276 lines
31 KiB
XML
276 lines
31 KiB
XML
<document id="92AC403A1E48A360BA734294D84CD3D5" ID-DOI="10.5281/zenodo.175330" ID-GBIF-Dataset="a0f6b836-6169-4a28-9f3b-a5cc210f5f1d" ID-ISSN="11755326" ID-Zenodo-Dep="175330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459804323977" checkinUser="plazi" docAuthor="Wilms, Thomas M. & Schmitz, Andreas" docDate="2007" docId="03E2ED68640DDF78FF37F9F4FE8899AB" docLanguage="en" docName="zt01394p023.pdf" docOrigin="Zootaxa 1394" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Uromastyx yemenensis subsp. shobraki Wilms & Schmitz, 2007, ssp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFDB95106402DF6BFFA0FF92FC0A9971" masterDocTitle="A new polytypic species of the genus Uromastyx MERREM 1820 (Reptilia: Squamata: Agamidae: Leiolepidinae) from southwestern Arabia" masterLastPageNumber="23" masterPageNumber="1" pageNumber="16" updateTime="1698220469858" updateUser="plazi">
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<mods:titleInfo id="ED5043F3158A7DE64979633B7758D323">
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<mods:title id="72502E87352F3DE5AEDE23246431FE6A">A new polytypic species of the genus Uromastyx MERREM 1820 (Reptilia: Squamata: Agamidae: Leiolepidinae) from southwestern Arabia</mods:title>
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</mods:titleInfo>
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<mods:name id="4865756A921A9172903CDAD58EA7AEF0" type="personal">
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<mods:role id="E50F5A0BA66ED8AB67F55BC30E44E5BC">
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<mods:roleTerm id="93A31751A3B9E6A6173E9A3FEB8F8935">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="343B1BE45E9BAD2D403958CB96A3E9AF">Wilms, Thomas M.</mods:namePart>
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</mods:name>
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<mods:name id="BA9E2619F34F4133DEB5DF527A4B7D61" type="personal">
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<mods:role id="8FC158F5582D96F191CE7CDBC971FED6">
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<mods:roleTerm id="F1C20236CE9F8ECD55DDF6BFD3DBB700">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4F69315D062B2E67238AB4870698E28A">Schmitz, Andreas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="65957021D0742B7605D96BEA3B8FD014">text</mods:typeOfResource>
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<mods:relatedItem id="962AE4C17875404EBE874C9B230D8B6D" type="host">
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<mods:titleInfo id="8955E7272A3F1C38FD2A2FD4614EA805">
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<mods:title id="AB4E99D55C0023FC82C21BE071540312">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="81E267F7D42CA13C48C27FFE7D8F243E">
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<mods:date id="266B6231BBFC7187E7CFE966262C39C8">2007</mods:date>
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<mods:detail id="6A0753F228A6C044079A96549C038B6D" type="volume">
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<mods:number id="4B301363A8FCD8D496A7384F8E7DA58E">1394</mods:number>
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<mods:start id="3205CE462B081A8E8984E227CB6CBD22">1</mods:start>
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<mods:end id="F4A0EFBC2396112A0CF911FF1D6A4193">23</mods:end>
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</mods:relatedItem>
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<mods:classification id="9A804DA423A763F71ED128B109ABA03E">journal article</mods:classification>
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<mods:identifier id="671F166FE646F279D4667AD049CF424D" type="DOI">10.5281/zenodo.175330</mods:identifier>
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<mods:identifier id="84B0550F04DD8EA80C35284768FE7DEE" type="GBIF-Dataset">a0f6b836-6169-4a28-9f3b-a5cc210f5f1d</mods:identifier>
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<mods:identifier id="331F5A1783313C65C22B9FB9B791127E" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="EE36D90D821B052E5EA4D3B408ED8A17" type="Zenodo-Dep">175330</mods:identifier>
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</mods:mods>
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<treatment id="03E2ED68640DDF78FF37F9F4FE8899AB" ID-DOI="http://doi.org/10.5281/zenodo.6243357" ID-GBIF-Taxon="119352696" ID-Zenodo-Dep="6243357" LSID="urn:lsid:plazi:treatment:03E2ED68640DDF78FF37F9F4FE8899AB" httpUri="http://treatment.plazi.org/id/03E2ED68640DDF78FF37F9F4FE8899AB" lastPageId="19" lastPageNumber="19" pageId="15" pageNumber="16">
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<subSubSection id="C3510FF5640DDF64FF37F9F4FE849FF1" box="[151,654,1638,1664]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BF45C7E640DDF64FF37F9F4FE849FF1" blockId="15.[151,654,1638,1664]" box="[151,654,1638,1664]" pageId="15" pageNumber="16">
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<heading id="D0BCEB12640DDF64FF37F9F4FE849FF1" bold="true" box="[151,654,1638,1664]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="B93F806C640DDF64FF37F9F4FE849FF1" bold="true" box="[151,654,1638,1664]" pageId="15" pageNumber="16">
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<taxonomicName id="4C4B27FD640DDF64FF37F9F4FE149FF1" box="[151,542,1638,1664]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="yemenensis" status="subsp. nov." subSpecies="shobraki">
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<emphasis id="B93F806C640DDF64FF37F9F4FE149FF1" bold="true" box="[151,542,1638,1664]" italics="true" pageId="15" pageNumber="16">Uromastyx yemenensis shobraki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C3D17640DDF64FD85F9F4FE849FF1" box="[549,654,1638,1664]" pageId="15" pageNumber="16" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3510FF5640DDF7BFF37F922FD2E99C3" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8BF45C7E640DDF64FF37F922FDB39F82" blockId="15.[151,1440,1712,2019]" pageId="15" pageNumber="16">
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<emphasis id="B93F806C640DDF64FF37F922FD1A9FBB" bold="true" box="[151,272,1712,1738]" pageId="15" pageNumber="16">
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<typeStatus id="54F0E2DC640DDF64FF37F922FD019FBB" box="[151,267,1712,1738]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED5AC4BB640DDF64FE81F923FD799FBA" LSID="urn:lsid:biocol.org:col:34613" box="[289,371,1713,1739]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="16">ZFMK</collectionCode>
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48681, adult male, Mafraq Mocca (Mafraq alMukha), km 13.5,
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<collectingCountry id="F35C1CEE640DDF64FBDEF923F96D9FBA" box="[1150,1383,1713,1739]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, leg. B. Schätti,
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<date id="FFF57ABE640DDF64FEB9F94BFDBF9F82" box="[281,437,1753,1779]" pageId="15" pageNumber="16" value="1988-04-05" valueMax="1988-04-06" valueMin="1988-04-05">5.–6.IV.1988</date>
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.
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</paragraph>
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<paragraph id="8BF45C7E640DDF7BFF66F893FD2E99C3" blockId="15.[151,1440,1712,2019]" lastBlockId="16.[151,1437,152,1669]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<typeStatus id="54F0E2DC640DDF64FF66F893FD379E6A" box="[198,317,1793,1819]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED5AC4BB640DDF64FEF2F893FDA99E6A" LSID="urn:lsid:biocol.org:col:34613" box="[338,419,1793,1819]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="16">ZFMK</collectionCode>
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48680, adult male, Mafraq Mocca (Mafraq alMukha), km 13.5,
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<collectingCountry id="F35C1CEE640DDF64FB11F893F9909E6A" box="[1201,1434,1793,1819]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, leg. B. Schätti,
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<date id="FFF57ABE640DDF64FEF1F8BBFDE79E32" box="[337,493,1833,1859]" pageId="15" pageNumber="16" value="1988-04-05" valueMax="1988-04-06" valueMin="1988-04-05">5.–6.IV.1988</date>
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.
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<collectionCode id="ED5AC4BB640DDF64FE5AF8BBFE5E9E32" box="[506,596,1833,1859]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16">MHNG</collectionCode>
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2496.55, adult female, Tihama, between Mocca (alMukha) and Oued Zabid,
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<collectingCountry id="F35C1CEE640DDF64FF4DF8C3FDD29E1A" box="[237,472,1873,1899]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, coll. P. A. Panchaud, leg. M. Guillod,
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<date id="FFF57ABE640DDF64FC18F8C3F8059E1A" box="[952,1039,1873,1899]" pageId="15" pageNumber="16" value="1990-05">V.1990</date>
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.
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<collectionCode id="ED5AC4BB640DDF64FBBDF8C3F87D9E1A" box="[1053,1143,1873,1899]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16">MHNG</collectionCode>
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2496.56, male, Tihama, between Mocca (alMukha) and Oued. Zabid,
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<collectingCountry id="F35C1CEE640DDF64FD6FF8EBFFB79EE2" box="[719,957,1913,1939]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, coll. P. A. Panchaud, leg. M. Guillod,
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<date id="FFF57ABE640DDF64FF37F833FCE49ECA" box="[151,238,1953,1979]" pageId="15" pageNumber="16" value="1990-05">V.1990</date>
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.
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<collectionCode id="ED5AC4BB640DDF64FEA2F833FD569ECA" box="[258,348,1953,1979]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16">MHNG</collectionCode>
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2553.56, semiadult, Mafraq alMukha, km 1.5,
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<collectingCountry id="F35C1CEE640DDF64FC70F833F8C29ECA" box="[976,1224,1953,1979]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, leg. B. Schätti,
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<date id="FFF57ABE640DDF64FF37F85BFD169E92" box="[151,284,1993,2019]" pageId="15" pageNumber="16" value="1994-04-05">05.IV.1994</date>
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.
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<collectionCode id="ED5AC4BB640DDF64FE88F85BFD889E92" box="[296,386,1993,2019]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16">MHNG</collectionCode>
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2455.100, adult female, Mafraq alMukha, km 13.5,
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<collectingCountry id="F35C1CEE640DDF64FC5AF85BF8EB9E92" box="[1018,1249,1993,2019]" name="Yemen" pageId="15" pageNumber="16">Republic of Yemen</collectingCountry>
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, leg. B. Schätti,
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<date id="FFF57ABE6412DF7BFF37FF0AFD1599C3" box="[151,287,152,178]" pageId="16" pageNumber="17" value="1989-06-19">19.VI.1989</date>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3510FF56412DF7BFF66FF2DFD8E98BB" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="8BF45C7E6412DF7BFF66FF2DFD8E98BB" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">
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<emphasis id="B93F806C6412DF7BFF66FF2DFD4C99A8" bold="true" box="[198,326,191,217]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
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<taxonomicName id="4C4B27FD6412DF7BFEF0FF52FEDA99A8" box="[336,720,192,217]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
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<emphasis id="B93F806C6412DF7BFEF0FF52FEDA99A8" box="[336,720,192,217]" italics="true" pageId="16" pageNumber="17">Uromastyx yemenensis shobraki</emphasis>
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</taxonomicName>
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shows the distinctive features listed in the diagnosis of the nominate subspecies,
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<taxonomicName id="4C4B27FD6412DF7BFE3FFF7AFE619870" box="[415,619,232,257]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="yemenensis">
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<emphasis id="B93F806C6412DF7BFE3FFF7AFE619870" box="[415,619,232,257]" italics="true" pageId="16" pageNumber="17">U. y. yemenensis</emphasis>
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</taxonomicName>
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. It differs from
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<taxonomicName id="4C4B27FD6412DF7BFC98FF7AFF919870" box="[824,923,232,257]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benti">
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<emphasis id="B93F806C6412DF7BFC98FF7AFF919870" box="[824,923,232,257]" italics="true" pageId="16" pageNumber="17">U. benti</emphasis>
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</taxonomicName>
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in having smaller scales around midbody (
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<quantity id="4CB3F19B6412DF7BFF3FFE82FD82985B" box="[159,392,272,298]" metricMagnitude="0" metricUnit="m" metricValue="4.817872" metricValueMax="5.144008" metricValueMin="4.491736" pageId="16" pageNumber="17" unit="in" value="189.68" valueMax="202.52" valueMin="176.84">189.68 +/ 12.84 in</quantity>
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||
<taxonomicName id="4C4B27FD6412DF7BFE31FE82FEA79858" box="[401,685,272,297]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
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||
<emphasis id="B93F806C6412DF7BFE31FE82FEA79858" box="[401,685,272,297]" italics="true" pageId="16" pageNumber="17">U. yemenensis shobraki</emphasis>
|
||
</taxonomicName>
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||
vs.
|
||
<quantity id="4CB3F19B6412DF7BFD40FE82FFB3985B" box="[736,953,272,298]" metricMagnitude="0" metricUnit="m" metricValue="4.06527" metricValueMax="4.293362" metricValueMin="3.837178" pageId="16" pageNumber="17" unit="in" value="160.05" valueMax="169.03" valueMin="151.07">160.05 +/ 8.98 in</quantity>
|
||
<taxonomicName id="4C4B27FD6412DF7BFC62FE82F82B9858" box="[962,1057,272,297]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benti">
|
||
<emphasis id="B93F806C6412DF7BFC62FE82F82B9858" box="[962,1057,272,297]" italics="true" pageId="16" pageNumber="17">U. benti</emphasis>
|
||
</taxonomicName>
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||
) and smaller ventrals (
|
||
<quantity id="4CB3F19B6412DF7BFA96FE82FCE29823" metricMagnitude="0" metricUnit="m" metricValue="2.2075139999999998" metricValueMax="2.3378159999999997" metricValueMin="2.0772120000000003" pageId="16" pageNumber="17" unit="in" value="86.91" valueMax="92.03999999999999" valueMin="81.78">86.91+/ 5.13 in</quantity>
|
||
<taxonomicName id="4C4B27FD6412DF7BFF4FFEAAFD9E9820" box="[239,404,312,337]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6412DF7BFF4FFEAAFD9E9820" box="[239,404,312,337]" italics="true" pageId="16" pageNumber="17">U. y. shobraki</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<quantity id="4CB3F19B6412DF7BFE63FEAAFE699823" box="[451,611,312,338]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" metricValueMax="1.9817079999999998" metricValueMin="1.777492" pageId="16" pageNumber="17" unit="in" value="74.0" valueMax="78.02" valueMin="69.98">74 +/ 4.02 in</quantity>
|
||
<taxonomicName id="4C4B27FD6412DF7BFDCAFEAAFEC29820" box="[618,712,312,337]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benti">
|
||
<emphasis id="B93F806C6412DF7BFDCAFEAAFEC29820" box="[618,712,312,337]" italics="true" pageId="16" pageNumber="17">U. benti</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C4B27FD6412DF7BFD40FEAAFFE29820" box="[736,1000,312,337]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6412DF7BFD40FEAAFFE29820" box="[736,1000,312,337]" italics="true" pageId="16" pageNumber="17">Uromastyx y. shobraki</emphasis>
|
||
</taxonomicName>
|
||
is differentiated from
|
||
<taxonomicName id="4C4B27FD6412DF7BFB4EFEAAFCBC9808" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="yemenensis">
|
||
<emphasis id="B93F806C6412DF7BFB4EFEAAFCBC9808" italics="true" pageId="16" pageNumber="17">U. y. yemenensis</emphasis>
|
||
</taxonomicName>
|
||
not only by its larger maximum size [
|
||
<quantity id="4CB3F19B6412DF7BFD28FEF2FEE6980B" box="[648,748,352,378]" metricMagnitude="-1" metricUnit="m" metricValue="3.9299999999999997" pageId="16" pageNumber="17" unit="cm" value="39.3">39.3 cm</quantity>
|
||
in
|
||
<taxonomicName id="4C4B27FD6412DF7BFCB7FEF2FFC89808" box="[791,962,352,377]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6412DF7BFCB7FEF2FFC89808" box="[791,962,352,377]" italics="true" pageId="16" pageNumber="17">U. y. shobraki</emphasis>
|
||
</taxonomicName>
|
||
(MHNG 2538.47) vs.
|
||
<quantity id="4CB3F19B6412DF7BFB76FEF2F932980B" box="[1238,1336,352,378]" metricMagnitude="-1" metricUnit="m" metricValue="3.32" pageId="16" pageNumber="17" unit="cm" value="33.2">33.2 cm</quantity>
|
||
in
|
||
<taxonomicName id="4C4B27FD6412DF7BFAC3FEF2FD1798D0" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="yemenensis">
|
||
<emphasis id="B93F806C6412DF7BFAC3FEF2FD1798D0" italics="true" pageId="16" pageNumber="17">U. y. yemenensis</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1946.8.11.68)] but also in different colour pattern (see Plate 5) and in significant genetic differences (
|
||
<tableCitation id="C6C969C56412DF7BFE88FE22FD7F98BB" box="[296,373,432,458]" captionStart="TABLE 5" captionStartId="9.[124,201,572,592]" captionTargetBox="[153,1697,626,1829]" captionTargetPageId="9" captionText="TABLE 5. Summary of the uncorrected pdistances for the 16 S data set." httpUri="http://table.plazi.org/id/DF340CF6640BDF62FFDCFDAEFF159B21" pageId="16" pageNumber="17" tableUuid="DF340CF6640BDF62FFDCFDAEFF159B21">Tab. 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3510FF56412DF7BFF66FE45F91C9F7C" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BF45C7E6412DF7BFF66FE45F89D9A4A" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">
|
||
<emphasis id="B93F806C6412DF7BFF66FE45FE2D9880" bold="true" box="[198,551,471,497]" pageId="16" pageNumber="17">
|
||
Description of the
|
||
<typeStatus id="54F0E2DC6412DF7BFE14FE45FE289880" box="[436,546,471,497]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Snoutventlength:
|
||
<quantity id="4CB3F19B6412DF7BFCB7FE4AFF719883" box="[791,891,472,498]" metricMagnitude="-1" metricUnit="m" metricValue="1.61" pageId="16" pageNumber="17" unit="cm" value="16.1">16.1 cm</quantity>
|
||
; tail length
|
||
<quantity id="4CB3F19B6412DF7BFBACFE4AF8519883" box="[1036,1115,472,498]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="cm" value="15.0">15 cm</quantity>
|
||
; head length
|
||
<quantity id="4CB3F19B6412DF7BFB5EFE4AF95C9883" box="[1278,1366,472,498]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="17" unit="mm" value="32.0">32 mm</quantity>
|
||
; head width
|
||
<quantity id="4CB3F19B6412DF7BFF41FD90FD3D9B6D" box="[225,311,514,540]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="17" unit="mm" value="33.0">33 mm</quantity>
|
||
; width of the tail between the 4th and 5th whorl:
|
||
<quantity id="4CB3F19B6412DF7BFCC7FD90FFB79B6D" box="[871,957,514,540]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="17" unit="mm" value="21.0">21 mm</quantity>
|
||
; maximum tail width at the 5th whorl:
|
||
<quantity id="4CB3F19B6412DF7BFADFFD90FCC29B35" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="17" unit="mm" value="29.0">29 mm</quantity>
|
||
; 207 scales around midbody; 97 scale rows between gular and inguinal fold, 30 gular scales counted from a line between the anterior margins of the ear openings to the mental scale; 28 (left) and 27 (right) scales between the mid of the ear openings under the jaws to the mental scale; On each side 4 scale rows between the supralabials and the enlarged suboculars; 38 scales around the 5th whorl, 27 tail whorls, the last 21 whorls are made up as single continuous scale rows; 17 subdigital lamellae beneath the 4th toe; no preanal/femoral pores, but more or less enlarged callose scales along the thighs in the position normally occupied by femoral/preanal pores. The
|
||
<typeStatus id="54F0E2DC6412DF7BFEBAFCB3FD8B9A4A" box="[282,385,801,827]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
has a total length of
|
||
<quantity id="4CB3F19B6412DF7BFDD7FCB3FED89A4A" box="[631,722,801,827]" metricMagnitude="-1" metricUnit="m" metricValue="3.11" pageId="16" pageNumber="17" unit="cm" value="31.1">31.1cm</quantity>
|
||
. The tail length is 93.17% of the SVL.
|
||
</paragraph>
|
||
<paragraph id="8BF45C7E6412DF7BFF66FCDBF8EB9AAA" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">Head covered with irregularly arranged scales of different size; the smallest above the eyes. Occipital and neck scales small and smooth. Anterior margins of the ear openings not covered with enlarged scales. Scales of the forelegs smooth, scales on the sole with a well developed median keel. Subdigital scales only slightly keeled. Gular scales and ventral scales smooth. Forelegs and back without enlarged tubercles.</paragraph>
|
||
<paragraph id="8BF45C7E6412DF7BFF66FC7BFE459DD4" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">Thigh without enlarged tubercular scales; scales on the backside of the thigh distinctly smaller than those on the front. Ventral surface of the lower leg with smooth rhomboid scales, larger than the ventral scales. Dorsal surface with enlarged tubercular scales intermixed with very small smooth scales. Scales on the upper side of the feet slightly keeled and pointed. A patch of enlarged triangular and pointed scales present between the base of the 4th and 5th toe to the ankle.</paragraph>
|
||
<paragraph id="8BF45C7E6412DF7BFF66FB21FE339D84" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">Scales on the dorsal side of the tail, strongly keeled, with the median rows only slightly pointed while the marginal rows are strongly pointed.</paragraph>
|
||
<paragraph id="8BF45C7E6412DF7BFF66FA91FF9F9C94" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">
|
||
<emphasis id="B93F806C6412DF7BFF66FA91FE269C6C" bold="true" box="[198,556,1283,1309]" pageId="16" pageNumber="17">Colouration in preservative.</emphasis>
|
||
Dorsal surface of head, body and hind limbs dark brown, tail somewhat lighter. Light brown roundish dots (diameter 4–5 scales) are present on the dorsum, tending to form transverse rows. In addition, irregular light brown dots are present on the whole dorsum. Colouration of the hands not different from the colouration of the forearm. Head dark brown, with light brown pattern. Ventral side of the specimen yellowish brown. Ventral side of head and chest marbled with anthracite and dark brown. On the sides of the belly exists an anthracitecoloured stripelike pattern.
|
||
</paragraph>
|
||
<paragraph id="8BF45C7E6412DF7BFF66FA61F91C9F7C" blockId="16.[151,1437,152,1669]" box="[198,1302,1523,1549]" pageId="16" pageNumber="17">
|
||
<emphasis id="B93F806C6412DF7BFF66FA61FD4B9F7C" bold="true" box="[198,321,1523,1549]" pageId="16" pageNumber="17">Variation.</emphasis>
|
||
For measurements and scale counts of the
|
||
<typeStatus id="54F0E2DC6412DF7BFC9CFA61FF649F7C" box="[828,878,1523,1549]" pageId="16" pageNumber="17">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C4B27FD6412DF7BFC7FFA66F88E9F7C" box="[991,1156,1524,1549]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6412DF7BFC7FFA66F88E9F7C" box="[991,1156,1524,1549]" italics="true" pageId="16" pageNumber="17">U. y. shobraki</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<tableCitation id="C6C969C56412DF7BFB18FA61F9189F7C" box="[1208,1298,1523,1549]" captionStart="TABLE 7" captionStartId="14.[151,242,1466,1490]" captionTargetBox="[159,1425,1542,1967]" captionTargetPageId="14" captionText="TABLE 7. Measurements (in mm) and scale counts of the type series of U. y. yemenensis and U. y. shobrak" httpUri="http://table.plazi.org/id/DF340CF6640CDF65FF37FA28F91E9CA0" pageId="16" pageNumber="17" tableUuid="DF340CF6640CDF65FF37FA28F91E9CA0">Table 7</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3510FF56412DF7AFF66F989F9749E52" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="8BF45C7E6412DF7BFF66F989FD8F9FF4" blockId="16.[151,1437,152,1669]" pageId="16" pageNumber="17">
|
||
<emphasis id="B93F806C6412DF7BFF66F989FD599F44" bold="true" box="[198,339,1563,1589]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
This taxon is named in honour of Dr. Mohammed Shobrak, Director of the “National Wildlife Research Centre, NWRC” in Taif,
|
||
<collectingCountry id="F35C1CEE6412DF7BFD90F9D1FEC79F2C" box="[560,717,1603,1629]" name="Saudi Arabia" pageId="16" pageNumber="17">Saudi Arabia</collectingCountry>
|
||
, for his outstanding achievements regarding the conservation of Arabian Wildlife.
|
||
</paragraph>
|
||
<paragraph id="8BF45C7E6413DF7AFF37F899F9749E52" blockId="17.[151,1406,1803,1827]" box="[151,1406,1803,1827]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93F806C6413DF7AFF37F899FD029E52" bold="true" box="[151,264,1803,1827]" pageId="17" pageNumber="18">PLATE 4.</emphasis>
|
||
Dorsal & ventral view of ZFMK 47860 and ZFMK 49036 (
|
||
<typeStatus id="54F0E2DC6413DF7AFC28F89EFFF89E52" box="[904,1010,1804,1827]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B27FD6413DF7AFBB5F89EF8A59E52" box="[1045,1199,1804,1827]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="yemenensis">
|
||
<emphasis id="B93F806C6413DF7AFBB5F89EF8A59E52" box="[1045,1199,1804,1827]" italics="true" pageId="17" pageNumber="18">U. yemenensis</emphasis>
|
||
</taxonomicName>
|
||
) Photos: T. Wilms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3510FF56410DF79FF37FB3DF90A9DB7" box="[151,1280,1199,1223]" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="8BF45C7E6410DF79FF37FB3DF90A9DB7" blockId="18.[151,1280,1199,1223]" box="[151,1280,1199,1223]" pageId="18" pageNumber="19">
|
||
<emphasis id="B93F806C6410DF79FF37FB3DFD029DB6" bold="true" box="[151,264,1199,1223]" pageId="18" pageNumber="19">PLATE 5.</emphasis>
|
||
<typeStatus id="54F0E2DC6410DF79FEAFFB3DFD789DB7" box="[271,370,1199,1222]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B27FD6410DF79FE36FB22FE999DB6" box="[406,659,1200,1223]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6410DF79FE36FB22FE999DB6" box="[406,659,1200,1223]" italics="true" pageId="18" pageNumber="19">U. yemenensis shobraki</emphasis>
|
||
</taxonomicName>
|
||
(ZFMK 48681), dorsal & ventral view. Photos: T. Wilms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF340CF66410DF79FF37F892FDB09E29" httpUri="https://zenodo.org/record/175334/files/figure.png" pageId="18" pageNumber="19" targetBox="[415,1171,1267,1760]" targetPageId="18">
|
||
<paragraph id="8BF45C7E6410DF79FF37F892FDB09E29" blockId="18.[151,1435,1792,1880]" pageId="18" pageNumber="19">
|
||
<emphasis id="B93F806C6410DF79FF37F892FD139E69" bold="true" box="[151,281,1792,1816]" pageId="18" pageNumber="19">FIGURE 4.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C4B27FD6410DF79FE64F893FEBD9E69" box="[452,695,1793,1816]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="yemenensis">
|
||
<emphasis id="B93F806C6410DF79FE64F893FEBD9E69" box="[452,695,1793,1816]" italics="true" pageId="18" pageNumber="19">Uromastyx yemenensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4B27FD6410DF79FD65F891FF419E69" box="[709,843,1795,1816]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="shobraki">
|
||
<emphasis id="B93F806C6410DF79FD65F891FF419E69" box="[709,843,1795,1816]" italics="true" pageId="18" pageNumber="19">U. y. shobraki</emphasis>
|
||
</taxonomicName>
|
||
: 1 AlMukha; 2 Taizz; 3 Zabid.
|
||
<taxonomicName id="4C4B27FD6410DF79FB60F891F96A9E69" box="[1216,1376,1795,1816]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="yemenensis" subSpecies="yemenensis">
|
||
<emphasis id="B93F806C6410DF79FB60F891F96A9E69" box="[1216,1376,1795,1816]" italics="true" pageId="18" pageNumber="19">U. y. yemenensis</emphasis>
|
||
</taxonomicName>
|
||
: 1 Lahij; 2 50 mls of Aden; 3 Zinjibar; 4 AbianArea / AbianMountains; 5 Lawdar.
|
||
<taxonomicName id="4C4B27FD6410DF79FBADF8B1F8569E49" box="[1037,1116,1827,1848]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="benti">
|
||
<emphasis id="B93F806C6410DF79FBADF8B1F8569E49" box="[1037,1116,1827,1848]" italics="true" pageId="18" pageNumber="19">U. benti</emphasis>
|
||
</taxonomicName>
|
||
: 1 Azzan; 2 AlMukalla; 3 Vicinity of Mirbat (Oman).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3510FF56410DF78FF66F81AFE8899AB" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8BF45C7E6410DF78FF66F81AFE8899AB" blockId="18.[151,1439,1928,2035]" lastBlockId="19.[151,1433,152,218]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
<emphasis id="B93F806C6410DF79FF66F81AFD6D9ED3" bold="true" box="[198,359,1928,1954]" pageId="18" pageNumber="19">Distribution.</emphasis>
|
||
<taxonomicName id="4C4B27FD6410DF79FECFF81BFE779ED3" box="[367,637,1929,1954]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="yemenensis">
|
||
<emphasis id="B93F806C6410DF79FECFF81BFE779ED3" box="[367,637,1929,1954]" italics="true" pageId="18" pageNumber="19">Uromastyx yemenensis</emphasis>
|
||
</taxonomicName>
|
||
is distributed in the southwestern parts of the
|
||
<collectingCountry id="F35C1CEE6410DF79FB10F81BF9929ED2" box="[1200,1432,1929,1955]" name="Yemen" pageId="18" pageNumber="19">Republic of Yemen</collectingCountry>
|
||
, with the range extending from the vicinity of Lawdar west to the Red Sea and from Zabid south to the Gulf of Aden. For the distribution areas of the subspecies of
|
||
<taxonomicName id="4C4B27FD6410DF79FCAEF84BFFB39E83" box="[782,953,2009,2034]" class="Reptilia" family="Agamidae" genus="Uromastyx" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="yemenensis">
|
||
<emphasis id="B93F806C6410DF79FCAEF84BFFB39E83" box="[782,953,2009,2034]" italics="true" pageId="18" pageNumber="19">U. yemenensis</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<figureCitation id="137040FB6410DF79FC4FF84BF8339E82" box="[1007,1081,2009,2035]" captionStart="FIGURE 4" captionStartId="18.[151,255,1792,1816]" captionTargetBox="[415,1171,1267,1760]" captionTargetId="figure@18.[415,1171,1267,1789]" captionTargetPageId="18" captionText="FIGURE 4. Distribution of Uromastyx yemenensis. U. y. shobraki: 1 AlMukha; 2 Taizz; 3 Zabid. U. y. yemenensis: 1 Lahij; 2 50 mls of Aden; 3 Zinjibar; 4 AbianArea / AbianMountains; 5 Lawdar. U. benti: 1 Azzan; 2 AlMukalla; 3 Vicinity of Mirbat (Oman)." httpUri="https://zenodo.org/record/175334/files/figure.png" pageId="18" pageNumber="19">Fig. 4</figureCitation>
|
||
. The areas of both subspecies are separated by a series of mountains and mountain ranges (example: Jebel Manar, Jebel Sawraq, Jebel Sabir, Jebel Dawran) with heights up to
|
||
<quantity id="4CB3F19B6411DF78FD82FF52FE7499AB" box="[546,638,192,218]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="19" pageNumber="20" unit="m" value="3200.0">3200 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |