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<document id="BBF92E18001254BA47734955B9F24A04" ID-DOI="10.26515/rzsi/v121/i3/2021/158157" ID-ISSN="2581-8686" ID-Zenodo-Dep="13184780" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722610335945" checkinUser="felipe" docAuthor="Shangningam, Bungdon &amp; Kosygin, Laishram" docDate="2021" docId="03E18799FFE8CB32ED3BE62E4B667904" docLanguage="en" docName="RecZoolSurvIndia.121.3.319-324.pdf" docOrigin="Records of the Zoological Survey of India 121 (3)" docStyle="DocumentStyle:30C78A88DFAFB68AE5A0604D5A6B2FFB.3:RecZoolSurvIndia.2017-.journal_article" docStyleId="30C78A88DFAFB68AE5A0604D5A6B2FFB" docStyleName="RecZoolSurvIndia.2017-.journal_article" docStyleVersion="3" docTitle="Psilorhynchus magnaoculus Shangningam &amp; Kosygin, 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="322" masterDocId="FFD8FFE1FFE8CB31EE65E06A4F027C5A" masterDocTitle="Psilorhynchus magnaoculus, a new species of torrent minnow (Teleostei: Psilorhynchidae) from Myanmar" masterLastPageNumber="324" masterPageNumber="319" pageNumber="319" updateTime="1722660594501" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D3A72C4E2CE7F76C9867DE709998E870" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="60E8952FE8E1360D596B6C1AC4AF685C">
<mods:title id="F0F152DB75E0ED98AB2B5866494897C6">Psilorhynchus magnaoculus, a new species of torrent minnow (Teleostei: Psilorhynchidae) from Myanmar</mods:title>
</mods:titleInfo>
<mods:name id="A9B6C2430307AE368BC1C446A5990CE5" type="personal">
<mods:role id="18E39BD7B32E073ED35D2920A2875E77">
<mods:roleTerm id="DF3F7346B6BA7B9FC3C75ABDD9873B10">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A60D0C3BC1C403E07A26CA1F8E6DE83">Shangningam, Bungdon</mods:namePart>
</mods:name>
<mods:name id="96B909647C3FDAE8EABB1115402B6DCC" type="personal">
<mods:role id="04574799B29315ECC8B3C7719C2AB303">
<mods:roleTerm id="20D5A7DA629D81842BD9966C40E98C3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2968E8F8E71BE993DA0D1E779D8D2166">Kosygin, Laishram</mods:namePart>
</mods:name>
<mods:typeOfResource id="3011F2BFE934BB6B19ED877A0CCC9EA3">text</mods:typeOfResource>
<mods:relatedItem id="3F3A60CA08465497CEB02C78B2AEC172" type="host">
<mods:titleInfo id="0F6352C1AD3998E3A571EED38EA300CE">
<mods:title id="42A2FE5B3E7FFB2072D2CA5CE71B7925">Records of the Zoological Survey of India</mods:title>
</mods:titleInfo>
<mods:part id="0215E2D9AC1559560C279A59DA430EA7">
<mods:date id="898808E31538AA615BA5FF1B0D856965">2021</mods:date>
<mods:detail id="A5F36CEC1BA4803F2B3CFAEC2BEA883A" type="volume">
<mods:number id="6B938D875281CF6E26172C5F2192A883">121</mods:number>
</mods:detail>
<mods:detail id="70B61A6619624132DCEE8345C8C52A03" type="issue">
<mods:number id="AA94EBCC1DFF93A82D9CECE00B5EE798">3</mods:number>
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<mods:start id="13BEA57C184C2BBDC3F9796E75955FAE">319</mods:start>
<mods:end id="40D1DA2CFE95617A5CF07B1058AFEE6D">324</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="1298370443B7263D330FA9B2A732A3C1">journal article</mods:classification>
<mods:identifier id="03C3CDB263E96B84B11907DF4B903389" type="DOI">10.26515/rzsi/v121/i3/2021/158157</mods:identifier>
<mods:identifier id="2EBAFD44DF66C6C011B8E5A591ED3497" type="ISSN">2581-8686</mods:identifier>
<mods:identifier id="3282C3F321BAEE91BF77FF530250D4BA" type="Zenodo-Dep">13184780</mods:identifier>
</mods:mods>
<treatment id="03E18799FFE8CB32ED3BE62E4B667904" LSID="urn:lsid:plazi:treatment:03E18799FFE8CB32ED3BE62E4B667904" httpUri="http://treatment.plazi.org/id/03E18799FFE8CB32ED3BE62E4B667904" lastPageId="3" lastPageNumber="322" pageId="0" pageNumber="319">
<subSubSection id="C3526504FFE8CB31ED3BE62E4B1B7AEE" pageId="0" pageNumber="319" type="nomenclature">
<paragraph id="8BF7368FFFE8CB31ED3BE62E4AB07A05" blockId="0.[862,1458,1604,1631]" box="[862,1458,1604,1631]" pageId="0" pageNumber="319">
<heading id="D0BF81E3FFE8CB31ED3BE62E4AB07A05" box="[862,1458,1604,1631]" fontSize="11" level="2" pageId="0" pageNumber="319" reason="4">
<taxonomicName id="4C484D0CFFE8CB31ED3BE62E4B9C7A05" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[862,1182,1604,1631]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="319" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFE8CB31ED3BE62E4B9C7A05" bold="true" box="[862,1182,1604,1631]" italics="true" pageId="0" pageNumber="319">Psilorhynchus magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFE8CB31EAC6E62F4BF57A05" box="[1187,1271,1605,1631]" pageId="0" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13732A0AFFE8CB31EB61E62E4A6B7A05" box="[1284,1385,1604,1631]" captionStart="Figure 1" captionStartId="1.[160,236,1814,1840]" captionTargetBox="[160,797,1600,1757]" captionTargetId="figure-578@1.[160,797,1600,1757]" captionTargetPageId="1" captionText="Figure 1. P. magnaoculus sp. nov., holotype, ZSI FF 8211, 47.5 mm SL; a. dorsal b. lateral c. ventral view." figureDoi="http://doi.org/10.5281/zenodo.13184782" httpUri="https://zenodo.org/record/13184782/files/figure.png" pageId="0" pageNumber="319">Figures 1</figureCitation>
a-c, 2)
</heading>
</paragraph>
<paragraph id="8BF7368FFFE8CB31ED3BE61C4B1B7AEE" blockId="0.[862,1499,1654,1716]" pageId="0" pageNumber="319">
<uri id="FFD93A8DFFE8CB31ED3BE61C4B1B7AEE" pageId="0" pageNumber="319">
urn:lsid:zoobank.org:act:
<uuid id="FFEE0C5AFFE8CB31EA0AE61C4B1B7AEE" pageId="0" pageNumber="319">A7429508-AD0F-4CFD-8D43- C88B4417C3CD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3526504FFE8CB32ED3BE6A64B667904" lastPageId="3" lastPageNumber="322" pageId="0" pageNumber="319" type="materials_examined">
<paragraph id="8BF7368FFFE8CB30ED3BE6A64C1F7D4E" blockId="0.[862,1499,1739,1802]" lastBlockId="1.[159,797,214,456]" lastPageId="1" lastPageNumber="320" pageId="0" pageNumber="319">
<emphasis id="B93CEA9DFFE8CB31ED3BE6A64B017ABF" box="[862,1027,1739,1765]" italics="true" pageId="0" pageNumber="319">Type material</emphasis>
:
<materialsCitation id="3B203CD2FFE8CB30EA79E6A14EF77D4E" collectingDate="2018-12-28" country="Myanmar" county="Tamu District" lastPageId="1" lastPageNumber="320" latitude="24.226946" location="Tamu" longLatPrecision="21" longitude="94.31833" municipality="Yu River" pageId="0" pageNumber="319" specimenCount="1" stateProvince="Sagaing" typeStatus="holotype">
<typeStatus id="54F3882DFFE8CB31EA79E6A14B857ABF" box="[1052,1159,1739,1765]" pageId="0" pageNumber="319" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F35F761FFFE8CB31EAF8E6A64A097ABC" box="[1181,1291,1740,1766]" name="Myanmar" pageId="0" pageNumber="319">Myanmar</collectingCountry>
,
<collectingRegion id="498CF86DFFE8CB31EB46E6A14AD47ABF" box="[1315,1494,1739,1765]" country="Myanmar" name="Sagaing" pageId="0" pageNumber="319">Sagaing region</collectingRegion>
,
<collectingCounty id="62964E03FFE8CB31ED3BE69A4B0A7B53" box="[862,1032,1775,1802]" pageId="0" pageNumber="319">Tamu District</collectingCounty>
,
<collectingMunicipality id="6B93ACF5FFE8CB31EA78E69A4B8B7B53" box="[1053,1161,1775,1802]" pageId="0" pageNumber="319">Yu River</collectingMunicipality>
at
<location id="8E976054FFE8CB31EAA4E69A4A037B50" LSID="urn:lsid:plazi:treatment:03E18799FFE8CB32ED3BE62E4B667904:8E976054FFE8CB31EAA4E69A4A037B50" box="[1217,1281,1776,1802]" country="Myanmar" county="Tamu District" latitude="24.226946" longLatPrecision="21" longitude="94.31833" municipality="Yu River" name="Tamu" pageId="0" pageNumber="319" stateProvince="Sagaing">Tamu</location>
(
<location id="8E976054FFE8CB31EB7FE6854AD97B53" LSID="urn:lsid:plazi:treatment:03E18799FFE8CB32ED3BE62E4B667904:8E976054FFE8CB31EB7FE6854AD97B53" box="[1306,1499,1775,1801]" country="Myanmar" county="Tamu District" latitude="24.226946" longLatPrecision="21" longitude="94.31833" municipality="Yu River" name="Chindwin River" pageId="0" pageNumber="319" stateProvince="Sagaing">Chindwin River</location>
basin),
<geoCoordinate id="EE7C5048FFE9CB30EE9EE0BC4E7E7CAA" box="[251,380,214,240]" degrees="24" direction="north" minutes="13" orientation="latitude" pageId="1" pageNumber="320" precision="15" seconds="37" value="24.226946">24°13’37’’N</geoCoordinate>
<geoCoordinate id="EE7C5048FFE9CB30EFEFE0BC4D0A7CAA" box="[394,520,214,240]" degrees="94" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="320" precision="15" seconds="06" value="94.31833">94°19’06’’E</geoCoordinate>
,
<date id="FFF6104FFFE9CB30EC7EE0BC4D9C7CAA" box="[539,670,214,240]" pageId="1" pageNumber="320" value="2018-12-28">
<collectingDate id="EFB2E9A7FFE9CB30EC7EE0BC4D9C7CAA" box="[539,670,214,240]" pageId="1" pageNumber="320" value="2018-12-28">28-xii-2018</collectingDate>
</date>
, collector? Shangningam (ZSI FF 8211)
</materialsCitation>
.
<typeStatus id="54F3882DFFE9CB30EC62E0904D747D4E" box="[519,630,250,276]" pageId="1" pageNumber="320" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4EFD06FFE9CB30ECE2E0904C1B7D4E" box="[647,793,250,276]" count="3" pageId="1" pageNumber="320" type="generic">3 specimens</specimenCount>
,
</paragraph>
<paragraph id="8BF7368FFFE9CB30EEC5E1744E177D92" blockId="1.[159,797,214,456]" pageId="1" pageNumber="320">
32.0–
<quantity id="4CB09B6AFFE9CB30EEB8E1744E3D7D62" box="[221,319,286,312]" metricMagnitude="-2" metricUnit="m" metricValue="4.029999999999999" pageId="1" pageNumber="320" unit="mm" value="40.3">40.3 mm</quantity>
SL, same locality and label data as
<typeStatus id="54F3882DFFE9CB30ECDEE1744C1F7D62" box="[699,797,286,312]" pageId="1" pageNumber="320" type="holotype">holotype</typeStatus>
(ZSI FF 8212). The specimens bear a red printed label: “
<taxonomicName id="4C484D0CFFE9CB30EECEE10C4EE97DDA" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[171,491,358,384]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="320" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFE9CB30EECEE10C4EE97DDA" box="[171,491,358,384]" italics="true" pageId="1" pageNumber="320">Psilorhynchus magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFE9CB30EC66E10C4D687DDA" box="[515,618,358,384]" pageId="1" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="54F3882DFFE9CB30ECE0E10C4DEF7DDA" box="[645,749,358,384]" pageId="1" pageNumber="320" type="holotype">Holotype</typeStatus>
or
<typeStatus id="54F3882DFFE9CB30EEC5E1E04E067DFE" box="[160,260,394,420]" pageId="1" pageNumber="320" type="paratype">Paratype</typeStatus>
, Bungdon Shangningam* &amp; Laishram Kosygin, Det. 2021.
</paragraph>
<paragraph id="8BF7368FFFE9CB30EEC5E18A4DBE7E47" blockId="1.[160,797,479,541]" pageId="1" pageNumber="320">
<materialsCitation id="3B203CD2FFE9CB30EEC5E18A4D0F7E47" country="India" location="Zoological Survey" pageId="1" pageNumber="320" specimenCount="2" typeStatus="holotype">
<emphasis id="B93CEA9DFFE9CB30EEC5E18A4E4E7DA3" box="[160,332,479,505]" italics="true" pageId="1" pageNumber="320">
<typeStatus id="54F3882DFFE9CB30EEC5E18A4FD67DA3" box="[160,212,480,505]" pageId="1" pageNumber="320">Type</typeStatus>
depository
</emphasis>
: The
<typeStatus id="54F3882DFFE9CB30EFEEE18A4EB97DA0" box="[395,443,480,506]" pageId="1" pageNumber="320">type</typeStatus>
specimens are deposited in the
<location id="8E976054FFE9CB30EEC4E2694E6E7E47" LSID="urn:lsid:plazi:treatment:03E18799FFE8CB32ED3BE62E4B667904:8E976054FFE9CB30EEC4E2694E6E7E47" box="[161,364,515,541]" country="India" name="Zoological Survey" pageId="1" pageNumber="320">Zoological Survey</location>
of
<collectingCountry id="F35F761FFFE9CB30EFEBE2694ECB7E47" box="[398,457,515,541]" name="India" pageId="1" pageNumber="320">India</collectingCountry>
(ZSI)
</materialsCitation>
, Kolkata,
<collectingCountry id="F35F761FFFE9CB30EC1FE2694DBA7E47" box="[634,696,515,541]" name="India" pageId="1" pageNumber="320">India</collectingCountry>
.
</paragraph>
<paragraph id="8BF7368FFFE9CB30EEC5E25F4E4D7F7C" blockId="1.[159,800,564,806]" pageId="1" pageNumber="320">
<emphasis id="B93CEA9DFFE9CB30EEC5E25F4E1C7E14" box="[160,286,565,590]" italics="true" pageId="1" pageNumber="320">Description</emphasis>
(
<typeStatus id="54F3882DFFE9CB30EF55E25E4E9D7E14" box="[304,415,564,590]" pageId="1" pageNumber="320" type="holotype">Holotype</typeStatus>
): Biometric data of
<typeStatus id="54F3882DFFE9CB30ECE2E25E4DEB7E14" box="[647,745,564,590]" pageId="1" pageNumber="320" type="holotype">holotype</typeStatus>
and
<typeStatus id="54F3882DFFE9CB30EEC5E2334E0E7E29" box="[160,268,601,627]" pageId="1" pageNumber="320" type="paratype">paratypes</typeStatus>
as in
<tableCitation id="C6CA0334FFE9CB30EF3AE2324EB77E29" box="[351,437,600,627]" captionStart="Table 1" captionStartId="2.[133,196,603,629]" captionText="Table 1. Biometric data of holotype and paratypes of P. magnaoculus sp. nov. (n=4)." pageId="1" pageNumber="320">Table 1</tableCitation>
. Body elongate, deep, greatest depth at dorsal-fin origin. Dorsal profile arched, rising gradually to dorsal-fin origin, sloping gently towards caudal peduncle. Ventral profile straight from lower jaw to anal-fin origin dorsally inclined from anal-fin origin to caudal-fin base.
</paragraph>
<paragraph id="8BF7368FFFE9CB30EEADE3544A487E03" blockId="1.[159,799,830,1036]" lastBlockId="1.[861,1501,215,1430]" pageId="1" pageNumber="320">Head large. Mouth inferior, relatively wide. Eye large, contributing to 42.6–43.6 percent of head length, dorsolateral located its diameter almost equal to snout length. Snout squarish anteriorly with a notch situated at the level of the ethmoid region anterior to nares, ventral surface bordered by deep longitudinal groove on each side. Rostral cap fused with upper lip, separated by shallow groove. Lower jaw covered by thick squarish cushion that can be folded backwards, cushion composed of two adnate tissue layers: anteriorly located deeper layer, lower lip, smooth, not continuous with upper lip around corner of mouth; and posteriorly located superficial layer densely papillated, thick, continuous with skin of isthmus. Superficial layer connected with rostral cap by narrow strip of skin around corner of mouth, extended posteriorly, broadened as ridged, slightly papillated skin fold at posterolateral-most corner of mouth.</paragraph>
<paragraph id="8BF7368FFFE9CB30EDE0E2094AB17853" blockId="1.[861,1501,215,1430]" pageId="1" pageNumber="320">Dorsal fin with ii.9 rays, high, tip rounded, posterior edge weakly concave. Anal fin with ii.6 rays, long, posterior edge concave, reaching base of caudal fin. Caudal fin weakly forked, upper lobe slightly longer than lower. Principal caudal-fin rays 10+9, procurrent rays 4 dorsally and 3 ventrally. Pelvic-fin rays ii.7. Pelvic-fin origin posterior to dorsal-fin origin, opposite to 3rd branched dorsal-fin ray. Pectoral with fin v, 10(1) or vi, 11(3) rays. Pectoral fin reaching horizontal through dorsal-fin origin, reaching to two scale rows anterior to pelvic-fin origin when depressed. Skin on ventral surface of unbranched pectoral and pelvic-fin rays thickened.</paragraph>
<paragraph id="8BF7368FFFE9CB30EDE0E4794CD179CC" blockId="1.[861,1501,215,1430]" pageId="1" pageNumber="320">
Scales cycloid, fairly large, thin and more or less regularly arranged on body. Scales along lateral line 31(1) or 32(3), plus 2(4) on base of caudal fin. Transverse scale rows from dorsal-fin origin to pelvic-fin origin 3.5/1/2, scale rows around caudal peduncle 10, predorsal scales 11(3)–12(1). Anus close to pelvic fin, position 2(1)–3(3) scales posterior to pelvic-fin origin. Scales between anus and anal-fin origin 9(1) or 10(3). Scales absent from midventral region between pectoral fins. Total number of vertebrae 33, comprised of 22 abdominal and 11 caudal (
<figureCitation id="13732A0AFFE9CB30ED00E5114CC479CC" box="[869,966,1403,1430]" captionStart="Figure 2" captionStartId="2.[201,277,507,533]" captionTargetBox="[201,1405,215,493]" captionTargetId="figure-21@2.[201,1405,215,493]" captionTargetPageId="2" captionText="Figure 2. Radiograph of P. magnaoculus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13184784" httpUri="https://zenodo.org/record/13184784/files/figure.png" pageId="1" pageNumber="320">Figure 2</figureCitation>
).
</paragraph>
<footnote id="E8532A81FFE9CB30EFABE5414EED791F" box="[462,495,1323,1349]" pageId="1" pageNumber="320">
<paragraph id="8BF7368FFFE9CB30EFABE5414EED791F" blockId="1.[462,495,1323,1349]" box="[462,495,1323,1349]" pageId="1" pageNumber="320">(a)</paragraph>
</footnote>
<paragraph id="8BF7368FFFE9CB32ED38E5C74D807990" blockId="1.[859,1501,1453,1875]" lastBlockId="3.[158,799,1276,1482]" lastPageId="3" lastPageNumber="322" pageId="1" pageNumber="320">
<emphasis id="B93CEA9DFFE9CB30ED38E5C74CE3799D" box="[861,993,1453,1479]" italics="true" pageId="1" pageNumber="320">Colouration</emphasis>
: In 70% ethanol, body background greyish brown dorsally extends up to3/4
<superScript id="7C3D9BC7FFE9CB30EAA5E5BB4BCC7985" attach="left" box="[1216,1230,1489,1503]" fontSize="6" pageId="1" pageNumber="320">th</superScript>
laterally, then pale white ventrally. Dorsum of head brown, five brown saddles, first, 3–4 scales broad posterior to occiput and extending up to pectoral-fin base, second, 4 scales wide, between 3–9 branched dorsal-fin rays, extending oblique up to anterior of pelvic-fin base; third and fourth respectively between dorsal-fin origin and caudal-fin base, and fifth, 2 scale wide, around caudal-fin base. Squarish to rectangular black patches on lateral line, patches obscured by the background pigmentation in region between lateral-line origin and middle of dorsal-fin base, posterior four
<taxonomicName id="4C484D0CFFEBCB32EEC5E01A4EBE7CDD" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[160,444,112,135]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="species" species="magnaoculus">
<emphasis id="B93CEA9DFFEBCB32EEC5E01A4EBE7CDD" box="[160,444,112,135]" italics="true" pageId="3" pageNumber="322">Psilorhynchus magnaoculus</emphasis>
</taxonomicName>
, a new species of torrent minnow (Teleostei:
<taxonomicName id="4C484D0CFFEBCB32EDF1E01A4B3F7CDD" authorityName="Hora" authorityYear="1926" box="[916,1085,112,135]" family="Psilorhynchidae" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="family">Psilorhynchidae</taxonomicName>
) from
<collectingCountry id="F35F761FFFEBCB32EAE5E01A4BE67CDD" box="[1152,1252,112,135]" name="Myanmar" pageId="3" pageNumber="322">Myanmar</collectingCountry>
patches, last smallest one on caudal-fin base. Dorsal-fin rays with two rows of spots, one on base and another in middle of the fin. Anal fin immaculate. Paired fin rays with thin melanophores from middle to distal margin. Caudal fin with two black bars, one complete bar in middle, and another incomplete near base of caudal fin.
</paragraph>
<footnote id="E8532A81FFE9CB30EFA8E6634EF27A79" box="[461,496,1545,1571]" pageId="1" pageNumber="320">
<paragraph id="8BF7368FFFE9CB30EFA8E6634EF27A79" blockId="1.[461,496,1545,1571]" box="[461,496,1545,1571]" pageId="1" pageNumber="320">(b)</paragraph>
</footnote>
<footnote id="E8532A81FFE9CB30EFAAE6834EEC7B59" box="[463,494,1769,1795]" pageId="1" pageNumber="320">
<paragraph id="8BF7368FFFE9CB30EFAAE6834EEC7B59" blockId="1.[463,494,1769,1795]" box="[463,494,1769,1795]" pageId="1" pageNumber="320">(c)</paragraph>
</footnote>
<caption id="DF376607FFE9CB30EEC5E77C4ED37B29" ID-DOI="http://doi.org/10.5281/zenodo.13184782" ID-Zenodo-Dep="13184782" httpUri="https://zenodo.org/record/13184782/files/figure.png" pageId="1" pageNumber="320" startId="1.[160,236,1814,1840]" targetBox="[160,797,1600,1757]" targetPageId="1" targetType="figure">
<paragraph id="8BF7368FFFE9CB30EEC5E77C4ED37B29" blockId="1.[160,768,1814,1908]" pageId="1" pageNumber="320">
<emphasis id="B93CEA9DFFE9CB30EEC5E77C4E0A7B6A" bold="true" box="[160,264,1814,1840]" pageId="1" pageNumber="320">Figure 1.</emphasis>
<taxonomicName id="4C484D0CFFE9CB30EF4EE77D4ED77B6A" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[299,469,1814,1840]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="320" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFE9CB30EF4EE77D4ED77B6A" box="[299,469,1814,1840]" italics="true" pageId="1" pageNumber="320">P. magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFE9CB30EFB9E77C4D377B6A" box="[476,565,1814,1840]" pageId="1" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ZSI FF 8211, 47.5 mm SL;
<emphasis id="B93CEA9DFFE9CB30EC65E7524D0F7B08" bold="true" box="[512,525,1848,1874]" pageId="1" pageNumber="320">a</emphasis>
. dorsal
<emphasis id="B93CEA9DFFE9CB30EC00E7524D767B08" bold="true" box="[613,628,1848,1874]" pageId="1" pageNumber="320">b</emphasis>
. lateral
<emphasis id="B93CEA9DFFE9CB30EF4EE7334E357B29" bold="true" box="[299,311,1881,1907]" pageId="1" pageNumber="320">c</emphasis>
. ventral view.
</paragraph>
</caption>
<caption id="DF376607FFEACB33EEACE1914DF57E4C" ID-DOI="http://doi.org/10.5281/zenodo.13184784" ID-Zenodo-Dep="13184784" box="[201,759,507,534]" httpUri="https://zenodo.org/record/13184784/files/figure.png" pageId="2" pageNumber="321" startId="2.[201,277,507,533]" targetBox="[201,1405,215,493]" targetPageId="2" targetType="figure">
<paragraph id="8BF7368FFFEACB33EEACE1914DF57E4C" blockId="2.[201,759,507,534]" box="[201,759,507,534]" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EEACE1914E337E4F" bold="true" box="[201,305,507,533]" pageId="2" pageNumber="321">Figure 2.</emphasis>
Radiograph of
<taxonomicName id="4C484D0CFFEACB33EF96E1964D9F7E4F" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[499,669,507,533]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="321" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFEACB33EF96E1964D9F7E4F" box="[499,669,507,533]" italics="true" pageId="2" pageNumber="321">P. magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFEACB33ECC6E1964DF57E4C" box="[675,759,508,534]" pageId="2" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF376607FFEACB33EEE0E2314B3E7E2F" box="[133,1084,602,629]" pageId="2" pageNumber="321" startId="2.[133,196,603,629]" targetType="table">
<paragraph id="8BF7368FFFEACB33EEE0E2314B3E7E2F" blockId="2.[133,1084,602,629]" box="[133,1084,602,629]" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EEE0E2314FE27E2E" bold="true" box="[133,224,602,629]" pageId="2" pageNumber="321">Table 1.</emphasis>
Biometric data of holotype and paratypes of
<taxonomicName id="4C484D0CFFEACB33EC8EE2314C977E2E" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[747,917,602,628]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="321" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFEACB33EC8EE2314C977E2E" box="[747,917,602,628]" italics="true" pageId="2" pageNumber="321">P. magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFEACB33EDFEE2314CED7E2F" box="[923,1007,603,629]" pageId="2" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
(n=4).
</paragraph>
</caption>
<paragraph id="8BF7368FFFEACB33EC09E2FE4A897A1E" pageId="2" pageNumber="321">
<table id="F948C42FFFEA34CEEEF4E2FE4A8F7A1E" box="[145,1421,660,1604]" gridcols="5" gridrows="24" pageId="2" pageNumber="321">
<tr id="357834CDFFEA34CEEEF4E2FE4A8F7EF6" box="[145,1421,660,684]" gridrow="0" pageId="2" pageNumber="321" rowspan-0="1">
<th id="76A95DB1FFEA34CEEC09E2FE4C677EF6" box="[620,869,660,684]" gridcol="1" gridrow="0" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EC09E2FE4C677EF6" bold="true" box="[620,869,660,684]" pageId="2" pageNumber="321">Holotype (ZSI FF 8211)</emphasis>
</th>
<th id="76A95DB1FFEA34CEEDEEE2FE4B8B7EF6" box="[907,1161,660,684]" gridcol="2" gridrow="0" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EDEEE2FE4B8B7EF6" bold="true" box="[907,1161,660,684]" pageId="2" pageNumber="321">Paratypes (ZSI FF 8212)</emphasis>
</th>
<th id="76A95DB1FFEA34CEEAA4E2FE4A147EF6" box="[1217,1302,660,684]" gridcol="3" gridrow="0" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EAA4E2FE4A147EF6" bold="true" box="[1217,1302,660,684]" pageId="2" pageNumber="321">Average</emphasis>
</th>
<th id="76A95DB1FFEA34CEEB08E2FE4A8F7EF6" box="[1389,1421,660,684]" gridcol="4" gridrow="0" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EB08E2FE4A8F7EF6" bold="true" box="[1389,1421,660,684]" pageId="2" pageNumber="321">SD</emphasis>
</th>
</tr>
<tr id="357834CDFFEA34CEEEF4E2D64A8F7E8E" box="[145,1421,700,724]" gridrow="1" pageId="2" pageNumber="321" rowspan-3="1" rowspan-4="1">
<th id="76A95DB1FFEA34CEEEF4E2D64D2A7E8E" box="[145,552,700,724]" gridcol="0" gridrow="1" pageId="2" pageNumber="321">Standard length (in mm)</th>
<td id="76A95DB1FFEA34CEEC09E2D64C677E8E" box="[620,869,700,724]" gridcol="1" gridrow="1" pageId="2" pageNumber="321">47.5</td>
<td id="76A95DB1FFEA34CEEDEEE2D64B8B7E8E" box="[907,1161,700,724]" gridcol="2" gridrow="1" pageId="2" pageNumber="321">32.0–47.5</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E28E4A8F7EA6" box="[145,1421,740,764]" gridrow="2" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E28E4A8F7EA6" box="[145,1421,740,764]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EEF4E28E4EC67EA6" bold="true" box="[145,452,740,764]" pageId="2" pageNumber="321">In percent of standard length</emphasis>
</th>
</tr>
<tr id="357834CDFFEA34CEEEF4E3664A8F7F7E" box="[145,1421,780,804]" gridrow="3" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3664D2A7F7E" box="[145,552,780,804]" gridcol="0" gridrow="3" pageId="2" pageNumber="321">Body depth</th>
<td id="76A95DB1FFEA34CEEC09E3664C677F7E" box="[620,869,780,804]" gridcol="1" gridrow="3" pageId="2" pageNumber="321">19.4</td>
<td id="76A95DB1FFEA34CEEDEEE3664B8B7F7E" box="[907,1161,780,804]" gridcol="2" gridrow="3" pageId="2" pageNumber="321">15.6–19.4</td>
<td id="76A95DB1FFEA34CEEAA4E3664A147F7E" box="[1217,1302,780,804]" gridcol="3" gridrow="3" pageId="2" pageNumber="321">17.5</td>
<td id="76A95DB1FFEA34CEEB08E3664A8F7F7E" box="[1389,1421,780,804]" gridcol="4" gridrow="3" pageId="2" pageNumber="321">1.5</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E35E4A8F7F16" box="[145,1421,820,844]" gridrow="4" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E35E4D2A7F16" box="[145,552,820,844]" gridcol="0" gridrow="4" pageId="2" pageNumber="321">Head length</th>
<td id="76A95DB1FFEA34CEEC09E35E4C677F16" box="[620,869,820,844]" gridcol="1" gridrow="4" pageId="2" pageNumber="321">19.8</td>
<td id="76A95DB1FFEA34CEEDEEE35E4B8B7F16" box="[907,1161,820,844]" gridcol="2" gridrow="4" pageId="2" pageNumber="321">19.8–22.4</td>
<td id="76A95DB1FFEA34CEEAA4E35E4A147F16" box="[1217,1302,820,844]" gridcol="3" gridrow="4" pageId="2" pageNumber="321">21.3</td>
<td id="76A95DB1FFEA34CEEB08E35E4A8F7F16" box="[1389,1421,820,844]" gridcol="4" gridrow="4" pageId="2" pageNumber="321">1.1</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E3364A8F7F2E" box="[145,1421,860,884]" gridrow="5" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3364D2A7F2E" box="[145,552,860,884]" gridcol="0" gridrow="5" pageId="2" pageNumber="321">Pre-dorsal length</th>
<td id="76A95DB1FFEA34CEEC09E3364C677F2E" box="[620,869,860,884]" gridcol="1" gridrow="5" pageId="2" pageNumber="321">52.2</td>
<td id="76A95DB1FFEA34CEEDEEE3364B8B7F2E" box="[907,1161,860,884]" gridcol="2" gridrow="5" pageId="2" pageNumber="321">46.0–52.2</td>
<td id="76A95DB1FFEA34CEEAA4E3364A147F2E" box="[1217,1302,860,884]" gridcol="3" gridrow="5" pageId="2" pageNumber="321">48.8</td>
<td id="76A95DB1FFEA34CEEB08E3364A8F7F2E" box="[1389,1421,860,884]" gridcol="4" gridrow="5" pageId="2" pageNumber="321">3.0</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E3EE4A8F7FC6" box="[145,1421,900,924]" gridrow="6" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3EE4D2A7FC6" box="[145,552,900,924]" gridcol="0" gridrow="6" pageId="2" pageNumber="321">Pre-anal length</th>
<td id="76A95DB1FFEA34CEEC09E3EE4C677FC6" box="[620,869,900,924]" gridcol="1" gridrow="6" pageId="2" pageNumber="321">76.8</td>
<td id="76A95DB1FFEA34CEEDEEE3EE4B8B7FC6" box="[907,1161,900,924]" gridcol="2" gridrow="6" pageId="2" pageNumber="321">76.8–80.0</td>
<td id="76A95DB1FFEA34CEEAA4E3EE4A147FC6" box="[1217,1302,900,924]" gridcol="3" gridrow="6" pageId="2" pageNumber="321">78.3</td>
<td id="76A95DB1FFEA34CEEB08E3EE4A8F7FC6" box="[1389,1421,900,924]" gridcol="4" gridrow="6" pageId="2" pageNumber="321">1.3</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E3C64A8F7F9E" box="[145,1421,940,964]" gridrow="7" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3C64D2A7F9E" box="[145,552,940,964]" gridcol="0" gridrow="7" pageId="2" pageNumber="321">Pre-pelvic length</th>
<td id="76A95DB1FFEA34CEEC09E3C64C677F9E" box="[620,869,940,964]" gridcol="1" gridrow="7" pageId="2" pageNumber="321">52.6</td>
<td id="76A95DB1FFEA34CEEDEEE3C64B8B7F9E" box="[907,1161,940,964]" gridcol="2" gridrow="7" pageId="2" pageNumber="321">52.6–54.2</td>
<td id="76A95DB1FFEA34CEEAA4E3C64A147F9E" box="[1217,1302,940,964]" gridcol="3" gridrow="7" pageId="2" pageNumber="321">53.4</td>
<td id="76A95DB1FFEA34CEEB08E3C64A8F7F9E" box="[1389,1421,940,964]" gridcol="4" gridrow="7" pageId="2" pageNumber="321">0.7</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E3BE4A8F7FB6" box="[145,1421,980,1004]" gridrow="8" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3BE4D2A7FB6" box="[145,552,980,1004]" gridcol="0" gridrow="8" pageId="2" pageNumber="321">Pre-pectoral length</th>
<td id="76A95DB1FFEA34CEEC09E3BE4C677FB6" box="[620,869,980,1004]" gridcol="1" gridrow="8" pageId="2" pageNumber="321">18.9</td>
<td id="76A95DB1FFEA34CEEDEEE3BE4B8B7FB6" box="[907,1161,980,1004]" gridcol="2" gridrow="8" pageId="2" pageNumber="321">18.8–20.1</td>
<td id="76A95DB1FFEA34CEEAA4E3BE4A147FB6" box="[1217,1302,980,1004]" gridcol="3" gridrow="8" pageId="2" pageNumber="321">19.4</td>
<td id="76A95DB1FFEA34CEEB08E3BE4A8F7FB6" box="[1389,1421,980,1004]" gridcol="4" gridrow="8" pageId="2" pageNumber="321">0.6</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E3964A8F784E" box="[145,1421,1020,1044]" gridrow="9" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E3964D2A784E" box="[145,552,1020,1044]" gridcol="0" gridrow="9" pageId="2" pageNumber="321">Pelvic fin length</th>
<td id="76A95DB1FFEA34CEEC09E3964C67784E" box="[620,869,1020,1044]" gridcol="1" gridrow="9" pageId="2" pageNumber="321">22.1</td>
<td id="76A95DB1FFEA34CEEDEEE3964B8B784E" box="[907,1161,1020,1044]" gridcol="2" gridrow="9" pageId="2" pageNumber="321">22.1–25.1</td>
<td id="76A95DB1FFEA34CEEAA4E3964A14784E" box="[1217,1302,1020,1044]" gridcol="3" gridrow="9" pageId="2" pageNumber="321">23.7</td>
<td id="76A95DB1FFEA34CEEB08E3964A8F784E" box="[1389,1421,1020,1044]" gridcol="4" gridrow="9" pageId="2" pageNumber="321">1.4</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E44E4A8F7866" box="[145,1421,1060,1084]" gridrow="10" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E44E4D2A7866" box="[145,552,1060,1084]" gridcol="0" gridrow="10" pageId="2" pageNumber="321">Pectoral fin length</th>
<td id="76A95DB1FFEA34CEEC09E44E4C677866" box="[620,869,1060,1084]" gridcol="1" gridrow="10" pageId="2" pageNumber="321">30.1</td>
<td id="76A95DB1FFEA34CEEDEEE44E4B8B7866" box="[907,1161,1060,1084]" gridcol="2" gridrow="10" pageId="2" pageNumber="321">27.5–30.3</td>
<td id="76A95DB1FFEA34CEEAA4E44E4A147866" box="[1217,1302,1060,1084]" gridcol="3" gridrow="10" pageId="2" pageNumber="321">29.4</td>
<td id="76A95DB1FFEA34CEEB08E44E4A8F7866" box="[1389,1421,1060,1084]" gridcol="4" gridrow="10" pageId="2" pageNumber="321">1.2</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E4264A8F783E" box="[145,1421,1100,1124]" gridrow="11" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E4264D2A783E" box="[145,552,1100,1124]" gridcol="0" gridrow="11" pageId="2" pageNumber="321">Anus-anal distance</th>
<td id="76A95DB1FFEA34CEEC09E4264C67783E" box="[620,869,1100,1124]" gridcol="1" gridrow="11" pageId="2" pageNumber="321">15.8</td>
<td id="76A95DB1FFEA34CEEDEEE4264B8B783E" box="[907,1161,1100,1124]" gridcol="2" gridrow="11" pageId="2" pageNumber="321">14.1–15.8</td>
<td id="76A95DB1FFEA34CEEAA4E4264A14783E" box="[1217,1302,1100,1124]" gridcol="3" gridrow="11" pageId="2" pageNumber="321">14.7</td>
<td id="76A95DB1FFEA34CEEB08E4264A8F783E" box="[1389,1421,1100,1124]" gridcol="4" gridrow="11" pageId="2" pageNumber="321">0.7</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E41E4A8F78D6" box="[145,1421,1140,1164]" gridrow="12" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E41E4D2A78D6" box="[145,552,1140,1164]" gridcol="0" gridrow="12" pageId="2" pageNumber="321">Snout-anus distance</th>
<td id="76A95DB1FFEA34CEEC09E41E4C6778D6" box="[620,869,1140,1164]" gridcol="1" gridrow="12" pageId="2" pageNumber="321">64.4</td>
<td id="76A95DB1FFEA34CEEDEEE41E4B8B78D6" box="[907,1161,1140,1164]" gridcol="2" gridrow="12" pageId="2" pageNumber="321">59.7–64.4</td>
<td id="76A95DB1FFEA34CEEAA4E41E4A1478D6" box="[1217,1302,1140,1164]" gridcol="3" gridrow="12" pageId="2" pageNumber="321">61.6</td>
<td id="76A95DB1FFEA34CEEB08E41E4A8F78D6" box="[1389,1421,1140,1164]" gridcol="4" gridrow="12" pageId="2" pageNumber="321">2.2</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E4F64A8F78EF" box="[145,1421,1180,1205]" gridrow="13" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E4F64D2A78EF" box="[145,552,1180,1205]" gridcol="0" gridrow="13" pageId="2" pageNumber="321">Length of last unbranched anal-fin ray</th>
<td id="76A95DB1FFEA34CEEC09E4F64C6778EF" box="[620,869,1180,1205]" gridcol="1" gridrow="13" pageId="2" pageNumber="321">17.9</td>
<td id="76A95DB1FFEA34CEEDEEE4F64B8B78EF" box="[907,1161,1180,1205]" gridcol="2" gridrow="13" pageId="2" pageNumber="321">15.6–17.9</td>
<td id="76A95DB1FFEA34CEEAA4E4F64A1478EF" box="[1217,1302,1180,1205]" gridcol="3" gridrow="13" pageId="2" pageNumber="321">16.7</td>
<td id="76A95DB1FFEA34CEEB08E4F64A8F78EF" box="[1389,1421,1180,1205]" gridcol="4" gridrow="13" pageId="2" pageNumber="321">0.9</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E4AE4A8F7887" box="[145,1421,1220,1245]" gridrow="14" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E4AE4D2A7887" box="[145,552,1220,1245]" gridcol="0" gridrow="14" pageId="2" pageNumber="321">Length of last unbranched dorsal-fin ray</th>
<td id="76A95DB1FFEA34CEEC09E4AE4C677887" box="[620,869,1220,1245]" gridcol="1" gridrow="14" pageId="2" pageNumber="321">21.3</td>
<td id="76A95DB1FFEA34CEEDEEE4AE4B8B7887" box="[907,1161,1220,1245]" gridcol="2" gridrow="14" pageId="2" pageNumber="321">21.3–25.1</td>
<td id="76A95DB1FFEA34CEEAA4E4AE4A147887" box="[1217,1302,1220,1245]" gridcol="3" gridrow="14" pageId="2" pageNumber="321">23.4</td>
<td id="76A95DB1FFEA34CEEB08E4AE4A8F7887" box="[1389,1421,1220,1245]" gridcol="4" gridrow="14" pageId="2" pageNumber="321">1.9</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E4864A8F795E" box="[145,1421,1260,1284]" gridrow="15" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E4864D2A795E" box="[145,552,1260,1284]" gridcol="0" gridrow="15" pageId="2" pageNumber="321">Caudal peduncle length</th>
<td id="76A95DB1FFEA34CEEC09E4864C67795E" box="[620,869,1260,1284]" gridcol="1" gridrow="15" pageId="2" pageNumber="321">12.4</td>
<td id="76A95DB1FFEA34CEEDEEE4864B8B795E" box="[907,1161,1260,1284]" gridcol="2" gridrow="15" pageId="2" pageNumber="321">11.9–13.2</td>
<td id="76A95DB1FFEA34CEEAA4E4864A14795E" box="[1217,1302,1260,1284]" gridcol="3" gridrow="15" pageId="2" pageNumber="321">12.5</td>
<td id="76A95DB1FFEA34CEEB08E4864A8F795E" box="[1389,1421,1260,1284]" gridcol="4" gridrow="15" pageId="2" pageNumber="321">0.5</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E57E4A8F7976" box="[145,1421,1300,1324]" gridrow="16" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E57E4D2A7976" box="[145,552,1300,1324]" gridcol="0" gridrow="16" pageId="2" pageNumber="321">Caudal peduncle depth</th>
<td id="76A95DB1FFEA34CEEC09E57E4C677976" box="[620,869,1300,1324]" gridcol="1" gridrow="16" pageId="2" pageNumber="321">10.1</td>
<td id="76A95DB1FFEA34CEEDEEE57E4B8B7976" box="[907,1161,1300,1324]" gridcol="2" gridrow="16" pageId="2" pageNumber="321">09.0–10.2</td>
<td id="76A95DB1FFEA34CEEAA4E57E4A147976" box="[1217,1302,1300,1324]" gridcol="3" gridrow="16" pageId="2" pageNumber="321">09.8</td>
<td id="76A95DB1FFEA34CEEB08E57E4A8F7976" box="[1389,1421,1300,1324]" gridcol="4" gridrow="16" pageId="2" pageNumber="321">0.5</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E5564A8F790E" box="[145,1421,1340,1364]" gridrow="17" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E5564A8F790E" box="[145,1421,1340,1364]" colspan="5" colspanRight="4" gridcol="0" gridrow="17" pageId="2" pageNumber="321">
<emphasis id="B93CEA9DFFEACB33EEF4E5564E9B790E" bold="true" box="[145,409,1340,1364]" pageId="2" pageNumber="321">In percent of head length</emphasis>
</th>
</tr>
<tr id="357834CDFFEA34CEEEF4E50E4A8F7926" box="[145,1421,1380,1404]" gridrow="18" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E50E4D2A7926" box="[145,552,1380,1404]" gridcol="0" gridrow="18" pageId="2" pageNumber="321">Head depth</th>
<td id="76A95DB1FFEA34CEEC09E50E4C677926" box="[620,869,1380,1404]" gridcol="1" gridrow="18" pageId="2" pageNumber="321">63.8</td>
<td id="76A95DB1FFEA34CEEDEEE50E4B8B7926" box="[907,1161,1380,1404]" gridcol="2" gridrow="18" pageId="2" pageNumber="321">57.4–63.8</td>
<td id="76A95DB1FFEA34CEEAA4E50E4A147926" box="[1217,1302,1380,1404]" gridcol="3" gridrow="18" pageId="2" pageNumber="321">61.0</td>
<td id="76A95DB1FFEA34CEEB08E50E4A8F7926" box="[1389,1421,1380,1404]" gridcol="4" gridrow="18" pageId="2" pageNumber="321">2.7</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E5E64A8F79FF" box="[145,1421,1420,1445]" gridrow="19" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E5E64D2A79FF" box="[145,552,1420,1445]" gridcol="0" gridrow="19" pageId="2" pageNumber="321">Eye diameter</th>
<td id="76A95DB1FFEA34CEEC09E5E64C6779FF" box="[620,869,1420,1445]" gridcol="1" gridrow="19" pageId="2" pageNumber="321">43.6</td>
<td id="76A95DB1FFEA34CEEDEEE5E64B8B79FF" box="[907,1161,1420,1445]" gridcol="2" gridrow="19" pageId="2" pageNumber="321">42.6–43.6</td>
<td id="76A95DB1FFEA34CEEAA4E5E64A1479FF" box="[1217,1302,1420,1445]" gridcol="3" gridrow="19" pageId="2" pageNumber="321">43.0</td>
<td id="76A95DB1FFEA34CEEB08E5E64A8F79FF" box="[1389,1421,1420,1445]" gridcol="4" gridrow="19" pageId="2" pageNumber="321">0.4</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E5DE4A8F7996" box="[145,1421,1460,1484]" gridrow="20" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E5DE4D2A7996" box="[145,552,1460,1484]" gridcol="0" gridrow="20" pageId="2" pageNumber="321">Head width</th>
<td id="76A95DB1FFEA34CEEC09E5DE4C677996" box="[620,869,1460,1484]" gridcol="1" gridrow="20" pageId="2" pageNumber="321">74.5</td>
<td id="76A95DB1FFEA34CEEDEEE5DE4B8B7996" box="[907,1161,1460,1484]" gridcol="2" gridrow="20" pageId="2" pageNumber="321">73.0–74.5</td>
<td id="76A95DB1FFEA34CEEAA4E5DE4A147996" box="[1217,1302,1460,1484]" gridcol="3" gridrow="20" pageId="2" pageNumber="321">73.5</td>
<td id="76A95DB1FFEA34CEEB08E5DE4A8F7996" box="[1389,1421,1460,1484]" gridcol="4" gridrow="20" pageId="2" pageNumber="321">0.6</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E5B64A8F79AE" box="[145,1421,1500,1524]" gridrow="21" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E5B64D2A79AE" box="[145,552,1500,1524]" gridcol="0" gridrow="21" pageId="2" pageNumber="321">Inter-orbital distance</th>
<td id="76A95DB1FFEA34CEEC09E5B64C6779AE" box="[620,869,1500,1524]" gridcol="1" gridrow="21" pageId="2" pageNumber="321">45.7</td>
<td id="76A95DB1FFEA34CEEDEEE5B64B8B79AE" box="[907,1161,1500,1524]" gridcol="2" gridrow="21" pageId="2" pageNumber="321">42.2–45.7</td>
<td id="76A95DB1FFEA34CEEAA4E5B64A1479AE" box="[1217,1302,1500,1524]" gridcol="3" gridrow="21" pageId="2" pageNumber="321">43.3</td>
<td id="76A95DB1FFEA34CEEB08E5B64A8F79AE" box="[1389,1421,1500,1524]" gridcol="4" gridrow="21" pageId="2" pageNumber="321">1.6</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E66E4A8F7A46" box="[145,1421,1540,1564]" gridrow="22" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E66E4D2A7A46" box="[145,552,1540,1564]" gridcol="0" gridrow="22" pageId="2" pageNumber="321">Snout length</th>
<td id="76A95DB1FFEA34CEEC09E66E4C677A46" box="[620,869,1540,1564]" gridcol="1" gridrow="22" pageId="2" pageNumber="321">53.2</td>
<td id="76A95DB1FFEA34CEEDEEE66E4B8B7A46" box="[907,1161,1540,1564]" gridcol="2" gridrow="22" pageId="2" pageNumber="321">42.6–53.9</td>
<td id="76A95DB1FFEA34CEEAA4E66E4A147A46" box="[1217,1302,1540,1564]" gridcol="3" gridrow="22" pageId="2" pageNumber="321">49.6</td>
<td id="76A95DB1FFEA34CEEB08E66E4A8F7A46" box="[1389,1421,1540,1564]" gridcol="4" gridrow="22" pageId="2" pageNumber="321">5.1</td>
</tr>
<tr id="357834CDFFEA34CEEEF4E6464A8F7A1E" box="[145,1421,1580,1604]" gridrow="23" pageId="2" pageNumber="321">
<th id="76A95DB1FFEA34CEEEF4E6464D2A7A1E" box="[145,552,1580,1604]" gridcol="0" gridrow="23" pageId="2" pageNumber="321">Mouth width</th>
<td id="76A95DB1FFEA34CEEC09E6464C677A1E" box="[620,869,1580,1604]" gridcol="1" gridrow="23" pageId="2" pageNumber="321">35.0</td>
<td id="76A95DB1FFEA34CEEDEEE6464B8B7A1E" box="[907,1161,1580,1604]" gridcol="2" gridrow="23" pageId="2" pageNumber="321">35.0–41.0</td>
<td id="76A95DB1FFEA34CEEAA4E6464A147A1E" box="[1217,1302,1580,1604]" gridcol="3" gridrow="23" pageId="2" pageNumber="321">37.6</td>
<td id="76A95DB1FFEA34CEEB08E6464A8F7A1E" box="[1389,1421,1580,1604]" gridcol="4" gridrow="23" pageId="2" pageNumber="321">2.6</td>
</tr>
</table>
</paragraph>
<caption id="DF376607FFEBCB32EF06E4CB4B9A78E6" ID-DOI="http://doi.org/10.5281/zenodo.13184786" ID-Zenodo-Dep="13184786" box="[355,1176,1185,1212]" httpUri="https://zenodo.org/record/13184786/files/figure.png" pageId="3" pageNumber="322" startId="3.[355,431,1185,1211]" targetBox="[355,1303,215,1155]" targetPageId="3" targetType="figure">
<paragraph id="8BF7368FFFEBCB32EF06E4CB4B9A78E6" blockId="3.[355,1176,1185,1212]" box="[355,1176,1185,1212]" pageId="3" pageNumber="322">
<emphasis id="B93CEA9DFFEBCB32EF06E4CB4EC978E1" bold="true" box="[355,459,1185,1211]" pageId="3" pageNumber="322">Figure 3.</emphasis>
Map showing the type locality of
<taxonomicName id="4C484D0CFFEBCB32ED3CE4C84B0178E6" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[857,1027,1186,1212]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFEBCB32ED3CE4C84B0178E6" box="[857,1027,1186,1212]" italics="true" pageId="3" pageNumber="322">P. magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFEBCB32EA6CE4C84B5F78E6" box="[1033,1117,1186,1212]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
(▲).
</paragraph>
</caption>
<paragraph id="8BF7368FFFEBCB32EEC5E5884D967AF5" blockId="3.[159,798,1505,1712]" pageId="3" pageNumber="322">
<emphasis id="B93CEA9DFFEBCB32EEC5E5884E0479A1" box="[160,262,1506,1531]" italics="true" pageId="3" pageNumber="322">Diagnosis</emphasis>
:
<taxonomicName id="4C484D0CFFEBCB32EF72E58B4D3C79A1" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[279,574,1505,1531]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFEBCB32EF72E58B4D3C79A1" box="[279,574,1505,1531]" italics="true" pageId="3" pageNumber="322">Psilorhynchus magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFEBCB32EC2CE5884D9D79A6" box="[585,671,1506,1532]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C484D0CFFEBCB32EEADE66C4E337A45" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[200,305,1541,1567]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B93CEA9DFFEBCB32EEADE66C4E337A45" box="[200,305,1541,1567]" italics="true" pageId="3" pageNumber="322">P. balitora</emphasis>
</taxonomicName>
species group (sensu
<bibRefCitation id="EFD94B7EFFEBCB32EC72E66F4DB37A45" author="Conway, K. W." box="[535,689,1541,1567]" pageId="3" pageNumber="322" pagination="50 - 154" refId="ref3764" refString="Conway, K. W. 2011. Osteology of the South Asian genus Psilorhynchus McClelland, 1839 (Teleostei: Ostariophysi: Psilorhynchidae), with investigation of its phylogenetic relationships within the order Cypriniformes. Zool. J. Linn. Soc., 163: 50 - 154. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00698. x" type="journal article" year="2011">Conway, 2011</bibRefCitation>
). The new species is notable from all the congeners in having a squarish snout a larger eye, contributing 42.6–43.6 percent of head length, almost equal to the length of snout; a notch on the ethmoid region and caudal fin with two bars.
</paragraph>
<paragraph id="8BF7368FFFEBCB32EEC5E6AC4D7D7B5E" blockId="3.[160,798,1734,1797]" pageId="3" pageNumber="322">
<emphasis id="B93CEA9DFFEBCB32EEC5E6AC4E167ABA" box="[160,276,1734,1760]" italics="true" pageId="3" pageNumber="322">Etymology</emphasis>
: The species is named after the large eye (Latin), ‘magnaoculus’, noun in apposition.
</paragraph>
<paragraph id="8BF7368FFFEBCB32ED38E4964B667904" blockId="3.[861,1499,1276,1374]" pageId="3" pageNumber="322">
<emphasis id="B93CEA9DFFEBCB32ED38E4964CE1794C" box="[861,995,1276,1302]" italics="true" pageId="3" pageNumber="322">Distribution</emphasis>
:
<taxonomicName id="4C484D0CFFEBCB32ED9AE4964A3E794C" authorityName="Shangningam &amp; Kosygin" authorityYear="2021" box="[1023,1340,1276,1302]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="322" phylum="Chordata" rank="species" species="magnaoculus" status="sp. nov.">
<emphasis id="B93CEA9DFFEBCB32ED9AE4964A3E794C" box="[1023,1340,1276,1302]" italics="true" pageId="3" pageNumber="322">Psilorhynchus magnaoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57E6FFEBCB32EB34E4964AB6794C" box="[1361,1460,1276,1302]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
is presently known only from the Yu River, Chindwin basin in
<collectingCountry id="F35F761FFFEBCB32ED1FE52E4CEA7904" box="[890,1000,1348,1374]" name="Myanmar" pageId="3" pageNumber="322">Myanmar</collectingCountry>
(
<figureCitation id="13732A0AFFEBCB32ED93E52E4B547904" box="[1014,1110,1348,1374]" captionStart="Figure 3" captionStartId="3.[355,431,1185,1211]" captionTargetBox="[355,1303,215,1155]" captionTargetId="figure-255@3.[355,1303,215,1155]" captionTargetPageId="3" captionText="Figure 3. Map showing the type locality of P. magnaoculus sp. nov. (▲)." figureDoi="http://doi.org/10.5281/zenodo.13184786" httpUri="https://zenodo.org/record/13184786/files/figure.png" pageId="3" pageNumber="322">Figure 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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