166 lines
20 KiB
XML
166 lines
20 KiB
XML
<document id="C884E1F49F1C50778AD2965E20640E62" ID-DOI="10.11646/zootaxa.4790.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3891303" ID-ZooBank="F011C13E-8E90-45D8-ABA3-B7D58582648A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591959490517" checkinUser="plazi" docAuthor="Lambkin, Kevin J." docDate="2020" docId="03E187D0BA7C5128FF006179F3C5FDAF" docLanguage="en" docName="zootaxa.4790.3.7.pdf" docOrigin="Zootaxa 4790 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesocixiodes Tillyard 1922" docType="treatment" docVersion="6" lastPageNumber="534" masterDocId="FFD8FFA8BA745121FF976716F317FFCC" masterDocTitle="Revision of the Hylicellidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Hylicelloidea)" masterLastPageNumber="539" masterPageNumber="525" pageNumber="533" updateTime="1698774607034" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7C892E5C7EFEE2286105AE4852BEABB4">Revision of the Hylicellidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Hylicelloidea)</mods:title>
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<mods:namePart id="4233B1111DB6E24F49196210B53B997E">Lambkin, Kevin J.</mods:namePart>
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<mods:title id="47B15CC3AE91F1FD0293EF9B1298CFE2">Zootaxa</mods:title>
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<mods:date id="A6F27D47D6B7D7E0D9B24058A611C26B">2020</mods:date>
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<mods:number id="DD8104F99077B08460BDF715C4292D9A">2020-06-12</mods:number>
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<subSubSection id="C352654DBA7C5129FF006179F2E6F946" box="[151,497,1647,1674]" pageId="8" pageNumber="533" type="nomenclature">
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<paragraph id="8BF736C6BA7C5129FF006179F2E6F946" blockId="8.[151,497,1647,1674]" box="[151,497,1647,1674]" pageId="8" pageNumber="533">
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<heading id="D0BF81AABA7C5129FF006179F2E6F946" bold="true" box="[151,497,1647,1674]" fontSize="11" level="1" pageId="8" pageNumber="533" reason="1">
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<taxonomicName id="4C484D45BA7C5129FF006179F2E6F946" ID-CoL="8HM9C" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[151,497,1647,1674]" class="Insecta" family="Hylicellidae" genus="Mesocixiodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="B93CEAD4BA7C5129FF006179F2E6F946" bold="true" box="[151,497,1647,1674]" pageId="8" pageNumber="533">
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<emphasis id="B93CEAD4BA7C5129FF006179F22DF946" bold="true" box="[151,314,1647,1674]" italics="true" pageId="8" pageNumber="533">Mesocixiodes</emphasis>
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<bibRefCitation id="EFD94B37BA7C5129FED66166F2E6F946" author="Tillyard, R. J." box="[321,497,1648,1674]" pageId="8" pageNumber="533" pagination="447 - 470" refId="ref8374" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. Proceedings of the Linnean Society of New South Wales, 47, 447 - 470." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352654DBA7C5129FF0061A2F2E5F900" box="[151,498,1715,1740]" pageId="8" pageNumber="533" type="reference_group">
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<paragraph id="8BF736C6BA7C5129FF0061A2F2E5F900" blockId="8.[151,971,1715,1771]" box="[151,498,1715,1740]" pageId="8" pageNumber="533">
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<treatmentCitationGroup id="AB5811E8BA7C5129FF0061A2F2E5F900" box="[151,498,1715,1740]" pageId="8" pageNumber="533">
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<treatmentCitation id="0AE910D7BA7C5129FF0061A2F2F9F900" author="Tillyard, R. J." box="[151,494,1715,1740]" page="462" pageId="8" pageNumber="533" year="1922">
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<taxonomicName id="4C484D45BA7C5129FF0061A2F2F9F900" ID-CoL="8HM9C" authority="Tillyard, 1922: 462" authorityName="Tillyard" authorityPageNumber="462" authorityYear="1922" box="[151,494,1715,1740]" class="Insecta" family="Hylicellidae" genus="Mesocixiodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="B93CEAD4BA7C5129FF0061A2F236F900" box="[151,289,1716,1740]" italics="true" pageId="8" pageNumber="533">Mesocixiodes</emphasis>
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<bibRefCitation id="EFD94B37BA7C5129FEB061A5F2F9F900" author="Tillyard, R. J." box="[295,494,1715,1740]" pageId="8" pageNumber="533" pagination="447 - 470" refId="ref8374" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. Proceedings of the Linnean Society of New South Wales, 47, 447 - 470." type="journal article" year="1922">Tillyard, 1922: 462</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C352654DBA7C5128FF0061C5F3C5FDAF" lastPageId="9" lastPageNumber="534" pageId="8" pageNumber="533" type="type_taxon">
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<paragraph id="8BF736C6BA7C5129FF0061C5F0DCF927" blockId="8.[151,971,1715,1771]" box="[151,971,1746,1771]" pageId="8" pageNumber="533">
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<emphasis id="B93CEAD4BA7C5129FF0061C5F234F927" bold="true" box="[151,291,1746,1771]" inLineHeading="true" pageId="8" pageNumber="533" reason="1">
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<typeStatus id="54F38864BA7C5129FF0061C5F3DBF927" box="[151,204,1747,1771]" pageId="8" pageNumber="533">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C484D45BA7C5129FEBD61C5F1C7F927" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[298,720,1746,1771]" class="Insecta" family="Hylicellidae" genus="Mesocixiodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="533" phylum="Arthropoda" rank="species" species="termioneura">
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<emphasis id="B93CEAD4BA7C5129FEBD61C5F120F926" box="[298,567,1747,1771]" italics="true" pageId="8" pageNumber="533">Mesocixiodes termioneura</emphasis>
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<bibRefCitation id="EFD94B37BA7C5129FDAA61C4F1C7F927" author="Tillyard, R. J." box="[573,720,1746,1771]" pageId="8" pageNumber="533" pagination="447 - 470" refId="ref8374" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. Proceedings of the Linnean Society of New South Wales, 47, 447 - 470." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="8BF736C6BA7C5128FF00600FF640FE47" blockId="8.[151,1436,1816,2029]" lastBlockId="9.[151,1437,152,611]" lastPageId="9" lastPageNumber="534" pageId="8" pageNumber="533">
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<emphasis id="B93CEAD4BA7C5129FF00600FF290F8FF" bold="true" box="[151,391,1816,1843]" pageId="8" pageNumber="533">Emended diagnosis.</emphasis>
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<materialsCitation id="3B203C9BBA7C5128FE18600CF643FE47" collectionCode="RK" lastPageId="9" lastPageNumber="534" pageId="8" pageNumber="533" specimenCode="RK1, RK2" specimenCount="1" typeStatus="holotype">
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Tegmen
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<emphasis id="B93CEAD4BA7C5129FE66600CF2EAF8FF" box="[497,509,1818,1843]" italics="true" pageId="8" pageNumber="533">c</emphasis>
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. 3.5 times longer than wide, finely to somewhat coarsely punctate and tending to irregularly transversely rugose, extreme apical area possibly less so; costal margin broadly rounded, apex narrowly rounded; peripheral membrane of even width; costal space
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<emphasis id="B93CEAD4BA7C5129FCAE6074F052F8B7" box="[825,837,1890,1915]" italics="true" pageId="8" pageNumber="533">c</emphasis>
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. 3 times as wide as medial cell; R and M separating near apex of basal cell; basal cell quite broad proximally, slightly tapering and closed distally by a fusion for a short length of M and CuA; primary fork of R just before tegmen length, far basal to primary fork of M which is well beyond primary fork of RA and just proximal to very shallow CuA fork; RA
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<subScript id="17CC3483BA7C5129FC5560C9F0DDF821" attach="left" box="[962,970,2015,2029]" fontSize="6" pageId="8" pageNumber="533">1</subScript>
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simple, obliquely angled towards anterior margin; RA
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<subScript id="17CC3483BA7D5128FE8B67BCF233FF74" attach="left" box="[284,292,170,184]" fontSize="6" pageId="9" pageNumber="534">2</subScript>
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stem directed apically, more or less parallel with costal margin, with 4 or 5 long, sub-equal pectinate branches directed antero-apically; RA
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<subScript id="17CC3483BA7D5128FDA967D8F151FF10" attach="left" box="[574,582,206,220]" fontSize="6" pageId="9" pageNumber="534">1</subScript>
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+ RA
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<subScript id="17CC3483BA7D5128FD1F67D8F187FF10" attach="left" box="[648,656,206,220]" fontSize="6" pageId="9" pageNumber="534">2</subScript>
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field occupying entire antero-apical margin; RP deeply forked; one
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<emphasis id="B93CEAD4BA7D5128FA1467A8F681FF1B" box="[1411,1430,190,215]" italics="true" pageId="9" pageNumber="534">ir</emphasis>
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, running from RA
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<subScript id="17CC3483BA7D5128FECD67E4F275FECC" attach="left" box="[346,354,242,256]" fontSize="6" pageId="9" pageNumber="534">2</subScript>
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to RP
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<subScript id="17CC3483BA7D5128FE3E67E4F2A6FECC" attach="left" box="[425,433,242,256]" fontSize="6" pageId="9" pageNumber="534">1</subScript>
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; RP
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<subScript id="17CC3483BA7D5128FE7467E4F2FCFECC" attach="left" box="[483,491,242,256]" fontSize="6" pageId="9" pageNumber="534">2</subScript>
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fused with M
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<subScript id="17CC3483BA7D5128FD1F67E4F1B5FECC" attach="left" box="[648,674,242,256]" fontSize="6" pageId="9" pageNumber="534">1+2</subScript>
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for a short length (
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<specimenCode id="DBEE9EBDBA7D5128FCE067F7F0BBFF37" box="[887,940,225,251]" collectionCode="RK" pageId="9" pageNumber="534">RK1</specimenCode>
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and
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<specimenCode id="DBEE9EBDBA7D5128FC7367F7F70AFF37" box="[996,1053,225,251]" collectionCode="RK" pageId="9" pageNumber="534">RK2</specimenCode>
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), or separate, with a single
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<emphasis id="B93CEAD4BA7D5128FAC767F4F66FFF37" box="[1360,1400,226,251]" italics="true" pageId="9" pageNumber="534">r-m</emphasis>
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, in line with
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<emphasis id="B93CEAD4BA7D5128FF686610F205FED3" box="[255,274,262,287]" italics="true" pageId="9" pageNumber="534">ir</emphasis>
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(
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<typeStatus id="54F38864BA7D5128FE886613F290FEEC" box="[287,391,261,288]" pageId="9" pageNumber="534" type="holotype">holotype</typeStatus>
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); M with 4 or 5 branches;
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<emphasis id="B93CEAD4BA7D5128FD326610F1D6FED3" box="[677,705,262,287]" italics="true" pageId="9" pageNumber="534">im</emphasis>
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between M
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<subScript id="17CC3483BA7D5128FCD46600F05CFEE8" attach="left" box="[835,843,278,292]" fontSize="6" pageId="9" pageNumber="534">2</subScript>
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and M
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<subScript id="17CC3483BA7D5128FC0F6600F0B7FEE8" attach="left" box="[920,928,278,292]" fontSize="6" pageId="9" pageNumber="534">3</subScript>
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or M
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<subScript id="17CC3483BA7D5128FC4B6600F0E1FEE8" attach="left" box="[988,1014,278,292]" fontSize="6" pageId="9" pageNumber="534">3+4</subScript>
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;intra-medial cell quite broad but very small,
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<emphasis id="B93CEAD4BA7D5128FF74663CF3F8FE8F" box="[227,239,298,323]" italics="true" pageId="9" pageNumber="534">c</emphasis>
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. 2 times longer than wide, occupying a very small portion of post-nodal area; CuA branches short, CuA
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<subScript id="17CC3483BA7D5128FA03662CF68BFE84" attach="left" box="[1428,1436,314,328]" fontSize="6" pageId="9" pageNumber="534">1</subScript>
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strongly angled at
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<emphasis id="B93CEAD4BA7D5128FEF36658F2BDFEAB" box="[356,426,334,359]" italics="true" pageId="9" pageNumber="534">m-cua</emphasis>
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which joins M
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<subScript id="17CC3483BA7D5128FDD86648F17EFEA0" attach="left" box="[591,617,350,364]" fontSize="6" pageId="9" pageNumber="534">3+4</subScript>
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around midpoint of intra-medial cell; crossvein field tending to alignment at level of claval apex, thus creating a fairly even series of long, sub-equal apical cells; clavus not preserved
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8BF736C6BA7D5128FF506683F3C5FDAF" blockId="9.[151,1437,152,611]" pageId="9" pageNumber="534">
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<emphasis id="B93CEAD4BA7D5128FF506683F206FE63" bold="true" box="[199,273,405,431]" pageId="9" pageNumber="534">Notes.</emphasis>
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<taxonomicName id="4C484D45BA7D5128FE8F6683F2A7FE63" authorityName="Tillyard" authorityYear="1922" box="[280,432,405,431]" class="Insecta" family="Hylicellidae" genus="Mesocixiodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="534" phylum="Arthropoda" rank="genus">
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<emphasis id="B93CEAD4BA7D5128FE8F6683F2A7FE63" box="[280,432,405,431]" italics="true" pageId="9" pageNumber="534">Mesocixiodes</emphasis>
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||
</taxonomicName>
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||
was placed by
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||
<bibRefCitation id="EFD94B37BA7D5128FDCB6683F016FE63" author="Tillyard, R. J." box="[604,769,405,431]" pageId="9" pageNumber="534" pagination="447 - 470" refId="ref8374" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. Proceedings of the Linnean Society of New South Wales, 47, 447 - 470." type="journal article" year="1922">Tillyard (1922</bibRefCitation>
|
||
,
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||
<bibRefCitation id="EFD94B37BA7D5128FC9B6683F05FFE7C" author="Tillyard, R. J." box="[780,840,405,432]" pageId="9" pageNumber="534" pagination="265 - 272" refId="ref8422" refString="Tillyard, R. J. (1926) The Rhaetic " Crane-Flies from South America not Diptera but Homoptera. American Journal of Science, 11, 265 - 272. https: // doi. org / 10.2475 / ajs. s 5 - 11.63.265" type="journal article" year="1926">1926</bibRefCitation>
|
||
),
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||
<bibRefCitation id="EFD94B37BA7D5128FCCC6683F79CFE7C" author="Becker-Migdisova, E. E." box="[859,1163,405,432]" pageId="9" pageNumber="534" pagination="216 - 317" refId="ref6742" refString="Becker-Migdisova, E. E. (1962 a) Superorder Rhynchota. Insects with proboscis. In: Rohdendorf, B. B. (Ed.), Fundamentals of Paleontology. Vol. 9. Arthropoda, Tracheata, Chelicerata. Akademiya Nauk SSSR, Moscow, pp. 216 - 317. [in Russian; English translation 1991, Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C.]" type="journal article" year="1962">Becker-Migdisova (1962a)</bibRefCitation>
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||
, and
|
||
<bibRefCitation id="EFD94B37BA7D5128FB516683F628FE63" author="Jell, P. A." box="[1222,1343,405,431]" pageId="9" pageNumber="534" pagination="1 - 124" refId="ref7267" refString="Jell, P. A. (2004) The fossil insects of Australia. Memoirs of the Queensland Museum, 50, 1 - 124." type="journal article" year="2004">Jell (2004)</bibRefCitation>
|
||
in
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||
<taxonomicName id="4C484D45BA7D5128FAF46683F27DFE18" ID-CoL="89Z" authority="(Fulgoroidea)" baseAuthorityName="Fulgoroidea" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="534" phylum="Arthropoda" rank="family">Cixiidae (Fulgoroidea)</taxonomicName>
|
||
, by
|
||
<bibRefCitation id="EFD94B37BA7D5128FE0F66AFF173FE1F" author="Handlirsch, A." box="[408,612,441,467]" pageId="9" pageNumber="534" pagination="1 - 240" refId="ref7222" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Teil. Annalen des naturhistorischen Museums in Wien, 49, 1 - 240." type="journal article" year="1939">Handlirsch (1939)</bibRefCitation>
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||
in
|
||
<taxonomicName id="4C484D45BA7D5128FD1F66AFF013FE1F" ID-CoL="623TM" box="[648,772,441,467]" class="Insecta" family="Fulgoridae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="534" phylum="Arthropoda" rank="family">Fulgoridae</taxonomicName>
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, by
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<bibRefCitation id="EFD94B37BA7D5128FCA466AFF0DEFE1F" author="Evans, J. W." box="[819,969,440,467]" pageId="9" pageNumber="534" refId="ref6976" refString="Evans, J. W. (1956) Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165 - 158. https: // doi. org / 10.1071 / ZO 9560165" type="book" year="1956">Evans (1956)</bibRefCitation>
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in
|
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<taxonomicName id="4C484D45BA7D5128FC7A66AFF784FE1F" ID-CoL="8HBQF" authorityName="Evans" authorityYear="1956" box="[1005,1171,441,467]" class="Insecta" family="Chiliocyclidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="534" phylum="Arthropoda" rank="family">Chiliocyclidae</taxonomicName>
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, and was considered by
|
||
<bibRefCitation id="EFD94B37BA7D5128FF0066CBF24EFE3B" author="Carpenter, F. M." box="[151,345,477,503]" pageId="9" pageNumber="534" pagination="1 - 277" refId="ref6857" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vol. 3. The Geological Society of America Inc. Boulder, Colorado and The University of Kansas, Lawrence, Kansas, pp. i-xxi + 1 - 277." type="book chapter" year="1992">Carpenter (1992)</bibRefCitation>
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||
as ‘Family uncertain’. It was transferred to
|
||
<taxonomicName id="4C484D45BA7D5128FCC466CBF0CBFE3B" ID-CoL="8HBQP" authorityName="Evans" authorityYear="1956" box="[851,988,477,503]" class="Insecta" family="Hylicellidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="534" phylum="Arthropoda" rank="family">Hylicellidae</taxonomicName>
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||
by
|
||
<bibRefCitation id="EFD94B37BA7D5128FB9E66CBF7DEFE3B" author="Hamilton, K. G. A." box="[1033,1225,477,503]" pageId="9" pageNumber="534" pagination="423 - 430" refId="ref7164" refString="Hamilton, K. G. A. (1992) Lower Cretaceous Homoptera from the Koonwarra fossil bed in Australia, with a new Superfamily and synopsis of Mesozoic Homoptera. Annals of the Entomological Society of America, 85, 423 - 430. https: // doi. org / 10.1093 / aese / 85.4.423" type="journal article" year="1992">Hamilton (1992)</bibRefCitation>
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. It differs from all other hylicellids in the very late forking of M, well distal of that of RA, the very small intra-medial cell, and the tendency to alignment of the crossvein field at the claval apex to create a fairly even series of long, sub-equal apical cells.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |