E. frater (26), E. elongata (27): aedeagus in lateral (A) and ventral (B) view," figureDoi="http://doi.org/10.5281/zenodo.5742511" httpUri="https://zenodo.org/record/5742511/files/figure.png" pageId="23" pageNumber="298">Figs 25</figureCitation>
A-C, 30H
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFF2FFB4FCCFDC0CFBB2FA04" pageId="23" pageNumber="298" type="materials_examined">
<paragraph id="8BE07645FFF2FFB4FCCFDC0CFBB2FA04" blockId="23.[816,1421,1311,1432]" pageId="23" pageNumber="298">
<emphasis id="B92BAA57FFF2FFB4FCCFDC0CFC10FAAB" bold="true" box="[816,970,1311,1336]" pageId="23" pageNumber="298">Type material</emphasis>
:
<materialsCitation id="3B377C18FFF2FFB4FC26DC0CFBBEFA04" collectingDate="2014-04-26" collectionCode="UIC, TSC" collectorName="T. Struyve" country="Panama" latitude="8.52" location="Santa Fe" longLatPrecision="781" longitude="-81.13" municipality="Piedra Alta" pageId="23" pageNumber="298" specimenCount="2" typeStatus="holotype">
male,
<typeStatus id="54E4C8E7FFF2FFB4FBE3DC0CFB58FAA4" box="[1052,1154,1312,1336]" pageId="23" pageNumber="298" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34836D5FFF2FFB4FB71DC0CFB3CFAAB" box="[1166,1254,1312,1335]" name="Panama" pageId="23" pageNumber="298">Panama</collectingCountry>
:
<collectingCountry id="F34836D5FFF2FFB4FB0CDC0CFA92FAAB" box="[1267,1352,1312,1335]" name="Panama" pageId="23" pageNumber="298">Panama</collectingCountry>
Prov.,
<location id="8E80209EFFF2FFB4FCCFDC6CFC4BFAC4" LSID="urn:lsid:plazi:treatment:03F6C753FFF2FFBBFCA9DD86FEF1F824:8E80209EFFF2FFB4FCCFDC6CFC4BFAC4" box="[816,913,1344,1368]" country="Panama" latitude="8.52" longLatPrecision="781" longitude="-81.13" municipality="Piedra Alta" name="Santa Fe" pageId="23" pageNumber="298">Santa Fé</location>
,
<collectingMunicipality id="6B84EC3FFFF2FFB4FC5FDC6CFBC7FAC4" box="[928,1053,1344,1368]" pageId="23" pageNumber="298">Piedra Alta</collectingMunicipality>
(
<geoCoordinate id="EE6B1082FFF2FFB4FBCFDC6CFBA7FAC4" box="[1072,1149,1344,1368]" degrees="8.52" direction="north" orientation="latitude" pageId="23" pageNumber="298" precision="555" value="8.52">8.52°N</geoCoordinate>
, -
<geoCoordinate id="EE6B1082FFF2FFB4FB6BDC6CFB2CFAC4" box="[1172,1270,1344,1368]" degrees="81.13" direction="west" orientation="longitude" pageId="23" pageNumber="298" precision="555" value="-81.13">81.13°W</geoCoordinate>
), dead wood,
<date id="FFE15085FFF2FFB4FCCFDC4CFC42FAE4" box="[816,920,1376,1400]" pageId="23" pageNumber="298" value="2014-04-26">
<collectingDate id="EFA5A96DFFF2FFB4FCCFDC4CFC42FAE4" box="[816,920,1376,1400]" pageId="23" pageNumber="298" value="2014-04-26">26.4.2014</collectingDate>
</date>
, leg.
<collectorName id="26AA1393FFF2FFB4FC28DC4CFB98FAE4" box="[983,1090,1376,1400]" pageId="23" pageNumber="298">T. Struyve</collectorName>
(
<collectionCode id="ED4EEE80FFF2FFB4FBABDC4CFB5DFAE4" box="[1108,1159,1376,1400]" pageId="23" pageNumber="298">UIC</collectionCode>
).
<typeStatus id="54E4C8E7FFF2FFB4FB61DC4CFB24FAE4" box="[1182,1278,1376,1400]" pageId="23" pageNumber="298" type="paratype">Paratype</typeStatus>
: female with same data as holotype (
<collectionCode id="ED4EEE80FFF2FFB4FBD8DCACFB86FA04" box="[1063,1116,1408,1432]" country="0" httpUri="http://grbio.org/cool/vspn-j4n9" name="Tarleton State University, Tarleton State Collection" pageId="23" pageNumber="298">TSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFF2FFB4FCCFDC93FC54F8CB" pageId="23" pageNumber="298" type="diagnosis">
<paragraph id="8BE07645FFF2FFB4FCCFDC93FC54F8CB" blockId="23.[816,1421,1471,1880]" pageId="23" pageNumber="298">
<emphasis id="B92BAA57FFF2FFB4FCCFDC93FC45FA4B" bold="true" box="[816,927,1471,1495]" pageId="23" pageNumber="298">Diagnosis</emphasis>
: The species closely resembles
<taxonomicName id="4C5F0DC6FFF2FFB4FAECDCECFAB0FA44" authorityName="FAUVEL" authorityYear="1864" box="[1299,1386,1472,1496]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FAECDCECFAB0FA44" box="[1299,1386,1472,1496]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
in size, shape, and colouration. It is certainly the sister species of
<taxonomicName id="4C5F0DC6FFF2FFB4FC5DDF2CFC23F984" authorityName="FAUVEL" authorityYear="1864" box="[930,1017,1536,1560]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FC5DDF2CFC23F984" box="[930,1017,1536,1560]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
in
<collectingCountry id="F34836D5FFF2FFB4FBE1DF2CFBA2F98B" box="[1054,1144,1536,1559]" name="Panama" pageId="23" pageNumber="298">Panama</collectingCountry>
. It can be easily differentiated from
<taxonomicName id="4C5F0DC6FFF2FFB4FC4ADF0CFBD6F9A4" authorityName="FAUVEL" authorityYear="1864" box="[949,1036,1568,1592]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FC4ADF0CFBD6F9A4" box="[949,1036,1568,1592]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
by the microsculpture of the head and the elongate aedeagus. Whereas the head
<taxonomicName id="4C5F0DC6FFF2FFB4FAC9DF6CFA57F9C4" authorityName="FAUVEL" authorityYear="1864" box="[1334,1421,1600,1624]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FAC9DF6CFA57F9C4" box="[1334,1421,1600,1624]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
has only remains of microsculpture with large parts polished, the head of.
<taxonomicName id="4C5F0DC6FFF2FFB4FBDADFACFB11F90B" authorityName="Irmler" authorityYear="2017" box="[1061,1227,1664,1687]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="panamaensis">
<emphasis id="B92BAA57FFF2FFB4FBDADFACFB11F90B" box="[1061,1227,1664,1687]" italics="true" pageId="23" pageNumber="298">E. panamaensis</emphasis>
</taxonomicName>
is totally covered by microsculpture. The head of
<taxonomicName id="4C5F0DC6FFF2FFB4FB3BDF8CFAB0F92B" authorityName="Irmler" authorityYear="2017" box="[1220,1386,1696,1719]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="panamaensis">
<emphasis id="B92BAA57FFF2FFB4FB3BDF8CFAB0F92B" box="[1220,1386,1696,1719]" italics="true" pageId="23" pageNumber="298">E. panamaensis</emphasis>
</taxonomicName>
is impunctate, whereas the head of
<taxonomicName id="4C5F0DC6FFF2FFB4FB55DFECFADBF944" authorityName="FAUVEL" authorityYear="1864" box="[1194,1281,1728,1752]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FB55DFECFADBF944" box="[1194,1281,1728,1752]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
is distinctly punctate. The eyes are smaller and temples longer in
<taxonomicName id="4C5F0DC6FFF2FFB4FCCFDE2CFC5DF884" authorityName="FAUVEL" authorityYear="1864" box="[816,903,1792,1816]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FCCFDE2CFC5DF884" box="[816,903,1792,1816]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C5F0DC6FFF2FFB4FC1CDE2CFB53F88B" authorityName="Irmler" authorityYear="2017" box="[995,1161,1792,1815]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="panamaensis">
<emphasis id="B92BAA57FFF2FFB4FC1CDE2CFB53F88B" box="[995,1161,1792,1815]" italics="true" pageId="23" pageNumber="298">E. panamaensis</emphasis>
</taxonomicName>
. The central lobe of the aedeagus is nearly twice as long in
<taxonomicName id="4C5F0DC6FFF2FFB4FB52DE0CFA89F8AB" authorityName="Irmler" authorityYear="2017" box="[1197,1363,1824,1847]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="panamaensis">
<emphasis id="B92BAA57FFF2FFB4FB52DE0CFA89F8AB" box="[1197,1363,1824,1847]" italics="true" pageId="23" pageNumber="298">E. panamaensis</emphasis>
</taxonomicName>
as in
<taxonomicName id="4C5F0DC6FFF2FFB4FCCFDE6CFC5DF8C4" authorityName="FAUVEL" authorityYear="1864" box="[816,903,1856,1880]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="298" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BAA57FFF2FFB4FCCFDE6CFC5DF8C4" box="[816,903,1856,1880]" italics="true" pageId="23" pageNumber="298">E. frater</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFF2FFBBFCCFDE53FE85F8A4" lastPageId="24" lastPageNumber="299" pageId="23" pageNumber="298" type="description">
<paragraph id="8BE07645FFF2FFB4FCCFDE53FBB1F844" blockId="23.[816,1421,1919,2104]" pageId="23" pageNumber="298">
<emphasis id="B92BAA57FFF2FFB4FCCFDE53FC68F80B" bold="true" box="[816,946,1919,1943]" pageId="23" pageNumber="298">Description</emphasis>
: Length:
<quantity id="4CA7DBA0FFF2FFB4FBE8DEACFBB1F80B" box="[1047,1131,1920,1944]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="298" unit="mm" value="4.5">4.5 mm</quantity>
. Colouration: totally black; sides of abdominal segments lighter brown; legs light brown; antennae dark brown.
</paragraph>
<paragraph id="8BE07645FFF2FFBBFCCFDECCFE25FD8B" blockId="23.[816,1421,1919,2104]" lastBlockId="24.[212,817,256,1848]" lastPageId="24" lastPageNumber="299" pageId="23" pageNumber="298">
Head:
<quantity id="4CA7DBA0FFF2FFB4FC86DECCFC0DF86B" box="[889,983,2016,2040]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="23" pageNumber="298" unit="mm" value="0.54">0.54 mm</quantity>
long,
<quantity id="4CA7DBA0FFF2FFB4FBE1DECCFBA7F86B" box="[1054,1149,2016,2040]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="23" pageNumber="298" unit="mm" value="0.57">0.57 mm</quantity>
wide; eyes large; slightly prominent; temples slightly shorter than eyes; behind eyes shortly parallel; posterior angles obtuse; preocular sides nearly as long as temples; anterior margin of frons continuously narrowed to short straight central part; central prominent part slightly emarginate; supraocular groove deep; anterior part parallel to side; slightly curved in posterior part; impunctate except setiferous puncture at each end of supraocular groove; another pair of setiferous punctures on each side of central clypeal prominence; elongate microsculpture weak, but dense; longitudinally striate; surface slightly matt.
</paragraph>
<paragraph id="8BE07645FFFDFFBBFF2BDB0CFE52FCAB" blockId="24.[212,817,256,1848]" pageId="24" pageNumber="299">Antennae slightly shorter than head and pronotum combined; first antennomere slightly thicker than following antennomeres; second antennomere only half as long as first; third slightly longer than second; following antennomeres decreasing in length and increasing in width; fourth antennomere triangular; slightly longer than wide; tenth antennomere 1.6 times wider than long; fourth to eleventh antennomere pubescent; all antennomeres with long apical setae.</paragraph>
<paragraph id="8BE07645FFFDFFBBFF2BDA6CFE66FB64" blockId="24.[212,817,256,1848]" pageId="24" pageNumber="299">
Pronotum:
<quantity id="4CA7DBA0FFFDFFBBFEB0DA6CFE74FCCB" box="[335,430,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="24" pageNumber="299" unit="mm" value="0.52">0.52 mm</quantity>
long;
<quantity id="4CA7DBA0FFFDFFBBFE10DA6CFD95FCCB" box="[495,591,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="24" pageNumber="299" unit="mm" value="0.59">0.59 mm</quantity>
wide; widest close to anterior angles; sides in anterior half parallel; in posterior half narrowed to posterior margin in even curve; anterior angles shortly rounded; at posterior margin, central part slightly prominent; prominent part marked by deep impression margined at inner side by ridge; close to anterior angles, at anterior margin two deep impressions with setiferous punctures; another deeply impressed setiferous puncture at beginning of posterior half and at posterior margin close to ridge; very finely and sparsely punctate; elongate microsculpture similar to that of head, but midline and few parts on disc without microsculpture; longitudinally striate; surface shinier than on head.
</paragraph>
<paragraph id="8BE07645FFFDFFBBFF2BDC2CFDC3FA64" blockId="24.[212,817,256,1848]" pageId="24" pageNumber="299">
Elytra:
<quantity id="4CA7DBA0FFFDFFBBFED8DC2CFE5CFA8B" box="[295,390,1280,1304]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="299" unit="mm" value="0.73">0.73 mm</quantity>
long,
<quantity id="4CA7DBA0FFFDFFBBFE2ADC2CFDEEFA8B" box="[469,564,1280,1304]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="299" unit="mm" value="0.73">0.73 mm</quantity>
wide; humeral angles shortly rounded; sides slightly divergent to widely rounded posterior angles; on each side of suture, posterior margin convex; punctation weak and sparse; similar to pronotum; along lateral margin with setiferous punctures; on each side of suture setiferous puncture on central disc; elongate microsculpture weak; with shiny interstices; transversely striate.
</paragraph>
<paragraph id="8BE07645FFFDFFBBFF2BDF2CFE3EF9C4" blockId="24.[212,817,256,1848]" pageId="24" pageNumber="299">Abdomen finely punctate; each segment with transverse row of four setiferous punctures; elongate microsculpture weak; transversely striate.</paragraph>
<paragraph id="8BE07645FFFDFFBBFF2BDF4CFE85F8A4" blockId="24.[212,817,256,1848]" pageId="24" pageNumber="299">Aedeagus elongate; apical part of central lobe much longer than basal part; in lateral aspect, apex acute, but obtusely rounded; in ventral aspect, central lobe parallel; apex widely rounded; sensillae concentrating along apical midline; at apex, denser than closer to base; parameres shorter than central lobe; transparent lobe narrow; with few sensillae.</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFDFFBBFF2BDE4CFEF1F824" pageId="24" pageNumber="299" type="etymology">
<paragraph id="8BE07645FFFDFFBBFF2BDE4CFEF1F824" blockId="24.[212,816,1888,1976]" pageId="24" pageNumber="299">
<emphasis id="B92BAA57FFFDFFBBFF2BDE4CFE91F8E4" bold="true" box="[212,331,1888,1912]" pageId="24" pageNumber="299">Etymology</emphasis>
: The species name derived from the countrys name
<collectingCountry id="F34836D5FFFDFFBBFEE7DEACFEAAF80B" box="[280,368,1920,1943]" name="Panama" pageId="24" pageNumber="299">Panama</collectingCountry>
, where the species was collected by Tim Struyve.
</paragraph>
</subSubSection>
</treatment>
</document>