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<document id="C2700E9699C033130A09B2E34CC0A740" ID-DOI="10.11646/zootaxa.3709.4.1" ID-GBIF-Dataset="b3a7f752-0509-4e01-868d-97e4ab16245e" ID-ISSN="1175-5326" ID-PMC="PMC3914756" ID-PubMed="24511221" ID-Zenodo-Dep="249070" ID-ZooBank="6F6CC130-6A51-4FC0-8E3C-A82B598FFB49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673920785" checkinUser="plazi" docAuthor="Herler, Juergen, Bogorodsky, Sergey V. &amp; Suzuki, Toshiyuki" docDate="2013" docId="03E18789192DAD483AEBFD9DE8BCFE80" docLanguage="en" docName="zt03709p329.pdf" docOrigin="Zootaxa 3709 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gobiodon ater Herler, Bogorodsky &amp; Suzuki, 2013, sp. nov." docType="treatment" docUuid="20EECF5A-16E7-41E3-969F-686865CF8A16" docUuidSource="ZooBank" docVersion="10" lastPageNumber="317" masterDocId="FFD8FFF11920AD593A7CFF86EB3DFFE4" masterDocTitle="Four new species of coral gobies (Teleostei: Gobiidae: Gobiodon), with comments on their relationships within the genus" masterLastPageNumber="329" masterPageNumber="301" pageNumber="314" updateTime="1698349206470" updateUser="plazi">
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<mods:title id="5D51C738247068698C8E90CAB9E0C3BC">Four new species of coral gobies (Teleostei: Gobiidae: Gobiodon), with comments on their relationships within the genus</mods:title>
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<taxonomicName id="4C484D1C192DAD543AEBFD9DEA07FDD7" ID-CoL="6KN6C" box="[151,314,539,564]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="species" species="ater" status="sp. nov.">
<emphasis id="B93CEA8D192DAD543AEBFD9DEA07FDD7" bold="true" box="[151,314,539,564]" italics="true" pageId="13" pageNumber="314">Gobiodon ater</emphasis>
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, 
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<paragraph id="8BF7369F192DAD543AEBFDB9EA68FDBC" blockId="13.[151,588,539,637]" box="[151,341,575,600]" pageId="13" pageNumber="314">Black Coralgoby</paragraph>
<paragraph id="8BF7369F192DAD543AEBFDE2E971FD98" blockId="13.[151,588,539,637]" box="[151,588,612,637]" pageId="13" pageNumber="314">
<figureCitation id="13732A1A192DAD543AEBFDE2EBD9FD99" box="[151,228,612,637]" captionStart="FIGURE 5" captionStartId="14.[151,250,1138,1160]" captionTargetBox="[179,1407,195,1113]" captionTargetId="figure@14.[179,1408,195,1113]" captionTargetPageId="14" captionText="FIGURE 5. Gobiodon ater sp. nov., adult, holotype NMW 94612, 26.5 mm SL, Dahab, Egypt, Red Sea. A. freshly collected; B. ethanol-preserved (epidermal mucus partly removed). Photos by J. Herler." httpUri="https://zenodo.org/record/249075/files/figure.png" pageId="13" pageNumber="314">Figs. 5</figureCitation>
, 
<figureCitation id="13732A1A192DAD543A8DFDE2EA3CFD99" box="[241,257,612,637]" captionStart="FIGURE 6" captionStartId="14.[151,250,1917,1939]" captionTargetBox="[151,1436,1245,1894]" captionTargetId="figure@14.[151,1436,1245,1895]" captionTargetPageId="14" captionText="FIGURE 6. Gobiodon ater sp. nov. A. freshly collected, highly gravid female, CH 232 - 42 - 003, 18.5 mm SL, Dahab, Egypt, Red Sea; B. ethanol-preserved adult (epidermal mucus partly removed), paratype BMNH 2012.3. 20.2, 24.6 mm SL, Dahab, Egypt, Red Sea; C. ethanol-preserved adult (epidermal mucus removed), CH 232 - 42 - 001, 18.2 mm SL, Dahab, Egypt, Red Sea; D. freshly collected adult, CH Mal 243, 19.1 mm SL, Gulhi Island, Maldives, Indian Ocean. Photos by J. Herler." httpUri="https://zenodo.org/record/249076/files/figure.png" pageId="13" pageNumber="314">6</figureCitation>
, 
<figureCitation id="13732A1A192DAD543B71FDE2EA14FD98" box="[269,297,612,636]" captionStart="FIGURE 10" captionStartId="23.[151,250,1629,1651]" captionTargetBox="[224,1369,736,1597]" captionTargetId="figure@23.[209,1384,724,1615]" captionTargetPageId="23" captionText="FIGURE 10. First two principal components of a PCA of Procrustes coordinates of 15 landmarks and 6 semi-landmarks put on 111 specimens of Gobiodon. The holotype (MNHN A- 4015) of G. unicolor (Castelnau, 1873) is marked by a red circle. RS = Red Sea. For species abbreviations see Table 1." httpUri="https://zenodo.org/record/249080/files/figure.png" pageId="13" pageNumber="314">10</figureCitation>
&amp; 
<figureCitation id="13732A1A192DAD543B31FDE2EA56FD98" box="[333,363,612,636]" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="13" pageNumber="314">11</figureCitation>
; 
<tableCitation id="C6CA0324192DAD543B0AFDE2EAE9FD98" box="[374,468,612,637]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionTargetBox="[159,1431,362,1898]" captionTargetPageId="4" captionText="TABLE 1. GenBank accession numbers (GB No.) and references for published sequences (12 S and 16 S rRNA) of 25 taxa (4 new species are underlined) of Gobiodon (G.) and Paragobiodon (P.). Localities: GBR = Great Barrier Reef, Mal = Maldives, PNG = Papua New Guinea, RS = Red Sea, Tai = Taiwan. Abbreviations (PM) and (JH) refer to the first demonstration of sequenced but undescribed species, Munday et al. (1999) and Herler and Hilgers (2009), respectively. Superscript numbers in locality code indicate type status: a: Paratype MNHN 2012 - 0111, b: Paratype BMNH 2006.10. 6.1, c: Paratype NMW 95565." httpUri="http://table.plazi.org/id/DF3766171924AD5D3AEBFF1EE90BFEAB" pageId="13" pageNumber="314" tableUuid="DF3766171924AD5D3AEBFF1EE90BFEAB">Tables 1</tableCitation>
, 
<tableCitation id="C6CA0324192DAD543B9EFDE2EACFFD99" box="[482,498,612,637]" captionStart="TABLE 6" captionStartId="15.[151,239,152,175]" captionTargetBox="[159,1408,266,463]" captionTargetPageId="15" captionText="TABLE 6. Body proportions of holotype and five paratypes of Gobiodon ater sp. nov. from the northern Red Sea. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD). d = damaged." httpUri="http://table.plazi.org/id/DF376617192FAD563AEBFF1EE909FF0B" pageId="13" pageNumber="314" tableUuid="DF376617192FAD563AEBFF1EE909FF0B">6</tableCitation>
, 
<tableCitation id="C6CA0324192DAD543B82FDE3E931FD98" box="[510,524,613,636]" captionStart="TABLE 7" captionStartId="16.[151,239,152,175]" captionTargetBox="[159,1386,202,410]" captionTargetPageId="16" captionText="TABLE 7. Fin counts of holotype and five paratypes of Gobiodon ater sp. nov. from the northern Red Sea." httpUri="http://table.plazi.org/id/DF3766171930AD493AEBFF1EEE31FF4B" pageId="13" pageNumber="314" tableUuid="DF3766171930AD493AEBFF1EEE31FF4B">7</tableCitation>
&amp; 
<tableCitation id="C6CA0324192DAD54384CFDE2E971FD98" box="[560,588,612,636]" captionStart="TABLE 10" captionStartId="21.[151,239,670,693]" captionTargetBox="[159,1393,234,607]" captionTargetPageId="21" captionText="TABLE 10. Selection of genetic (p -) distances from pairwise comparisons of specimens of G. bilineatus sp. nov., G. irregularis sp. nov., G. ater sp. nov. and G. fuscoruber sp. nov. from different regions (GBR: Great Barrier Reef, Jap: Japan, Mal: Maldives, RS: Red Sea, Tai: Taiwan). For comparison of intra- and interspecific genetic distances across the genus, other species from different regions as well as “ G. unicolor ” (sensu Munday et al. (1999) and Harold et al. (2008 )) from the GBR are included. Italic numbers indicate intraspecific distances. Bold values indicate intraspecific distances of more than 2 %. For species abbreviations and Genbank accession numbers see Table 1." httpUri="http://table.plazi.org/id/DF3766171935AD4C3AEBFD18EF0CFCB1" pageId="13" pageNumber="314" tableUuid="DF3766171935AD4C3AEBFD18EF0CFCB1">10</tableCitation>
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<taxonomicName id="4C484D1C192DAD543AEBFD2AEBC1FD26" ID-CoL="4PQ8" box="[151,252,684,706]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D192DAD543AEBFD2AEBC1FD26" box="[151,252,684,706]" italics="true" pageId="13" pageNumber="314">Gobiodon</emphasis>
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sp. 2 
<treatmentCitation id="0AE9108E192DAD543B4BFD2AE95DFD26" author="Herler" box="[311,608,684,706]" page="121" pageId="13" pageNumber="314" year="2005">
<bibRefCitation id="EFD94B6E192DAD543B4BFD2AE912FD26" author="Herler" box="[311,559,684,706]" pageId="13" pageNumber="314" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler and Hilgers 2005</bibRefCitation>
: 121
</treatmentCitation>
, Fig. 14; Niedermüller 
<emphasis id="B93CEA8D192DAD543937FD2BE84AFD26" box="[843,887,684,706]" italics="true" pageId="13" pageNumber="314">et al</emphasis>
. (2009): 1501, 
<figureCitation id="13732A1A192DAD543E6EFD2AEF72FD26" box="[1042,1103,684,706]" captionStart="FIGURE 2" captionStartId="7.[151,250,1856,1878]" captionTargetBox="[151,1436,1118,1833]" captionTargetId="figure@7.[151,1436,1118,1834]" captionTargetPageId="7" captionText="FIGURE 2. Gobiodon bilineatus sp. nov. A. alive adult, PMR VP 2234, 32.7 mm SL, Hanish Island, Yemen, Red Sea; B. alive subadult, PMR VP 3200, 23.7 mm SL, Sharm el Moya, Egypt, Red Sea; C. freshly collected adult, paratype BMNH 2006.10. 6.1, 29.4 mm SL, Dahab, Egypt, Red Sea; D. freshly collected juvenile, CH 232 - 41 - 063, 18.0 mm SL, Dahab, Egypt, Red Sea. Photos by S. V. Bogorodsky (A, B), J. Herler (C, D)." httpUri="https://zenodo.org/record/249072/files/figure.png" pageId="13" pageNumber="314">Fig. 2</figureCitation>
; 
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Herler 
<emphasis id="B93CEA8D192DAD543EDCFD2BEFF0FD26" box="[1184,1229,684,706]" italics="true" pageId="13" pageNumber="314">et al</emphasis>
. (2009)
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: 733
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, 
<figureCitation id="13732A1A192DAD543F27FD2AEEA5FD26" box="[1371,1432,684,706]" captionStart="FIGURE 4" captionStartId="10.[151,250,1841,1863]" captionTargetBox="[151,1436,918,1819]" captionTargetId="figure@10.[151,1436,918,1819]" captionTargetPageId="10" captionText="FIGURE 4. Gobiodon irregularis sp. nov. A. freshly collected adult, paratype MNHN 2006 - 1699, 32.3 mm SL, Dahab, Egypt, Red Sea; B. freshly collected adult, CH 232 - 41 - 020, 31.2 mm SL, Dahab, Egypt, Red Sea; C. freshly collected adult, SMF uncatalogued, 27.6 mm SL, Farasan Archipelago, Red Sea; D. freshly collected subadult, paratype NMW 95565, 23.4 mm SL, Dahab, Egypt, Red Sea; E. alive subadult, PMR VP 3201, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; F. freshly collected juvenile, SAIAB 70430, 15 mm SL, Rodrigues, western Indian Ocean. Photos by J. Herler (A, B, D), S. V. Bogorodsky (C, E), P. Heemstra, © SAIAB (F)." httpUri="https://zenodo.org/record/249074/files/figure.png" pageId="13" pageNumber="314">Fig. 4</figureCitation>
.
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</paragraph>
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<typeStatus id="54F3883D192DAD543AEBFD68EA3BFCE3" box="[151,262,750,775]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="ED59AE5A192DAD543B69FD69EA61FCE2" box="[277,348,751,774]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="314">NMW</collectionCode>
94612, female, 
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SL, Gulf of Aqaba, 
<collectingCountry id="F35F760F192DAD543906FD69E8FDFCE3" box="[890,960,751,775]" name="Egypt" pageId="13" pageNumber="314">Egypt</collectingCountry>
, Dahab, “Napoleon Reef” (
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, 
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), 
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, coll. J. Herler, 
<date id="FFF6105F192DAD543865FC95E9C8FCCF" box="[537,757,786,811]" pageId="13" pageNumber="314" value="2005-11-18">18 November 2005</date>
.
</paragraph>
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<typeStatus id="54F3883D192DAD543ABBFCB0EA7CFCAB" box="[199,321,822,847]" pageId="13" pageNumber="314" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Five specimens. 
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94613, female, 
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SL, 
<date id="FFF6105F192DAD5439B6FCB1EF92FCAB" box="[970,1199,822,847]" pageId="13" pageNumber="314" value="2005-11-14">14 November 2005</date>
, other data same as 
<typeStatus id="54F3883D192DAD543AEBFCDCEBC0FC97" box="[151,253,858,883]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED59AE5A192DAD543B74FCDDEA61FC96" box="[264,348,859,882]" pageId="13" pageNumber="314">BMNH</collectionCode>
2012.3.20.1, male, 
<quantity id="4CB09B7A192DAD543842FCDDE99EFC96" box="[574,675,859,883]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="13" pageNumber="314" unit="mm" value="23.1">23.1 mm</quantity>
SL, other data same as 
<collectionCode id="ED59AE5A192DAD5439C8FCDDE8C6FC96" box="[948,1019,859,882]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="314">NMW</collectionCode>
94613. 
<collectionCode id="ED59AE5A192DAD543E25FCDDEF90FC96" box="[1113,1197,859,882]" pageId="13" pageNumber="314">BMNH</collectionCode>
2012.3.20.2, female, 
<quantity id="4CB09B7A192DAD543AEBFCF8EBC2FC72" box="[151,255,894,919]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="13" pageNumber="314" unit="mm" value="24.6">24.6 mm</quantity>
SL, other data same as 
<collectionCode id="ED59AE5A192DAD543858FCF9E951FC72" box="[548,620,895,918]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="314">NMW</collectionCode>
94613. 
<collectionCode id="ED59AE5A192DAD5438ACFCF9E818FC72" LSID="urn:lsid:biocol.org:col:34988" box="[720,805,895,918]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="314">MNHN</collectionCode>
2012-0110, female, 
<quantity id="4CB09B7A192DAD543E66FCF8EFBFFC72" box="[1050,1154,894,919]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="314" unit="mm" value="25.4">25.4 mm</quantity>
SL, other data same as 
<typeStatus id="54F3883D192DAD543AEBFC24EBC0FC5F" box="[151,253,930,955]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED59AE5A192DAD543B7AFC25EA66FC5E" LSID="urn:lsid:biocol.org:col:34988" box="[262,347,931,954]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="314">MNHN</collectionCode>
2012-0111, male, 
<quantity id="4CB09B7A192DAD543850FC24E9ADFC5E" box="[556,656,930,955]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="13" pageNumber="314" unit="mm" value="15.9">15.9 mm</quantity>
SL, northern Red Sea, 
<collectingCountry id="F35F760F192DAD5439E9FC25E8E6FC5F" box="[917,987,931,955]" name="Egypt" pageId="13" pageNumber="314">Egypt</collectingCountry>
, Marsa Alam (
<geoCoordinate id="EE7C5058192DAD543EFCFC24EE2AFC5E" box="[1152,1303,930,955]" direction="north" orientation="latitude" pageId="13" pageNumber="314" precision="15" value="25.066387">25°03’59’’ N</geoCoordinate>
, 
<geoCoordinate id="EE7C5058192DAD543F5FFC24EB96FC3B" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="15" value="34.90139">34°54’05’’ E</geoCoordinate>
), 
<quantity id="4CB09B7A192DAD543AC3FC41EBD1FC3A" box="[191,236,967,991]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="314" unit="m" value="1.0">1 m</quantity>
, coll. J. Herler, 
<date id="FFF6105F192DAD543BE1FC41E944FC3B" box="[413,633,966,991]" pageId="13" pageNumber="314" value="2005-11-29">29 November 2005</date>
.
</paragraph>
<paragraph id="8BF7369F192DAD543ABBFC6CE8DCFB8B" blockId="13.[151,1436,750,2002]" pageId="13" pageNumber="314">
<emphasis id="B93CEA8D192DAD543ABBFC6CEA81FBE7" bold="true" box="[199,444,1002,1027]" pageId="13" pageNumber="314">Additional material.</emphasis>
<collectingCountry id="F35F760F192DAD543BBAFC6DEAD0FBE7" box="[454,493,1003,1027]" name="Switzerland" pageId="13" pageNumber="314">CH</collectingCountry>
232-42-001, 
<quantity id="4CB09B7A192DAD5438F6FC6DE9CCFBE6" box="[650,753,1003,1027]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="13" pageNumber="314" unit="mm" value="18.2">18.2 mm</quantity>
SL, 
<collectingCountry id="F35F760F192DAD543957FC6DE84CFBE7" box="[811,881,1003,1027]" name="Egypt" pageId="13" pageNumber="314">Egypt</collectingCountry>
, Dahab, coll. J. Herler, 
<date id="FFF6105F192DAD543EFAFC6DEE5AFBE7" box="[1158,1383,1002,1027]" pageId="13" pageNumber="314" value="2003-11-30">30 November 2003</date>
; 
<collectingCountry id="F35F760F192DAD543F09FC6DEEA1FBE7" box="[1397,1436,1003,1027]" name="Switzerland" pageId="13" pageNumber="314">CH</collectingCountry>
232-42-003, 
<quantity id="4CB09B7A192DAD543B5BFB88EAB7FBC2" box="[295,394,1038,1063]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="13" pageNumber="314" unit="mm" value="18.5">18.5 mm</quantity>
SL, 
<collectingCountry id="F35F760F192DAD543BC2FB89E939FBC3" box="[446,516,1039,1063]" name="Egypt" pageId="13" pageNumber="314">Egypt</collectingCountry>
, Dahab, coll. J. Herler, 
<date id="FFF6105F192DAD543974FB88E8A2FBC3" box="[776,927,1038,1063]" pageId="13" pageNumber="314" value="2004-05-26">26 May 2004</date>
; 
<collectingCountry id="F35F760F192DAD5439D6FB89E8ECFBC3" box="[938,977,1039,1063]" name="Switzerland" pageId="13" pageNumber="314">CH</collectingCountry>
Mal 243 (
<collectionCode id="ED59AE5A192DAD543E3FFB89EFBFFBC3" LSID="urn:lsid:biocol.org:col:15536" box="[1091,1154,1039,1063]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="13" pageNumber="314">DNA</collectionCode>
sample and photograph), 
<quantity id="4CB09B7A192DAD543AEBFBB5EBC0FBAE" box="[151,253,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="13" pageNumber="314" unit="mm" value="19.1">19.1 mm</quantity>
SL, Indian Ocean, 
<collectingCountry id="F35F760F192DAD543BA2FBB4E974FBAF" box="[478,585,1074,1099]" name="Maldives" pageId="13" pageNumber="314">Maldives</collectingCountry>
, Gulhi Island, coll. J. Herler, 
<date id="FFF6105F192DAD5439E2FBB5EF6CFBAF" box="[926,1105,1074,1099]" pageId="13" pageNumber="314" value="2007-03-27">27 March 2007</date>
; 
<collectingCountry id="F35F760F192DAD543E22FBB5EFB8FBAF" box="[1118,1157,1075,1099]" name="Switzerland" pageId="13" pageNumber="314">CH</collectingCountry>
Tai 0 0 7 (
<collectionCode id="ED59AE5A192DAD543E89FBB5EE0EFBAF" LSID="urn:lsid:biocol.org:col:15536" box="[1269,1331,1075,1099]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="13" pageNumber="314">DNA</collectionCode>
sample), 
<quantity id="4CB09B7A192DAD543AEBFBD1EBC6FB8A" box="[151,251,1111,1135]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="13" pageNumber="314" unit="mm" value="18.2">18.2 mm</quantity>
SL, southern 
<collectingCountry id="F35F760F192DAD543BEBFBD0EAD1FB8B" box="[407,492,1110,1135]" name="Taiwan" pageId="13" pageNumber="314">Taiwan</collectingCountry>
, Kenting, coll. J. Herler, 0 
<date id="FFF6105F192DAD54396FFBD1E8E3FB8B" box="[787,990,1110,1135]" pageId="13" pageNumber="314" value="2008-12-04">4 December 2008</date>
.
</paragraph>
<paragraph id="8BF7369F192DAD543ABBFBFCEF20FB53" blockId="13.[151,1436,750,2002]" pageId="13" pageNumber="314">
<emphasis id="B93CEA8D192DAD543ABBFBFCEAE5FB77" bold="true" box="[199,472,1146,1171]" pageId="13" pageNumber="314">Comparative material.</emphasis>
<taxonomicName id="4C484D1C192DAD543BA3FBFDE8ACFB77" authority="Bleeker, 1853" authorityName="Bleeker" authorityYear="1853" box="[479,913,1146,1171]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D192DAD543BA3FBFDE9E8FB76" box="[479,725,1147,1171]" italics="true" pageId="13" pageNumber="314">Gobiodon ceramensis</emphasis>
(
<bibRefCitation id="EFD94B6E192DAD543899FBFCE8B5FB77" author="Bleeker" box="[741,904,1146,1171]" pageId="13" pageNumber="314" refString="Bleeker, P. (1853) Nieuwe bijdrage tot de kennis der ichthyologische fauna van Ceram. Natuurkundig Tijdschrift Nederlandsch Indie, 3 (5), 689 - 714." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
)
</taxonomicName>
: 
<collectionCode id="ED59AE5A192DAD5439E0FBFDE8CCFB77" box="[924,1009,1147,1171]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="13" pageNumber="314">OMNH</collectionCode>
P34042 (
<figureCitation id="13732A1A192DAD543E23FBFCEF9FFB77" box="[1119,1186,1146,1171]" captionStart="FIGURE 9" captionStartId="22.[151,250,1660,1682]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure@22.[151,1436,181,1641]" captionTargetPageId="22" captionText="FIGURE 9. Comparative material. Freshly collected adults: Gobiodon cf. bilineatus sp. nov. A. uncatalogued, 28.9 mm SL, Kagi Island, Maldives. B. uncatalogued, 22.9 mm SL, Kenting, southern Taiwan. C. Gobiodon prolixus, ROM 73338, holotype, male 26.2 mm SL, Nha Trang, Vietnam. D. Gobiodon sp. D sensu Munday et al. (1999), uncatalogued, 29.9 mm SL, Hembadhu Island, Maldives E. Gobiodon quinquestrigatus, uncatalogued, 28.7 mm SL, Kenting, southern Taiwan; F. Gobiodon cf. fulvus (sensu Winterbottom &amp; Emery 1986), OMNH P 40259, 20.8 mm SL, Iriomote Island, Japan. G. Gobiodon oculolineatus, OMNH P 40260, 22.6 mm SL, Okinawa Island, Japan. H. Gobiodon ceramensis, OMNH P 34042, 35 mm SL, Ryukyu Island, Japan. Gobiodon unicolor, preserved holotype, MNHN A- 4015, 31 mm SL, I. entire specimen. J. close-up of black opercular spot (arrow) on the right side. Photos by J. Herler (A, B, D, E, I, J), R. Winterbottom (C), T. Suzuki (F-H)." httpUri="https://zenodo.org/record/249079/files/figure.png" pageId="13" pageNumber="314">Fig. 9</figureCitation>
H), 
<quantity id="4CB09B7A192DAD543EB2FBFCEE23FB76" box="[1230,1310,1146,1171]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="314" unit="mm" value="35.0">35 mm</quantity>
SL, 
<collectingCountry id="F35F760F192DAD543F29FBFDEEA5FB76" box="[1365,1432,1147,1170]" name="Japan" pageId="13" pageNumber="314">Japan</collectingCountry>
, Ryukyu Islands, Ishigaki Island, coll. T. Suzuki and M. Suzuki, 
<date id="FFF6105F192DAD54391EFB19EF25FB53" box="[866,1048,1183,1207]" pageId="13" pageNumber="314" value="1997-08-17">17 August 1997</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526514192DAD543ABBFB44EB82FA8F" pageId="13" pageNumber="314" type="diagnosis">
<paragraph id="8BF7369F192DAD543ABBFB44EB82FA8F" blockId="13.[151,1436,750,2002]" pageId="13" pageNumber="314">
<emphasis id="B93CEA8D192DAD543ABBFB44EA7CFB3F" bold="true" box="[199,321,1218,1243]" pageId="13" pageNumber="314">Diagnosis.</emphasis>
Dorsal-fin rays VI + I,10; anal-fin rays I,8; head and body naked; deep curved groove on isthmus; body and caudal peduncle slender (depth 37–41.2% and 13.8–14.7% SL, respectively); head rounded in juveniles, large adults slightly hump-headed. Juveniles and adults uniformly black, including iris. Very small species (&lt;
<quantity id="4CB09B7A192DAD543FFCFA8DEBFEFAA2" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="314" unit="mm" value="28.0">28 mm</quantity>
SL); females of breeding pairs usually significantly larger than males, smallest highly gravid female 
<quantity id="4CB09B7A192DAD543F44FAA8EEA1FAA2" box="[1336,1436,1326,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="13" pageNumber="314" unit="mm" value="18.5">18.5 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="C3526514192DAD493ABBFAF0EF3FFCC9" lastPageId="16" lastPageNumber="317" pageId="13" pageNumber="314" type="description">
<paragraph id="8BF7369F192DAD543ABBFAF0E9CAF86D" blockId="13.[151,1436,750,2002]" pageId="13" pageNumber="314">
<emphasis id="B93CEA8D192DAD543ABBFAF0EA6CFA6B" bold="true" box="[199,337,1398,1423]" pageId="13" pageNumber="314">Description</emphasis>
(based on 6 
<typeStatus id="54F3883D192DAD543B96FAF1E918FA6B" box="[490,549,1399,1423]" pageId="13" pageNumber="314">types</typeStatus>
and several non-types (for osteology)). Large, compressed head; body rather cylindrical. Body proportions and meristics for 
<typeStatus id="54F3883D192DAD5438CCFA1DE9D6FA57" box="[688,747,1435,1459]" pageId="13" pageNumber="314">types</typeStatus>
are provided in 
<tableCitation id="C6CA0324192DAD5439D9FA1CEF3EFA57" box="[933,1027,1434,1459]" captionStart="TABLE 6" captionStartId="15.[151,239,152,175]" captionTargetBox="[159,1408,266,463]" captionTargetPageId="15" captionText="TABLE 6. Body proportions of holotype and five paratypes of Gobiodon ater sp. nov. from the northern Red Sea. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD). d = damaged." httpUri="http://table.plazi.org/id/DF376617192FAD563AEBFF1EE909FF0B" pageId="13" pageNumber="314" tableUuid="DF376617192FAD563AEBFF1EE909FF0B">Tables 6</tableCitation>
and 
<tableCitation id="C6CA0324192DAD543E40FA1DEF71FA56" box="[1084,1100,1435,1458]" captionStart="TABLE 7" captionStartId="16.[151,239,152,175]" captionTargetBox="[159,1386,202,410]" captionTargetPageId="16" captionText="TABLE 7. Fin counts of holotype and five paratypes of Gobiodon ater sp. nov. from the northern Red Sea." httpUri="http://table.plazi.org/id/DF3766171930AD493AEBFF1EEE31FF4B" pageId="13" pageNumber="314" tableUuid="DF3766171930AD493AEBFF1EEE31FF4B">7</tableCitation>
, respectively. Unusual for the genus, the sexes differ in size: females of breeding pairs are usually significantly larger than the males. Dorsal-fin rays VI + I,10 (n = 6); anal-fin rays I,8 (n = 6); pectoral-fin rays 
<emphasis id="B93CEA8D192DAD54390EFA64E8B3FA1F" bold="true" box="[882,910,1506,1531]" pageId="13" pageNumber="314">19</emphasis>
–20 (19:5, 20:1); pelvic-fin rays I,5; caudal fin with 16–17 branched and segmented rays. First dorsal-fin short and rounded. Second dorsal- and anal-fin with long posterior rays, rhomboid in shape. Pelvic disc well-developed, with frenum between spines. No scales. Gill opening as wide as pectoral-fin base, ending ventrally in opposite of 1st or 2nd lower pectoral-fin ray. Gill rakers 0– 1 + 6–7. Deep curved groove present between interopercle and isthmus. Mouth terminal, bending downwards posteriorly, upper jaw reaching to below anterior margin of orbit or mid-orbit. Upper lip more curved down than lower lip. A few rows of small, slender and recurved teeth in both jaws behind the outer row, which has 10 larger and slightly recurved teeth in upper and lower jaw. In lower jaw, one pair of large, postsymphysial canines on each side; one or both of these teeth often smaller or absent, probably due to tooth loss and replacement. Anterior and posterior nasal openings at the end of short tubes. Head sensory canals typical for 
<taxonomicName id="4C484D1C192DAD543E20F8AFEA0EF882" authority="Winterbottom &amp; Harold 2005" authorityName="Winterbottom &amp; Harold" authorityYear="2005" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D192DAD543E20F8AFEFF1F8A5" box="[1116,1228,1833,1857]" italics="true" pageId="13" pageNumber="314">Gobiodon</emphasis>
(
<bibRefCitation id="EFD94B6E192DAD543EA2F8AFEA16F882" author="Winterbottom" pageId="13" pageNumber="314" refString="Winterbottom, R. &amp; Harold, A. S. (2005) Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes: Gobiidae) from the Indo-west Pacific. Proceedings of the Biological Society of Washington, 118 (3), 582 - 589." type="journal article" year="2005">Winterbottom &amp; Harold 2005</bibRefCitation>
)
</taxonomicName>
, with anterior oculoscapular (pores NA (paired), AI, PI (unpaired), SO, 
<collectingCountry id="F35F760F192DAD543E01F8C8EF98F882" box="[1149,1189,1870,1894]" name="Austria" pageId="13" pageNumber="314">AO</collectingCountry>
and IT (paired)) and preopercular (three pores on each side) canals present.
</paragraph>
<paragraph id="8BF7369F192DAD543ABBF813E82AF836" blockId="13.[151,1436,750,2002]" pageId="13" pageNumber="314">
<emphasis id="B93CEA8D192DAD543ABBF813EAB2F84A" bold="true" box="[199,399,1941,1966]" pageId="13" pageNumber="314">Life colouration.</emphasis>
Uniformly black, including iris; thus eye hardly visible in living fish in the field (
<figureCitation id="13732A1A192DAD543F4FF813EEBCF84A" box="[1331,1409,1941,1966]" captionStart="FIGURE 5" captionStartId="14.[151,250,1138,1160]" captionTargetBox="[179,1407,195,1113]" captionTargetId="figure@14.[179,1408,195,1113]" captionTargetPageId="14" captionText="FIGURE 5. Gobiodon ater sp. nov., adult, holotype NMW 94612, 26.5 mm SL, Dahab, Egypt, Red Sea. A. freshly collected; B. ethanol-preserved (epidermal mucus partly removed). Photos by J. Herler." httpUri="https://zenodo.org/record/249075/files/figure.png" pageId="13" pageNumber="314">Figs. 5</figureCitation>
A, 6A). When stressed, black colouration can turn into grey.
</paragraph>
<caption id="DF376617192EAD573AEBFBF4E8A0FB42" httpUri="https://zenodo.org/record/249075/files/figure.png" pageId="14" pageNumber="315" targetBox="[179,1407,195,1113]" targetPageId="14">
<paragraph id="8BF7369F192EAD573AEBFBF4E8A0FB42" blockId="14.[151,1436,1138,1190]" pageId="14" pageNumber="315">
<emphasis id="B93CEA8D192EAD573AEBFBF4EA2EFB63" bold="true" box="[151,275,1138,1160]" pageId="14" pageNumber="315">FIGURE 5.</emphasis>
<taxonomicName id="4C484D1C192EAD573B66FBF4EA92FB6C" box="[282,431,1138,1160]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="315" phylum="Chordata" rank="species" species="ater" status="sp. nov.">
<emphasis id="B93CEA8D192EAD573B66FBF4EA92FB6C" box="[282,431,1138,1160]" italics="true" pageId="14" pageNumber="315">Gobiodon ater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192EAD573BCAFBF5E935FB6C" box="[438,520,1139,1160]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
, adult, holotype NMW 94612, 26.5 mm SL, Dahab, Egypt, Red Sea. 
<emphasis id="B93CEA8D192EAD573EB0FBF4EFD8FB63" bold="true" box="[1228,1253,1138,1159]" pageId="14" pageNumber="315">A.</emphasis>
freshly collected; 
<emphasis id="B93CEA8D192EAD573AEBFB17EB93FB42" bold="true" box="[151,174,1169,1190]" pageId="14" pageNumber="315">B.</emphasis>
ethanol-preserved (epidermal mucus partly removed). Photos by J.Herler.
</paragraph>
</caption>
<caption id="DF376617192EAD573AEBF8FBEE0BF80B" httpUri="https://zenodo.org/record/249076/files/figure.png" pageId="14" pageNumber="315" targetBox="[151,1436,1245,1894]" targetPageId="14">
<paragraph id="8BF7369F192EAD573AEBF8FBEE0BF80B" blockId="14.[151,1436,1916,2031]" pageId="14" pageNumber="315">
<emphasis id="B93CEA8D192EAD573AEBF8FBEA29F876" bold="true" box="[151,276,1917,1939]" pageId="14" pageNumber="315">FIGURE 6.</emphasis>
<taxonomicName id="4C484D1C192EAD573B60F8FAEA8FF877" box="[284,434,1916,1939]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="315" phylum="Chordata" rank="species" species="ater" status="sp. nov.">
<emphasis id="B93CEA8D192EAD573B60F8FAEA8FF877" box="[284,434,1916,1939]" italics="true" pageId="14" pageNumber="315">Gobiodon ater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192EAD573BC5F8FBE934F876" box="[441,521,1917,1938]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEA8D192EAD57386CF8FBE914F876" bold="true" box="[528,553,1917,1938]" pageId="14" pageNumber="315">A.</emphasis>
freshly collected, highly gravid female, CH 232-42-003, 18.5 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192EAD573A80F81AEA2EF855" bold="true" box="[252,275,1948,1969]" pageId="14" pageNumber="315">B.</emphasis>
ethanol-preserved adult (epidermal mucus partly removed), paratype BMNH 2012.3.20.2, 24.6 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192EAD573B34F83CEA5DF834" bold="true" box="[328,352,1978,2000]" pageId="14" pageNumber="315">C.</emphasis>
ethanol-preserved adult (epidermal mucus removed), CH 232-42-001, 18.2 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192EAD573AB5F85FEBDCF80A" bold="true" box="[201,225,2009,2030]" pageId="14" pageNumber="315">D.</emphasis>
freshly collected adult, CH Mal 243, 19.1 mm SL, Gulhi Island, Maldives, Indian Ocean. Photos by J.Herler.
</paragraph>
</caption>
<caption id="DF376617192FAD563AEBFF1EE909FF0B" ID-Table-UUID="DF376617192FAD563AEBFF1EE909FF0B" httpUri="http://table.plazi.org/id/DF376617192FAD563AEBFF1EE909FF0B" pageId="15" pageNumber="316" targetBox="[159,1408,266,463]" targetIsTable="true" targetPageId="15">
<paragraph id="8BF7369F192FAD563AEBFF1EE909FF0B" blockId="15.[151,1436,151,239]" pageId="15" pageNumber="316">
<emphasis id="B93CEA8D192FAD563AEBFF1EEA35FF4B" bold="true" box="[151,264,152,175]" pageId="15" pageNumber="316">TABLE 6.</emphasis>
Body proportions of holotype and five paratypes of 
<taxonomicName id="4C484D1C192FAD563949FF11E8EDFF4B" box="[821,976,151,175]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="ater" status="sp. nov.">
<emphasis id="B93CEA8D192FAD563949FF11E8EDFF4B" box="[821,976,151,175]" italics="true" pageId="15" pageNumber="316">Gobiodon ater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192FAD5639ABFF1EEF17FF4B" box="[983,1066,152,175]" pageId="15" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
from the northern Red Sea. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD). d = damaged.
</paragraph>
</caption>
<paragraph id="8BF7369F192FAD563AE3FE8CEE42FE54" blockId="15.[159,1408,266,1618]" pageId="15" pageNumber="316">
<table id="F948C43F192F52A63AE3FE8CEEBDFE2B" box="[159,1408,266,463]" gridcols="8" gridrows="3" pageId="15" pageNumber="316">
<tr id="357834DD192F52A63AE3FE8CEEBDFEBA" box="[159,1408,266,350]" gridrow="0" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FE8CEAB1FEBA" box="[159,396,266,350]" gridcol="0" gridrow="0" pageId="15" pageNumber="316">Status Coll.No.</th>
<th id="76A95DA1192F52A63BD0FE8CE937FEBA" box="[428,522,266,350]" gridcol="1" gridrow="0" pageId="15" pageNumber="316">Holotype NMW 94612</th>
<th id="76A95DA1192F52A63848FE8CE9B0FEBA" box="[564,653,266,350]" gridcol="2" gridrow="0" pageId="15" pageNumber="316">Paratype NMW 94613</th>
<th id="76A95DA1192F52A638C7FE8CE80EFEBA" box="[699,819,266,350]" gridcol="3" gridrow="0" pageId="15" pageNumber="316">Paratype BMNH 2012.3.20.1</th>
<th id="76A95DA1192F52A6392FFE8CE8F1FEBA" box="[851,972,266,350]" gridcol="4" gridrow="0" pageId="15" pageNumber="316">Paratype BMNH 2012.3.20.2</th>
<th id="76A95DA1192F52A63988FE8CEF5CFEBA" box="[1012,1121,266,350]" gridcol="5" gridrow="0" pageId="15" pageNumber="316">Paratype MNHN 2012-0110</th>
<th id="76A95DA1192F52A63EF1FE8CEFC4FEBA" box="[1165,1273,266,350]" gridcol="6" gridrow="0" pageId="15" pageNumber="316">Paratype MNHN 2012-0111</th>
<th id="76A95DA1192F52A63F61FE8CEEBDFEBA" box="[1309,1408,266,350]" gridcol="7" gridrow="0" pageId="15" pageNumber="316">MEAN (±SD)</th>
</tr>
<tr id="357834DD192F52A63AE3FEF7EEBDFE63" box="[159,1408,369,391]" gridrow="1" pageId="15" pageNumber="316" rowspan-7="1">
<th id="76A95DA1192F52A63AE3FEF7EAB1FE63" box="[159,396,369,391]" gridcol="0" gridrow="1" pageId="15" pageNumber="316">SL (mm)</th>
<td id="76A95DA1192F52A63BD0FEF7E937FE63" box="[428,522,369,391]" gridcol="1" gridrow="1" pageId="15" pageNumber="316">26.5</td>
<td id="76A95DA1192F52A63848FEF7E9B0FE63" box="[564,653,369,391]" gridcol="2" gridrow="1" pageId="15" pageNumber="316">26.4</td>
<td id="76A95DA1192F52A638C7FEF7E80EFE63" box="[699,819,369,391]" gridcol="3" gridrow="1" pageId="15" pageNumber="316">23.1</td>
<td id="76A95DA1192F52A6392FFEF7E8F1FE63" box="[851,972,369,391]" gridcol="4" gridrow="1" pageId="15" pageNumber="316">24.6</td>
<td id="76A95DA1192F52A63988FEF7EF5CFE63" box="[1012,1121,369,391]" gridcol="5" gridrow="1" pageId="15" pageNumber="316">25.4</td>
<td id="76A95DA1192F52A63EF1FEF7EFC4FE63" box="[1165,1273,369,391]" gridcol="6" gridrow="1" pageId="15" pageNumber="316">15.9</td>
</tr>
<tr id="357834DD192F52A63AE3FE1CEEBDFE2B" box="[159,1408,410,463]" gridrow="2" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FE1CEAB1FE2B" box="[159,396,410,463]" gridcol="0" gridrow="2" pageId="15" pageNumber="316">Snout to first dorsal-fin origin</th>
<td id="76A95DA1192F52A63BD0FE1CE937FE2B" box="[428,522,410,463]" gridcol="1" gridrow="2" pageId="15" pageNumber="316">36.7</td>
<td id="76A95DA1192F52A63848FE1CE9B0FE2B" box="[564,653,410,463]" gridcol="2" gridrow="2" pageId="15" pageNumber="316">38.0</td>
<td id="76A95DA1192F52A638C7FE1CE80EFE2B" box="[699,819,410,463]" gridcol="3" gridrow="2" pageId="15" pageNumber="316">37.7</td>
<td id="76A95DA1192F52A6392FFE1CE8F1FE2B" box="[851,972,410,463]" gridcol="4" gridrow="2" pageId="15" pageNumber="316">36.9</td>
<td id="76A95DA1192F52A63988FE1CEF5CFE2B" box="[1012,1121,410,463]" gridcol="5" gridrow="2" pageId="15" pageNumber="316">37.5</td>
<td id="76A95DA1192F52A63EF1FE1CEFC4FE2B" box="[1165,1273,410,463]" gridcol="6" gridrow="2" pageId="15" pageNumber="316">41.8</td>
<td id="76A95DA1192F52A63F61FE1CEEBDFE2B" box="[1309,1408,410,463]" gridcol="7" gridrow="2" pageId="15" pageNumber="316">38.4 (2.0)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF7369F192FAD563AE3FE64EE42F9D7" blockId="15.[159,1408,266,1618]" pageId="15" pageNumber="316">
<table id="F948C43F192F52A63AE3FE64EEBDF9B6" box="[159,1408,482,1618]" gridcols="7" gridrows="22" pageId="15" pageNumber="316">
<tr id="357834DD192F52A63AE3FE64EEBDFDF3" box="[159,1408,482,535]" gridrow="0" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FE64EAE5FDF3" box="[159,472,482,535]" gridcol="0" gridrow="0" pageId="15" pageNumber="316">First dorsal-fin origin to 25.7 second dorsal-fin origin</th>
<th id="76A95DA1192F52A63848FE64E95DFDF3" box="[564,608,482,535]" gridcol="1" gridrow="0" pageId="15" pageNumber="316">26.6</th>
<th id="76A95DA1192F52A638C7FE64E9DAFDF3" box="[699,743,482,535]" gridcol="2" gridrow="0" pageId="15" pageNumber="316">24.6</th>
<th id="76A95DA1192F52A6392FFE64E842FDF3" box="[851,895,482,535]" gridcol="3" gridrow="0" pageId="15" pageNumber="316">25.1</th>
<th id="76A95DA1192F52A63988FE64EF1DFDF3" box="[1012,1056,482,535]" gridcol="4" gridrow="0" pageId="15" pageNumber="316">26.8</th>
<th id="76A95DA1192F52A63EF0FE64EF84FDF3" box="[1164,1209,482,535]" gridcol="5" gridrow="0" pageId="15" pageNumber="316">24.5</th>
<th id="76A95DA1192F52A63F60FE64EEBDFDF3" box="[1308,1408,482,535]" gridcol="6" gridrow="0" pageId="15" pageNumber="316">25.5 (1.1)</th>
</tr>
<tr id="357834DD192F52A63AE3FDACEEBDFDBB" box="[159,1408,554,607]" gridrow="1" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FDACEAE5FDBB" box="[159,472,554,607]" gridcol="0" gridrow="1" pageId="15" pageNumber="316">Second dorsal-fin origin 31.5 to anal-fin origin</th>
<td id="76A95DA1192F52A63848FDACE95DFDBB" box="[564,608,554,607]" gridcol="1" gridrow="1" pageId="15" pageNumber="316">30.7</td>
<td id="76A95DA1192F52A638C7FDACE9DAFDBB" box="[699,743,554,607]" gridcol="2" gridrow="1" pageId="15" pageNumber="316">32.3</td>
<td id="76A95DA1192F52A6392FFDACE842FDBB" box="[851,895,554,607]" gridcol="3" gridrow="1" pageId="15" pageNumber="316">30.8</td>
<td id="76A95DA1192F52A63988FDACEF1DFDBB" box="[1012,1056,554,607]" gridcol="4" gridrow="1" pageId="15" pageNumber="316">29.5</td>
<td id="76A95DA1192F52A63EF0FDACEF84FDBB" box="[1164,1209,554,607]" gridcol="5" gridrow="1" pageId="15" pageNumber="316">29.2</td>
<td id="76A95DA1192F52A63F60FDACEEBDFDBB" box="[1308,1408,554,607]" gridcol="6" gridrow="1" pageId="15" pageNumber="316">30.5 (1.2)</td>
</tr>
<tr id="357834DD192F52A63AE3FDF4EEBDFD43" box="[159,1408,626,679]" gridrow="2" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FDF4EAE5FD43" box="[159,472,626,679]" gridcol="0" gridrow="2" pageId="15" pageNumber="316">Pelvic-fin origin to anal- 31.1 fin origin</th>
<td id="76A95DA1192F52A63848FDF4E95DFD43" box="[564,608,626,679]" gridcol="1" gridrow="2" pageId="15" pageNumber="316">31.7</td>
<td id="76A95DA1192F52A638C7FDF4E9DAFD43" box="[699,743,626,679]" gridcol="2" gridrow="2" pageId="15" pageNumber="316">30.6</td>
<td id="76A95DA1192F52A6392FFDF4E842FD43" box="[851,895,626,679]" gridcol="3" gridrow="2" pageId="15" pageNumber="316">30.4</td>
<td id="76A95DA1192F52A63988FDF4EF1DFD43" box="[1012,1056,626,679]" gridcol="4" gridrow="2" pageId="15" pageNumber="316">34.3</td>
<td id="76A95DA1192F52A63EF0FDF4EF84FD43" box="[1164,1209,626,679]" gridcol="5" gridrow="2" pageId="15" pageNumber="316">25.1</td>
<td id="76A95DA1192F52A63F60FDF4EEBDFD43" box="[1308,1408,626,679]" gridcol="6" gridrow="2" pageId="15" pageNumber="316">30.4 (3.3)</td>
</tr>
<tr id="357834DD192F52A63AE3FD3CEEBDFD0B" box="[159,1408,698,751]" gridrow="3" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FD3CEAE5FD0B" box="[159,472,698,751]" gridcol="0" gridrow="3" pageId="15" pageNumber="316">Snout to pelvic-fin 37.9 origin</th>
<td id="76A95DA1192F52A63848FD3CE95DFD0B" box="[564,608,698,751]" gridcol="1" gridrow="3" pageId="15" pageNumber="316">38.2</td>
<td id="76A95DA1192F52A638C7FD3CE9DAFD0B" box="[699,743,698,751]" gridcol="2" gridrow="3" pageId="15" pageNumber="316">40.3</td>
<td id="76A95DA1192F52A6392FFD3CE842FD0B" box="[851,895,698,751]" gridcol="3" gridrow="3" pageId="15" pageNumber="316">38.5</td>
<td id="76A95DA1192F52A63988FD3CEF1DFD0B" box="[1012,1056,698,751]" gridcol="4" gridrow="3" pageId="15" pageNumber="316">38.5</td>
<td id="76A95DA1192F52A63EF0FD3CEF84FD0B" box="[1164,1209,698,751]" gridcol="5" gridrow="3" pageId="15" pageNumber="316">39.3</td>
<td id="76A95DA1192F52A63F60FD3CEEBDFD0B" box="[1308,1408,698,751]" gridcol="6" gridrow="3" pageId="15" pageNumber="316">39.0 (0.9)</td>
</tr>
<tr id="357834DD192F52A63AE3FC84EEBDFCD3" box="[159,1408,770,823]" gridrow="4" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FC84EAE5FCD3" box="[159,472,770,823]" gridcol="0" gridrow="4" pageId="15" pageNumber="316">First dorsal-fin origin to 40.0 pelvic-fin origin</th>
<td id="76A95DA1192F52A63848FC84E95DFCD3" box="[564,608,770,823]" gridcol="1" gridrow="4" pageId="15" pageNumber="316">39.9</td>
<td id="76A95DA1192F52A638C7FC84E9DAFCD3" box="[699,743,770,823]" gridcol="2" gridrow="4" pageId="15" pageNumber="316">39.2</td>
<td id="76A95DA1192F52A6392FFC84E842FCD3" box="[851,895,770,823]" gridcol="3" gridrow="4" pageId="15" pageNumber="316">39.2</td>
<td id="76A95DA1192F52A63988FC84EF1DFCD3" box="[1012,1056,770,823]" gridcol="4" gridrow="4" pageId="15" pageNumber="316">40.6</td>
<td id="76A95DA1192F52A63EF0FC84EF84FCD3" box="[1164,1209,770,823]" gridcol="5" gridrow="4" pageId="15" pageNumber="316">37.0</td>
<td id="76A95DA1192F52A63F60FC84EEBDFCD3" box="[1308,1408,770,823]" gridcol="6" gridrow="4" pageId="15" pageNumber="316">39.2 (1.4)</td>
</tr>
<tr id="357834DD192F52A63AE3FCCCEEBDFC9B" box="[159,1408,842,895]" gridrow="5" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FCCCEAE5FC9B" box="[159,472,842,895]" gridcol="0" gridrow="5" pageId="15" pageNumber="316">First dorsal-fin origin to 46.5 anal-fin origin</th>
<td id="76A95DA1192F52A63848FCCCE95DFC9B" box="[564,608,842,895]" gridcol="1" gridrow="5" pageId="15" pageNumber="316">46.4</td>
<td id="76A95DA1192F52A638C7FCCCE9DAFC9B" box="[699,743,842,895]" gridcol="2" gridrow="5" pageId="15" pageNumber="316">45.6</td>
<td id="76A95DA1192F52A6392FFCCCE842FC9B" box="[851,895,842,895]" gridcol="3" gridrow="5" pageId="15" pageNumber="316">45.5</td>
<td id="76A95DA1192F52A63988FCCCEF1DFC9B" box="[1012,1056,842,895]" gridcol="4" gridrow="5" pageId="15" pageNumber="316">46.7</td>
<td id="76A95DA1192F52A63EF0FCCCEF84FC9B" box="[1164,1209,842,895]" gridcol="5" gridrow="5" pageId="15" pageNumber="316">42.7</td>
<td id="76A95DA1192F52A63F60FCCCEEBDFC9B" box="[1308,1408,842,895]" gridcol="6" gridrow="5" pageId="15" pageNumber="316">45.4 (1.6)</td>
</tr>
<tr id="357834DD192F52A63AE3FC14EEBDFC23" box="[159,1408,914,967]" gridrow="6" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FC14EAE5FC23" box="[159,472,914,967]" gridcol="0" gridrow="6" pageId="15" pageNumber="316">Pelvic-fin origin to 45.5 second dorsal-fin origin</th>
<td id="76A95DA1192F52A63848FC14E95DFC23" box="[564,608,914,967]" gridcol="1" gridrow="6" pageId="15" pageNumber="316">44.9</td>
<td id="76A95DA1192F52A638C7FC14E9DAFC23" box="[699,743,914,967]" gridcol="2" gridrow="6" pageId="15" pageNumber="316">42.2</td>
<td id="76A95DA1192F52A6392FFC14E842FC23" box="[851,895,914,967]" gridcol="3" gridrow="6" pageId="15" pageNumber="316">44.0</td>
<td id="76A95DA1192F52A63988FC14EF1DFC23" box="[1012,1056,914,967]" gridcol="4" gridrow="6" pageId="15" pageNumber="316">45.3</td>
<td id="76A95DA1192F52A63EF0FC14EF84FC23" box="[1164,1209,914,967]" gridcol="5" gridrow="6" pageId="15" pageNumber="316">39.8</td>
<td id="76A95DA1192F52A63F60FC14EEBDFC23" box="[1308,1408,914,967]" gridcol="6" gridrow="6" pageId="15" pageNumber="316">43.2 (2.3)</td>
</tr>
<tr id="357834DD192F52A63AE3FC5CEEBDFC14" box="[159,1408,986,1008]" gridrow="7" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FC5CEAE5FC14" box="[159,472,986,1008]" gridcol="0" gridrow="7" pageId="15" pageNumber="316">Head length 29.1</th>
<td id="76A95DA1192F52A63848FC5CE95DFC14" box="[564,608,986,1008]" gridcol="1" gridrow="7" pageId="15" pageNumber="316">29.5</td>
<td id="76A95DA1192F52A638C7FC5CE9DAFC14" box="[699,743,986,1008]" gridcol="2" gridrow="7" pageId="15" pageNumber="316">31.4</td>
<td id="76A95DA1192F52A6392FFC5CE842FC14" box="[851,895,986,1008]" gridcol="3" gridrow="7" pageId="15" pageNumber="316">30.0</td>
<td id="76A95DA1192F52A63988FC5CEF1DFC14" box="[1012,1056,986,1008]" gridcol="4" gridrow="7" pageId="15" pageNumber="316">28.9</td>
<td id="76A95DA1192F52A63EF0FC5CEF84FC14" box="[1164,1209,986,1008]" gridcol="5" gridrow="7" pageId="15" pageNumber="316">31.7</td>
<td id="76A95DA1192F52A63F60FC5CEEBDFC14" box="[1308,1408,986,1008]" gridcol="6" gridrow="7" pageId="15" pageNumber="316">30.3 (1.2)</td>
</tr>
<tr id="357834DD192F52A63AE3FB82EEBDFBFE" box="[159,1408,1028,1050]" gridrow="8" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FB82EAE5FBFE" box="[159,472,1028,1050]" gridcol="0" gridrow="8" pageId="15" pageNumber="316">Head depth 33.0</th>
<td id="76A95DA1192F52A63848FB82E95DFBFE" box="[564,608,1028,1050]" gridcol="1" gridrow="8" pageId="15" pageNumber="316">32.6</td>
<td id="76A95DA1192F52A638C7FB82E9DAFBFE" box="[699,743,1028,1050]" gridcol="2" gridrow="8" pageId="15" pageNumber="316">33.4</td>
<td id="76A95DA1192F52A6392FFB82E842FBFE" box="[851,895,1028,1050]" gridcol="3" gridrow="8" pageId="15" pageNumber="316">32.1</td>
<td id="76A95DA1192F52A63988FB82EF1DFBFE" box="[1012,1056,1028,1050]" gridcol="4" gridrow="8" pageId="15" pageNumber="316">33.8</td>
<td id="76A95DA1192F52A63EF0FB82EF84FBFE" box="[1164,1209,1028,1050]" gridcol="5" gridrow="8" pageId="15" pageNumber="316">29.3</td>
<td id="76A95DA1192F52A63F60FB82EEBDFBFE" box="[1308,1408,1028,1050]" gridcol="6" gridrow="8" pageId="15" pageNumber="316">32.2 (1.8)</td>
</tr>
<tr id="357834DD192F52A63AE3FBABEEBDFBA7" box="[159,1408,1069,1091]" gridrow="9" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FBABEAE5FBA7" box="[159,472,1069,1091]" gridcol="0" gridrow="9" pageId="15" pageNumber="316">Body depth 40.1</th>
<td id="76A95DA1192F52A63848FBABE95DFBA7" box="[564,608,1069,1091]" gridcol="1" gridrow="9" pageId="15" pageNumber="316">41.2</td>
<td id="76A95DA1192F52A638C7FBABE9DAFBA7" box="[699,743,1069,1091]" gridcol="2" gridrow="9" pageId="15" pageNumber="316">39.7</td>
<td id="76A95DA1192F52A6392FFBABE842FBA7" box="[851,895,1069,1091]" gridcol="3" gridrow="9" pageId="15" pageNumber="316">39.3</td>
<td id="76A95DA1192F52A63988FBABEF1DFBA7" box="[1012,1056,1069,1091]" gridcol="4" gridrow="9" pageId="15" pageNumber="316">40.5</td>
<td id="76A95DA1192F52A63EF0FBABEF84FBA7" box="[1164,1209,1069,1091]" gridcol="5" gridrow="9" pageId="15" pageNumber="316">37.0</td>
<td id="76A95DA1192F52A63F60FBABEEBDFBA7" box="[1308,1408,1069,1091]" gridcol="6" gridrow="9" pageId="15" pageNumber="316">39.5 (1.6)</td>
</tr>
<tr id="357834DD192F52A63AE3FBD0EEBDFB88" box="[159,1408,1110,1132]" gridrow="10" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FBD0EAE5FB88" box="[159,472,1110,1132]" gridcol="0" gridrow="10" pageId="15" pageNumber="316">Pelvic-fin length 15.4</th>
<td id="76A95DA1192F52A63848FBD0E95DFB88" box="[564,608,1110,1132]" gridcol="1" gridrow="10" pageId="15" pageNumber="316">15.9</td>
<td id="76A95DA1192F52A638C7FBD0E9DAFB88" box="[699,743,1110,1132]" gridcol="2" gridrow="10" pageId="15" pageNumber="316">17.5</td>
<td id="76A95DA1192F52A6392FFBD0E842FB88" box="[851,895,1110,1132]" gridcol="3" gridrow="10" pageId="15" pageNumber="316">16.4</td>
<td id="76A95DA1192F52A63988FBD0EF1DFB88" box="[1012,1056,1110,1132]" gridcol="4" gridrow="10" pageId="15" pageNumber="316">17.9</td>
<td id="76A95DA1192F52A63EF0FBD0EF84FB88" box="[1164,1209,1110,1132]" gridcol="5" gridrow="10" pageId="15" pageNumber="316">18.2</td>
<td id="76A95DA1192F52A63F60FBD0EEBDFB88" box="[1308,1408,1110,1132]" gridcol="6" gridrow="10" pageId="15" pageNumber="316">17.2 (1.0)</td>
</tr>
<tr id="357834DD192F52A63AE3FB06EEBDFB72" box="[159,1408,1152,1174]" gridrow="11" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FB06EAE5FB72" box="[159,472,1152,1174]" gridcol="0" gridrow="11" pageId="15" pageNumber="316">Anal-fin length 19.4</th>
<td id="76A95DA1192F52A63848FB06E95DFB72" box="[564,608,1152,1174]" gridcol="1" gridrow="11" pageId="15" pageNumber="316">17.2</td>
<td id="76A95DA1192F52A638C7FB06E9DAFB72" box="[699,743,1152,1174]" gridcol="2" gridrow="11" pageId="15" pageNumber="316">19.4</td>
<td id="76A95DA1192F52A6392FFB06E842FB72" box="[851,895,1152,1174]" gridcol="3" gridrow="11" pageId="15" pageNumber="316">19.2</td>
<td id="76A95DA1192F52A63988FB06EF1DFB72" box="[1012,1056,1152,1174]" gridcol="4" gridrow="11" pageId="15" pageNumber="316">17.2</td>
<td id="76A95DA1192F52A63EF0FB06EF84FB72" box="[1164,1209,1152,1174]" gridcol="5" gridrow="11" pageId="15" pageNumber="316">19.1</td>
<td id="76A95DA1192F52A63F60FB06EEBDFB72" box="[1308,1408,1152,1174]" gridcol="6" gridrow="11" pageId="15" pageNumber="316">18.4 (1.1)</td>
</tr>
<tr id="357834DD192F52A63AE3FB2FEEBDFB5B" box="[159,1408,1193,1215]" gridrow="12" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FB2FEAE5FB5B" box="[159,472,1193,1215]" gridcol="0" gridrow="12" pageId="15" pageNumber="316">Second dorsal-fin length 29.3</th>
<td id="76A95DA1192F52A63848FB2FE95DFB5B" box="[564,608,1193,1215]" gridcol="1" gridrow="12" pageId="15" pageNumber="316">30.5</td>
<td id="76A95DA1192F52A638C7FB2FE9DAFB5B" box="[699,743,1193,1215]" gridcol="2" gridrow="12" pageId="15" pageNumber="316">29.0</td>
<td id="76A95DA1192F52A6392FFB2FE842FB5B" box="[851,895,1193,1215]" gridcol="3" gridrow="12" pageId="15" pageNumber="316">28.6</td>
<td id="76A95DA1192F52A63988FB2FEF1DFB5B" box="[1012,1056,1193,1215]" gridcol="4" gridrow="12" pageId="15" pageNumber="316">28.9</td>
<td id="76A95DA1192F52A63EF0FB2FEF84FB5B" box="[1164,1209,1193,1215]" gridcol="5" gridrow="12" pageId="15" pageNumber="316">29.4</td>
<td id="76A95DA1192F52A63F60FB2FEEBDFB5B" box="[1308,1408,1193,1215]" gridcol="6" gridrow="12" pageId="15" pageNumber="316">29.3 (0.7)</td>
</tr>
<tr id="357834DD192F52A63AE3FB54EEBDFB0C" box="[159,1408,1234,1256]" gridrow="13" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FB54EAE5FB0C" box="[159,472,1234,1256]" gridcol="0" gridrow="13" pageId="15" pageNumber="316">Caudal-fin length 20.2</th>
<td id="76A95DA1192F52A63848FB54E95DFB0C" box="[564,608,1234,1256]" gridcol="1" gridrow="13" pageId="15" pageNumber="316">21.2</td>
<td id="76A95DA1192F52A638C7FB54E9DAFB0C" box="[699,743,1234,1256]" gridcol="2" gridrow="13" pageId="15" pageNumber="316">23.0</td>
<td id="76A95DA1192F52A6392FFB54E842FB0C" box="[851,895,1234,1256]" gridcol="3" gridrow="13" pageId="15" pageNumber="316">21.8</td>
<td id="76A95DA1192F52A63988FB54EF1DFB0C" box="[1012,1056,1234,1256]" gridcol="4" gridrow="13" pageId="15" pageNumber="316">20.9</td>
<td id="76A95DA1192F52A63EF0FB54EF84FB0C" box="[1164,1209,1234,1256]" gridcol="5" gridrow="13" pageId="15" pageNumber="316">22.7</td>
<td id="76A95DA1192F52A63F60FB54EEBDFB0C" box="[1308,1408,1234,1256]" gridcol="6" gridrow="13" pageId="15" pageNumber="316">21.9 (0.9)</td>
</tr>
<tr id="357834DD192F52A63AE3FB7AEEBDFAF6" box="[159,1408,1276,1298]" gridrow="14" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FB7AEAE5FAF6" box="[159,472,1276,1298]" gridcol="0" gridrow="14" pageId="15" pageNumber="316">Pectoral-fin length 19.1</th>
<td id="76A95DA1192F52A63848FB7AE95DFAF6" box="[564,608,1276,1298]" gridcol="1" gridrow="14" pageId="15" pageNumber="316">19.7</td>
<td id="76A95DA1192F52A638C7FB7AE9DAFAF6" box="[699,743,1276,1298]" gridcol="2" gridrow="14" pageId="15" pageNumber="316">d</td>
<td id="76A95DA1192F52A6392FFB7AE842FAF6" box="[851,895,1276,1298]" gridcol="3" gridrow="14" pageId="15" pageNumber="316">23.0</td>
<td id="76A95DA1192F52A63988FB7AEF1DFAF6" box="[1012,1056,1276,1298]" gridcol="4" gridrow="14" pageId="15" pageNumber="316">22.6</td>
<td id="76A95DA1192F52A63EF0FB7AEF84FAF6" box="[1164,1209,1276,1298]" gridcol="5" gridrow="14" pageId="15" pageNumber="316">25.0</td>
<td id="76A95DA1192F52A63F60FB7AEEBDFAF6" box="[1308,1408,1276,1298]" gridcol="6" gridrow="14" pageId="15" pageNumber="316">22.6 (2.2)</td>
</tr>
<tr id="357834DD192F52A63AE3FAA3EEBDFADF" box="[159,1408,1317,1339]" gridrow="15" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FAA3EAE5FADF" box="[159,472,1317,1339]" gridcol="0" gridrow="15" pageId="15" pageNumber="316">Caudal-peduncle length 22.8</th>
<td id="76A95DA1192F52A63848FAA3E95DFADF" box="[564,608,1317,1339]" gridcol="1" gridrow="15" pageId="15" pageNumber="316">21.6</td>
<td id="76A95DA1192F52A638C7FAA3E9DAFADF" box="[699,743,1317,1339]" gridcol="2" gridrow="15" pageId="15" pageNumber="316">22.5</td>
<td id="76A95DA1192F52A6392FFAA3E842FADF" box="[851,895,1317,1339]" gridcol="3" gridrow="15" pageId="15" pageNumber="316">21.8</td>
<td id="76A95DA1192F52A63988FAA3EF1DFADF" box="[1012,1056,1317,1339]" gridcol="4" gridrow="15" pageId="15" pageNumber="316">20.3</td>
<td id="76A95DA1192F52A63EF0FAA3EF84FADF" box="[1164,1209,1317,1339]" gridcol="5" gridrow="15" pageId="15" pageNumber="316">21.7</td>
<td id="76A95DA1192F52A63F60FAA3EEBDFADF" box="[1308,1408,1317,1339]" gridcol="6" gridrow="15" pageId="15" pageNumber="316">21.6 (0.8)</td>
</tr>
<tr id="357834DD192F52A63AE3FAC8EEBDFA80" box="[159,1408,1358,1380]" gridrow="16" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FAC8EAE5FA80" box="[159,472,1358,1380]" gridcol="0" gridrow="16" pageId="15" pageNumber="316">Caudal-peduncle depth 14.2</th>
<td id="76A95DA1192F52A63848FAC8E95DFA80" box="[564,608,1358,1380]" gridcol="1" gridrow="16" pageId="15" pageNumber="316">13.8</td>
<td id="76A95DA1192F52A638C7FAC8E9DAFA80" box="[699,743,1358,1380]" gridcol="2" gridrow="16" pageId="15" pageNumber="316">14.3</td>
<td id="76A95DA1192F52A6392FFAC8E842FA80" box="[851,895,1358,1380]" gridcol="3" gridrow="16" pageId="15" pageNumber="316">14.7</td>
<td id="76A95DA1192F52A63988FAC8EF1DFA80" box="[1012,1056,1358,1380]" gridcol="4" gridrow="16" pageId="15" pageNumber="316">13.9</td>
<td id="76A95DA1192F52A63EF0FAC8EF84FA80" box="[1164,1209,1358,1380]" gridcol="5" gridrow="16" pageId="15" pageNumber="316">14.5</td>
<td id="76A95DA1192F52A63F60FAC8EEBDFA80" box="[1308,1408,1358,1380]" gridcol="6" gridrow="16" pageId="15" pageNumber="316">14.2 (0.4)</td>
</tr>
<tr id="357834DD192F52A63AE3FAFEEEBDFA6A" box="[159,1408,1400,1422]" gridrow="17" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FAFEEAE5FA6A" box="[159,472,1400,1422]" gridcol="0" gridrow="17" pageId="15" pageNumber="316">Interorbital width 16.3</th>
<td id="76A95DA1192F52A63848FAFEE95DFA6A" box="[564,608,1400,1422]" gridcol="1" gridrow="17" pageId="15" pageNumber="316">15.0</td>
<td id="76A95DA1192F52A638C7FAFEE9DAFA6A" box="[699,743,1400,1422]" gridcol="2" gridrow="17" pageId="15" pageNumber="316">17.0</td>
<td id="76A95DA1192F52A6392FFAFEE842FA6A" box="[851,895,1400,1422]" gridcol="3" gridrow="17" pageId="15" pageNumber="316">16.3</td>
<td id="76A95DA1192F52A63988FAFEEF1DFA6A" box="[1012,1056,1400,1422]" gridcol="4" gridrow="17" pageId="15" pageNumber="316">16.3</td>
<td id="76A95DA1192F52A63EF0FAFEEF84FA6A" box="[1164,1209,1400,1422]" gridcol="5" gridrow="17" pageId="15" pageNumber="316">17.9</td>
<td id="76A95DA1192F52A63F60FAFEEEBDFA6A" box="[1308,1408,1400,1422]" gridcol="6" gridrow="17" pageId="15" pageNumber="316">16.5 (1.0)</td>
</tr>
<tr id="357834DD192F52A63AE3FA27EEBDFA53" box="[159,1408,1441,1463]" gridrow="18" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FA27EAE5FA53" box="[159,472,1441,1463]" gridcol="0" gridrow="18" pageId="15" pageNumber="316">Horizontal eye diameter 19.3</th>
<td id="76A95DA1192F52A63848FA27E95DFA53" box="[564,608,1441,1463]" gridcol="1" gridrow="18" pageId="15" pageNumber="316">20.9</td>
<td id="76A95DA1192F52A638C7FA27E9DAFA53" box="[699,743,1441,1463]" gridcol="2" gridrow="18" pageId="15" pageNumber="316">21.4</td>
<td id="76A95DA1192F52A6392FFA27E842FA53" box="[851,895,1441,1463]" gridcol="3" gridrow="18" pageId="15" pageNumber="316">19.3</td>
<td id="76A95DA1192F52A63988FA27EF1DFA53" box="[1012,1056,1441,1463]" gridcol="4" gridrow="18" pageId="15" pageNumber="316">21.3</td>
<td id="76A95DA1192F52A63EF0FA27EF84FA53" box="[1164,1209,1441,1463]" gridcol="5" gridrow="18" pageId="15" pageNumber="316">23.4</td>
<td id="76A95DA1192F52A63F60FA27EEBDFA53" box="[1308,1408,1441,1463]" gridcol="6" gridrow="18" pageId="15" pageNumber="316">21.3 (1.5)</td>
</tr>
<tr id="357834DD192F52A63AE3FA4CEEBDFA04" box="[159,1408,1482,1504]" gridrow="19" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FA4CEAE5FA04" box="[159,472,1482,1504]" gridcol="0" gridrow="19" pageId="15" pageNumber="316">Snout length 27.8</th>
<td id="76A95DA1192F52A63848FA4CE95DFA04" box="[564,608,1482,1504]" gridcol="1" gridrow="19" pageId="15" pageNumber="316">26.6</td>
<td id="76A95DA1192F52A638C7FA4CE9DAFA04" box="[699,743,1482,1504]" gridcol="2" gridrow="19" pageId="15" pageNumber="316">29.2</td>
<td id="76A95DA1192F52A6392FFA4CE842FA04" box="[851,895,1482,1504]" gridcol="3" gridrow="19" pageId="15" pageNumber="316">26.6</td>
<td id="76A95DA1192F52A63988FA4CEF1DFA04" box="[1012,1056,1482,1504]" gridcol="4" gridrow="19" pageId="15" pageNumber="316">28.3</td>
<td id="76A95DA1192F52A63EF0FA4CEF84FA04" box="[1164,1209,1482,1504]" gridcol="5" gridrow="19" pageId="15" pageNumber="316">28.0</td>
<td id="76A95DA1192F52A63F60FA4CEEBDFA04" box="[1308,1408,1482,1504]" gridcol="6" gridrow="19" pageId="15" pageNumber="316">27.7 (1.1)</td>
</tr>
<tr id="357834DD192F52A63AE3FA72EEBDF9EE" box="[159,1408,1524,1546]" gridrow="20" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3FA72EAE5F9EE" box="[159,472,1524,1546]" gridcol="0" gridrow="20" pageId="15" pageNumber="316">Upper-jaw length 34.9</th>
<td id="76A95DA1192F52A63848FA72E95DF9EE" box="[564,608,1524,1546]" gridcol="1" gridrow="20" pageId="15" pageNumber="316">36.6</td>
<td id="76A95DA1192F52A638C7FA72E9DAF9EE" box="[699,743,1524,1546]" gridcol="2" gridrow="20" pageId="15" pageNumber="316">36.4</td>
<td id="76A95DA1192F52A6392FFA72E842F9EE" box="[851,895,1524,1546]" gridcol="3" gridrow="20" pageId="15" pageNumber="316">35.1</td>
<td id="76A95DA1192F52A63988FA72EF1DF9EE" box="[1012,1056,1524,1546]" gridcol="4" gridrow="20" pageId="15" pageNumber="316">38.0</td>
<td id="76A95DA1192F52A63EF0FA72EF84F9EE" box="[1164,1209,1524,1546]" gridcol="5" gridrow="20" pageId="15" pageNumber="316">36.5</td>
<td id="76A95DA1192F52A63F60FA72EEBDF9EE" box="[1308,1408,1524,1546]" gridcol="6" gridrow="20" pageId="15" pageNumber="316">36.5 (1.0)</td>
</tr>
<tr id="357834DD192F52A63AE3F99BEEBDF9B6" box="[159,1408,1565,1618]" gridrow="21" pageId="15" pageNumber="316">
<th id="76A95DA1192F52A63AE3F99BEAE5F9B6" box="[159,472,1565,1618]" gridcol="0" gridrow="21" pageId="15" pageNumber="316">First dorsal spine to first 51.5 pectoral ray</th>
<td id="76A95DA1192F52A63848F99BE95DF9B6" box="[564,608,1565,1618]" gridcol="1" gridrow="21" pageId="15" pageNumber="316">53.9</td>
<td id="76A95DA1192F52A638C7F99BE9DAF9B6" box="[699,743,1565,1618]" gridcol="2" gridrow="21" pageId="15" pageNumber="316">52.1</td>
<td id="76A95DA1192F52A6392FF99BE842F9B6" box="[851,895,1565,1618]" gridcol="3" gridrow="21" pageId="15" pageNumber="316">54.3</td>
<td id="76A95DA1192F52A63988F99BEF1DF9B6" box="[1012,1056,1565,1618]" gridcol="4" gridrow="21" pageId="15" pageNumber="316">62.7</td>
<td id="76A95DA1192F52A63EF0F99BEF84F9B6" box="[1164,1209,1565,1618]" gridcol="5" gridrow="21" pageId="15" pageNumber="316">48.4</td>
<td id="76A95DA1192F52A63F60F99BEEBDF9B6" box="[1308,1408,1565,1618]" gridcol="6" gridrow="21" pageId="15" pageNumber="316">54.3 (5.2)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF7369F192FAD563ABBF917EBF2F916" blockId="15.[151,1436,1681,2030]" pageId="15" pageNumber="316">
<emphasis id="B93CEA8D192FAD563ABBF917EAF3F94E" bold="true" box="[199,462,1681,1706]" pageId="15" pageNumber="316">Preserved colouration</emphasis>
. After formalin fixation and ethanol preservation, the general colour is dark brownishgrey to black, eyes opaque. Some patches on the cheek, pectoral and median fins may appear lighter (
<figureCitation id="13732A1A192FAD563F73F933EE6CF92A" box="[1295,1361,1717,1742]" captionStart="FIGURE 5" captionStartId="14.[151,250,1138,1160]" captionTargetBox="[179,1407,195,1113]" captionTargetId="figure@14.[179,1408,195,1113]" captionTargetPageId="14" captionText="FIGURE 5. Gobiodon ater sp. nov., adult, holotype NMW 94612, 26.5 mm SL, Dahab, Egypt, Red Sea. A. freshly collected; B. ethanol-preserved (epidermal mucus partly removed). Photos by J. Herler." httpUri="https://zenodo.org/record/249075/files/figure.png" pageId="15" pageNumber="316">Fig. 5</figureCitation>
B, 6A, 6C).
</paragraph>
<paragraph id="8BF7369F192FAD563ABBF97BEBC4F80A" blockId="15.[151,1436,1681,2030]" pageId="15" pageNumber="316">
<emphasis id="B93CEA8D192FAD563ABBF97BEA8EF8F2" bold="true" box="[199,435,1789,1814]" pageId="15" pageNumber="316">Molecular genetics.</emphasis>
The present analysis includes the sequence of the a: 
<typeStatus id="54F3883D192FAD563E51F978EFB3F8F2" box="[1069,1166,1790,1814]" pageId="15" pageNumber="316" type="paratype">paratype</typeStatus>
MNHN 2012-0111 (as G.sp.2_RN1) from the northern Red Sea (see 
<tableCitation id="C6CA0324192FAD5638E2F8A7E9CFF8DE" box="[670,754,1825,1850]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionTargetBox="[159,1431,362,1898]" captionTargetPageId="4" captionText="TABLE 1. GenBank accession numbers (GB No.) and references for published sequences (12 S and 16 S rRNA) of 25 taxa (4 new species are underlined) of Gobiodon (G.) and Paragobiodon (P.). Localities: GBR = Great Barrier Reef, Mal = Maldives, PNG = Papua New Guinea, RS = Red Sea, Tai = Taiwan. Abbreviations (PM) and (JH) refer to the first demonstration of sequenced but undescribed species, Munday et al. (1999) and Herler and Hilgers (2009), respectively. Superscript numbers in locality code indicate type status: a: Paratype MNHN 2012 - 0111, b: Paratype BMNH 2006.10. 6.1, c: Paratype NMW 95565." httpUri="http://table.plazi.org/id/DF3766171924AD5D3AEBFF1EE90BFEAB" pageId="15" pageNumber="316" tableUuid="DF3766171924AD5D3AEBFF1EE90BFEAB">Table 1</tableCitation>
for Genbank accession numbers). Derived from the present analysis, the genetically closest species are 
<taxonomicName id="4C484D1C192FAD5638F4F8C0E83BF8B9" box="[648,774,1861,1885]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="axillaris">
<emphasis id="B93CEA8D192FAD5638F4F8C0E83BF8B9" box="[648,774,1861,1885]" italics="true" pageId="15" pageNumber="316">G. axillaris</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192FAD56393CF8C0E8E5F8B9" box="[832,984,1861,1885]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="fuscoruber">
<emphasis id="B93CEA8D192FAD56393CF8C0E8E5F8B9" box="[832,984,1861,1885]" italics="true" pageId="15" pageNumber="316">G. fuscoruber</emphasis>
</taxonomicName>
. The 
<emphasis id="B93CEA8D192FAD563E60F8C0EF17F8B9" box="[1052,1066,1862,1885]" italics="true" pageId="15" pageNumber="316">p</emphasis>
-distance between 
<taxonomicName id="4C484D1C192FAD563E81F8C0EE77F8B9" box="[1277,1354,1862,1885]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D192FAD563E81F8C0EE77F8B9" box="[1277,1354,1862,1885]" italics="true" pageId="15" pageNumber="316">G. ater</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192FAD563FF8F8C0EA2DF865" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="fuscoruber">
<emphasis id="B93CEA8D192FAD563FF8F8C0EA2DF865" italics="true" pageId="15" pageNumber="316">G. fuscoruber</emphasis>
</taxonomicName>
in the present study ranges from 0.023 to 0.032 (
<tableCitation id="C6CA0324192FAD563948F8EFE8A4F866" box="[820,921,1897,1922]" captionStart="TABLE 10" captionStartId="21.[151,239,670,693]" captionTargetBox="[159,1393,234,607]" captionTargetPageId="21" captionText="TABLE 10. Selection of genetic (p -) distances from pairwise comparisons of specimens of G. bilineatus sp. nov., G. irregularis sp. nov., G. ater sp. nov. and G. fuscoruber sp. nov. from different regions (GBR: Great Barrier Reef, Jap: Japan, Mal: Maldives, RS: Red Sea, Tai: Taiwan). For comparison of intra- and interspecific genetic distances across the genus, other species from different regions as well as “ G. unicolor ” (sensu Munday et al. (1999) and Harold et al. (2008 )) from the GBR are included. Italic numbers indicate intraspecific distances. Bold values indicate intraspecific distances of more than 2 %. For species abbreviations and Genbank accession numbers see Table 1." httpUri="http://table.plazi.org/id/DF3766171935AD4C3AEBFD18EF0CFCB1" pageId="15" pageNumber="316" tableUuid="DF3766171935AD4C3AEBFD18EF0CFCB1">Table 10</tableCitation>
), making up a genetic difference of more than 2%. 
<taxonomicName id="4C484D1C192FAD563AB7F80BEAA2F841" box="[203,415,1933,1957]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="axillaris">
<emphasis id="B93CEA8D192FAD563AB7F80BEAA2F841" box="[203,415,1933,1957]" italics="true" pageId="15" pageNumber="316">Gobiodon axillaris</emphasis>
</taxonomicName>
seems genetically even closer to 
<taxonomicName id="4C484D1C192FAD563964F808E85EF841" box="[792,867,1934,1957]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="316" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D192FAD563964F808E85EF841" box="[792,867,1934,1957]" italics="true" pageId="15" pageNumber="316">G. ater</emphasis>
</taxonomicName>
, but since the NJ-bootstrap value between the two was very low (21), this relationship is highly questionable and requires a more detailed genetic and morphological analysis.
</paragraph>
<caption id="DF3766171930AD493AEBFF1EEE31FF4B" ID-Table-UUID="DF3766171930AD493AEBFF1EEE31FF4B" box="[151,1292,151,175]" httpUri="http://table.plazi.org/id/DF3766171930AD493AEBFF1EEE31FF4B" pageId="16" pageNumber="317" targetBox="[159,1386,202,410]" targetIsTable="true" targetPageId="16">
<paragraph id="8BF7369F1930AD493AEBFF1EEE31FF4B" blockId="16.[151,1292,151,175]" box="[151,1292,151,175]" pageId="16" pageNumber="317">
<emphasis id="B93CEA8D1930AD493AEBFF1EEA35FF4B" bold="true" box="[151,264,152,175]" pageId="16" pageNumber="317">TABLE 7.</emphasis>
Fin counts of holotype and five paratypes of 
<taxonomicName id="4C484D1C1930AD493893FF11E8B6FF4B" box="[751,907,151,175]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater" status="sp. nov.">
<emphasis id="B93CEA8D1930AD493893FF11E8B6FF4B" box="[751,907,151,175]" italics="true" pageId="16" pageNumber="317">Gobiodon ater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61930AD4939EEFF1EE8D9FF4B" box="[914,996,152,175]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
from the northern Red Sea.
</paragraph>
</caption>
<paragraph id="8BF7369F1930AD493AE3FF4CEE36FE7E" blockId="16.[159,1386,202,575]" pageId="16" pageNumber="317">
<table id="F948C43F193052A63AE3FF4CEE57FE7E" box="[159,1386,202,410]" gridcols="4" gridrows="4" pageId="16" pageNumber="317">
<tr id="357834DD193052A63AE3FF4CEE57FEFA" box="[159,1386,202,286]" gridrow="0" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FF4CEBCAFEFA" box="[159,247,202,286]" gridcol="0" gridrow="0" pageId="16" pageNumber="317">Status Coll.No.</th>
<th id="76A95DA1193052A63B29FF4CE837FEFA" box="[341,778,202,286]" gridcol="1" gridrow="0" pageId="16" pageNumber="317">Holotype Paratype Paratype NMW 94612 NMW 94613 BMNH 2012.3.20.1</th>
<th id="76A95DA1193052A63922FF4CE8EAFEFA" box="[862,983,202,286]" gridcol="2" gridrow="0" pageId="16" pageNumber="317">Paratype BMNH 2012.3.20.2</th>
<th id="76A95DA1193052A63E52FF4CEE57FEFA" box="[1070,1386,202,286]" gridcol="3" gridrow="0" pageId="16" pageNumber="317">Paratype Paratype MNHN 2012-0110 MNHN 2012-0111</th>
</tr>
<tr id="357834DD193052A63AE3FEB7EE57FEA3" box="[159,1386,305,327]" gridrow="1" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FEB7EBCAFEA3" box="[159,247,305,327]" gridcol="0" gridrow="1" pageId="16" pageNumber="317">D1</th>
<td id="76A95DA1193052A63B29FEB7E837FEA3" box="[341,778,305,327]" gridcol="1" gridrow="1" pageId="16" pageNumber="317">VI VI VI</td>
<td id="76A95DA1193052A63922FEB7E8EAFEA3" box="[862,983,305,327]" gridcol="2" gridrow="1" pageId="16" pageNumber="317">VI</td>
<td id="76A95DA1193052A63E52FEB7EE57FEA3" box="[1070,1386,305,327]" gridcol="3" gridrow="1" pageId="16" pageNumber="317">VI VI</td>
</tr>
<tr id="357834DD193052A63AE3FEDCEE57FE94" box="[159,1386,346,368]" gridrow="2" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FEDCEBCAFE94" box="[159,247,346,368]" gridcol="0" gridrow="2" pageId="16" pageNumber="317">D2</th>
<td id="76A95DA1193052A63B29FEDCE837FE94" box="[341,778,346,368]" gridcol="1" gridrow="2" pageId="16" pageNumber="317">10 10 10</td>
<td id="76A95DA1193052A63922FEDCE8EAFE94" box="[862,983,346,368]" gridcol="2" gridrow="2" pageId="16" pageNumber="317">10</td>
<td id="76A95DA1193052A63E52FEDCEE57FE94" box="[1070,1386,346,368]" gridcol="3" gridrow="2" pageId="16" pageNumber="317">10 10</td>
</tr>
<tr id="357834DD193052A63AE3FE02EE57FE7E" box="[159,1386,388,410]" gridrow="3" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FE02EBCAFE7E" box="[159,247,388,410]" gridcol="0" gridrow="3" pageId="16" pageNumber="317">A</th>
<td id="76A95DA1193052A63B29FE02E837FE7E" box="[341,778,388,410]" gridcol="1" gridrow="3" pageId="16" pageNumber="317">8 8 8</td>
<td id="76A95DA1193052A63922FE02E8EAFE7E" box="[862,983,388,410]" gridcol="2" gridrow="3" pageId="16" pageNumber="317">8</td>
<td id="76A95DA1193052A63E52FE02EE57FE7E" box="[1070,1386,388,410]" gridcol="3" gridrow="3" pageId="16" pageNumber="317">8 8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF7369F1930AD493AE3FE2BEE79FDDB" blockId="16.[159,1386,202,575]" pageId="16" pageNumber="317">
<table id="F948C43F193052A63AE3FE2BEE57FDDB" box="[159,1386,429,575]" gridcols="6" gridrows="4" pageId="16" pageNumber="317">
<tr id="357834DD193052A63AE3FE2BEE57FE27" box="[159,1386,429,451]" gridrow="0" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FE2BEAA1FE27" box="[159,412,429,451]" gridcol="0" gridrow="0" pageId="16" pageNumber="317">C (segmented) 17</th>
<th id="76A95DA1193052A63B8EFE2BE905FE27" box="[498,568,429,451]" gridcol="1" gridrow="0" pageId="16" pageNumber="317">17</th>
<th id="76A95DA1193052A638EEFE2BE9E5FE27" box="[658,728,429,451]" gridcol="2" gridrow="0" pageId="16" pageNumber="317">17</th>
<th id="76A95DA1193052A63922FE2BE899FE27" box="[862,932,429,451]" gridcol="3" gridrow="0" pageId="16" pageNumber="317">17</th>
<th id="76A95DA1193052A63E52FE2BEF49FE27" box="[1070,1140,429,451]" gridcol="4" gridrow="0" pageId="16" pageNumber="317">16</th>
<th id="76A95DA1193052A63E82FE2BEE57FE27" box="[1278,1386,429,451]" gridcol="5" gridrow="0" pageId="16" pageNumber="317">17</th>
</tr>
<tr id="357834DD193052A63AE3FE50EE57FE08" box="[159,1386,470,492]" gridrow="1" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FE50EAA1FE08" box="[159,412,470,492]" gridcol="0" gridrow="1" pageId="16" pageNumber="317">C (branched) 17</th>
<td id="76A95DA1193052A63B8EFE50E905FE08" box="[498,568,470,492]" gridcol="1" gridrow="1" pageId="16" pageNumber="317">17</td>
<td id="76A95DA1193052A638EEFE50E9E5FE08" box="[658,728,470,492]" gridcol="2" gridrow="1" pageId="16" pageNumber="317">17</td>
<td id="76A95DA1193052A63922FE50E899FE08" box="[862,932,470,492]" gridcol="3" gridrow="1" pageId="16" pageNumber="317">17</td>
<td id="76A95DA1193052A63E52FE50EF49FE08" box="[1070,1140,470,492]" gridcol="4" gridrow="1" pageId="16" pageNumber="317">16</td>
<td id="76A95DA1193052A63E82FE50EE57FE08" box="[1278,1386,470,492]" gridcol="5" gridrow="1" pageId="16" pageNumber="317">17</td>
</tr>
<tr id="357834DD193052A63AE3FD86EE57FDF2" box="[159,1386,512,534]" gridrow="2" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FD86EAA1FDF2" box="[159,412,512,534]" gridcol="0" gridrow="2" pageId="16" pageNumber="317">P 19</th>
<td id="76A95DA1193052A63B8EFD86E905FDF2" box="[498,568,512,534]" gridcol="1" gridrow="2" pageId="16" pageNumber="317">19</td>
<td id="76A95DA1193052A638EEFD86E9E5FDF2" box="[658,728,512,534]" gridcol="2" gridrow="2" pageId="16" pageNumber="317">19</td>
<td id="76A95DA1193052A63922FD86E899FDF2" box="[862,932,512,534]" gridcol="3" gridrow="2" pageId="16" pageNumber="317">19</td>
<td id="76A95DA1193052A63E52FD86EF49FDF2" box="[1070,1140,512,534]" gridcol="4" gridrow="2" pageId="16" pageNumber="317">20</td>
<td id="76A95DA1193052A63E82FD86EE57FDF2" box="[1278,1386,512,534]" gridcol="5" gridrow="2" pageId="16" pageNumber="317">19</td>
</tr>
<tr id="357834DD193052A63AE3FDAFEE57FDDB" box="[159,1386,553,575]" gridrow="3" pageId="16" pageNumber="317">
<th id="76A95DA1193052A63AE3FDAFEAA1FDDB" box="[159,412,553,575]" gridcol="0" gridrow="3" pageId="16" pageNumber="317">V I/5+I/5</th>
<td id="76A95DA1193052A63B8EFDAFE905FDDB" box="[498,568,553,575]" gridcol="1" gridrow="3" pageId="16" pageNumber="317">I/5+I/5</td>
<td id="76A95DA1193052A638EEFDAFE9E5FDDB" box="[658,728,553,575]" gridcol="2" gridrow="3" pageId="16" pageNumber="317">I/5+I/5</td>
<td id="76A95DA1193052A63922FDAFE899FDDB" box="[862,932,553,575]" gridcol="3" gridrow="3" pageId="16" pageNumber="317">I/5+I/5</td>
<td id="76A95DA1193052A63E52FDAFEF49FDDB" box="[1070,1140,553,575]" gridcol="4" gridrow="3" pageId="16" pageNumber="317">I/5+I/5</td>
<td id="76A95DA1193052A63E82FDAFEE57FDDB" box="[1278,1386,553,575]" gridcol="5" gridrow="3" pageId="16" pageNumber="317">I/5+I/5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF7369F1930AD493ABBFD02EF3FFCC9" blockId="16.[151,1436,644,2037]" pageId="16" pageNumber="317">
<emphasis id="B93CEA8D1930AD493ABBFD02EA17FD79" bold="true" box="[199,298,644,669]" pageId="16" pageNumber="317">Habitat.</emphasis>
<taxonomicName id="4C484D1C1930AD493B4FFD02EAE7FD78" box="[307,474,644,668]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493B4FFD02EAE7FD78" box="[307,474,644,668]" italics="true" pageId="16" pageNumber="317">Gobiodon ater</emphasis>
</taxonomicName>
is highly specialized and prefers fine-branched corals such as 
<taxonomicName id="4C484D1C1930AD493ED1FD02EE57FD78" box="[1197,1386,644,668]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="selago">
<emphasis id="B93CEA8D1930AD493ED1FD02EE57FD78" box="[1197,1386,644,668]" italics="true" pageId="16" pageNumber="317">Acropora selago</emphasis>
</taxonomicName>
and small tabulate colonies of 
<taxonomicName id="4C484D1C1930AD493BC7FD2EE96EFD24" box="[443,595,680,704]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="hyacinthus">
<emphasis id="B93CEA8D1930AD493BC7FD2EE96EFD24" box="[443,595,680,704]" italics="true" pageId="16" pageNumber="317">A. hyacinthus</emphasis>
</taxonomicName>
. Less often, it also occurs in other 
<taxonomicName id="4C484D1C1930AD4939A5FD2EEF7EFD5B" box="[985,1091,680,703]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="genus">
<emphasis id="B93CEA8D1930AD4939A5FD2EEF7EFD5B" box="[985,1091,680,703]" italics="true" pageId="16" pageNumber="317">Acropora</emphasis>
</taxonomicName>
species, such as 
<taxonomicName id="4C484D1C1930AD493F7CFD2EEEA8FD5B" box="[1280,1429,680,703]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="acuminata">
<emphasis id="B93CEA8D1930AD493F7CFD2EEEA8FD5B" box="[1280,1429,680,703]" italics="true" pageId="16" pageNumber="317">A. acuminata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493AEBFD4AEA11FD00" box="[151,300,716,740]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="samoensis">
<emphasis id="B93CEA8D1930AD493AEBFD4AEA11FD00" box="[151,300,716,740]" italics="true" pageId="16" pageNumber="317">A. samoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493B41FD4AEAE8FD00" box="[317,469,716,740]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="eurystoma">
<emphasis id="B93CEA8D1930AD493B41FD4AEAE8FD00" box="[317,469,716,740]" italics="true" pageId="16" pageNumber="317">A. eurystoma</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1930AD49386EFD4AE8E9FD00" authority="Dirnwoeber &amp; Herler 2007" authorityName="Dirnwoeber &amp; Herler" authorityYear="2007" box="[530,980,716,741]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="species" species="valida">
<emphasis id="B93CEA8D1930AD49386EFD4AE947FD00" box="[530,634,716,740]" italics="true" pageId="16" pageNumber="317">A. valida</emphasis>
(
<bibRefCitation id="EFD94B6E1930AD4938F0FD4AE8F0FD00" author="Dirnwoeber" box="[652,973,716,741]" pageId="16" pageNumber="317" refString="Dirnwoeber, M. &amp; Herler, J. (2007) Microhabitat specialisation and ecological consequences for coral gobies of the genus Gobiodon in the Gulf of Aqaba, northern Red Sea. Marine Ecology Progress Series, 342, 265 - 275." type="journal article" year="2007">Dirnwoeber &amp; Herler 2007</bibRefCitation>
)
</taxonomicName>
. The species is found in shallow water regions such as protected reef flats and reef crests. Due to its black colouration it is sometimes difficult to detect this species within the dark interstices of narrow-branched 
<taxonomicName id="4C484D1C1930AD493950FC92E8AAFCCF" box="[812,919,788,811]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="16" pageNumber="317" phylum="Cnidaria" rank="genus">
<emphasis id="B93CEA8D1930AD493950FC92E8AAFCCF" box="[812,919,788,811]" italics="true" pageId="16" pageNumber="317">Acropora</emphasis>
</taxonomicName>
colonies.
</paragraph>
</subSubSection>
<subSubSection id="C35265141930AD493ABBFCB1EEA1FCB4" box="[199,1436,823,848]" pageId="16" pageNumber="317" type="distribution">
<paragraph id="8BF7369F1930AD493ABBFCB1EEA1FCB4" blockId="16.[151,1436,644,2037]" box="[199,1436,823,848]" pageId="16" pageNumber="317">
<emphasis id="B93CEA8D1930AD493ABBFCB1EA5DFCB4" bold="true" box="[199,352,823,848]" pageId="16" pageNumber="317">Distribution.</emphasis>
So far this species is known from the Red Sea, the central Indian and the western Pacific Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C35265141930AD493ABBFCDAE8EEFC7C" pageId="16" pageNumber="317" type="etymology">
<paragraph id="8BF7369F1930AD493ABBFCDAE8EEFC7C" blockId="16.[151,1436,644,2037]" pageId="16" pageNumber="317">
<emphasis id="B93CEA8D1930AD493ABBFCDAEA73FC91" bold="true" box="[199,334,860,885]" pageId="16" pageNumber="317">Etymology.</emphasis>
This species is named after its uniformly black life colouration (“ 
<taxonomicName id="4C484D1C1930AD493E4EFCDAEF5CFC90" box="[1074,1121,860,884]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">ater</taxonomicName>
” = Latin word for “black”), which even includes the eye. Suggested common name: Black Coralgoby.
</paragraph>
</subSubSection>
<subSubSection id="C35265141930AD483ABBFC22E8BCFE80" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="discussion">
<paragraph id="8BF7369F1930AD483ABBFC22E8BCFE80" blockId="16.[151,1436,644,2037]" lastBlockId="17.[151,1436,151,357]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">
<emphasis id="B93CEA8D1930AD493ABBFC22EA06FC59" bold="true" box="[199,315,932,957]" pageId="16" pageNumber="317">Remarks.</emphasis>
This species was designated as 
<taxonomicName id="4C484D1C1930AD4938CBFC22E81AFC58" box="[695,807,932,956]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D1930AD4938CBFC22E81AFC58" box="[695,807,932,956]" italics="true" pageId="16" pageNumber="317">Gobiodon</emphasis>
</taxonomicName>
sp. 2 by 
<bibRefCitation id="EFD94B6E1930AD4939E7FC22EFF5FC59" author="Herler" box="[923,1224,932,957]" pageId="16" pageNumber="317" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler and Hilgers (2005)</bibRefCitation>
and these authors assumed that it may be identical with one of the other entirely black species from the western Pacific, in particular with 
<taxonomicName id="4C484D1C1930AD493AA9FC6AEA45FBE0" box="[213,376,1004,1028]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD493AA9FC6AEA45FBE0" box="[213,376,1004,1028]" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
. The 
<typeStatus id="54F3883D1930AD493BB8FC6BE91BFBE1" box="[452,550,1005,1029]" pageId="16" pageNumber="317" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C484D1C1930AD49382AFC6AE9C5FBE0" box="[598,760,1004,1028]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD49382AFC6AE9C5FBE0" box="[598,760,1004,1028]" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
cannot be identified from the more than 
<specimenCount id="9D4EFD161930AD493E8DFC6AEEA6FBE1" box="[1265,1435,1004,1029]" pageId="16" pageNumber="317" type="generic">100 specimens</specimenCount>
collected by Bleeker (Ronald de Ruiter, pers. comm.), but when compared with the original description of 
<bibRefCitation id="EFD94B6E1930AD493F39FB89EBDEFBA9" author="Bleeker" pageId="16" pageNumber="317" refString="Bleeker, P. (1853) Nieuwe bijdrage tot de kennis der ichthyologische fauna van Ceram. Natuurkundig Tijdschrift Nederlandsch Indie, 3 (5), 689 - 714." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
, 
<taxonomicName id="4C484D1C1930AD493A8DFBB2EAADFBA8" box="[241,400,1076,1100]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD493A8DFBB2EAADFBA8" box="[241,400,1076,1100]" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
has a higher fin-ray A count (9–10 rays versus 8), a lower P count (17 versus 19–20) and larger eyes (25 versus 21% of head length) than 
<taxonomicName id="4C484D1C1930AD4938B8FBDEE82DFB94" box="[708,784,1112,1136]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD4938B8FBDEE82DFB94" box="[708,784,1112,1136]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
. The fin meristics of 
<bibRefCitation id="EFD94B6E1930AD493E76FBD1EF89FB94" author="Bleeker" box="[1034,1204,1111,1136]" pageId="16" pageNumber="317" refString="Bleeker, P. (1853) Nieuwe bijdrage tot de kennis der ichthyologische fauna van Ceram. Natuurkundig Tijdschrift Nederlandsch Indie, 3 (5), 689 - 714." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
are corroborated by specimens from 
<collectingCountry id="F35F760F1930AD493B29FBFBEAA5FB70" box="[341,408,1149,1172]" name="Japan" pageId="16" pageNumber="317">Japan</collectingCountry>
, which are considered to represent 
<taxonomicName id="4C484D1C1930AD49394CFBFAE8F2FB70" box="[816,975,1148,1172]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD49394CFBFAE8F2FB70" box="[816,975,1148,1172]" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
(
<figureCitation id="13732A1A1930AD49399DFBFAEF1BFB70" box="[993,1062,1148,1173]" captionStart="FIGURE 9" captionStartId="22.[151,250,1660,1682]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure@22.[151,1436,181,1641]" captionTargetPageId="22" captionText="FIGURE 9. Comparative material. Freshly collected adults: Gobiodon cf. bilineatus sp. nov. A. uncatalogued, 28.9 mm SL, Kagi Island, Maldives. B. uncatalogued, 22.9 mm SL, Kenting, southern Taiwan. C. Gobiodon prolixus, ROM 73338, holotype, male 26.2 mm SL, Nha Trang, Vietnam. D. Gobiodon sp. D sensu Munday et al. (1999), uncatalogued, 29.9 mm SL, Hembadhu Island, Maldives E. Gobiodon quinquestrigatus, uncatalogued, 28.7 mm SL, Kenting, southern Taiwan; F. Gobiodon cf. fulvus (sensu Winterbottom &amp; Emery 1986), OMNH P 40259, 20.8 mm SL, Iriomote Island, Japan. G. Gobiodon oculolineatus, OMNH P 40260, 22.6 mm SL, Okinawa Island, Japan. H. Gobiodon ceramensis, OMNH P 34042, 35 mm SL, Ryukyu Island, Japan. Gobiodon unicolor, preserved holotype, MNHN A- 4015, 31 mm SL, I. entire specimen. J. close-up of black opercular spot (arrow) on the right side. Photos by J. Herler (A, B, D, E, I, J), R. Winterbottom (C), T. Suzuki (F-H)." httpUri="https://zenodo.org/record/249079/files/figure.png" pageId="16" pageNumber="317">Fig. 9</figureCitation>
H) by one of us (TS); these also have fin-ray counts of D 
<date id="FFF6105F1930AD493BD7FB26EAF7FB5C" box="[427,458,1184,1208]" pageId="16" pageNumber="317">2 I</date>
,10 and A I,9, the range recorded by 
<bibRefCitation id="EFD94B6E1930AD493915FB19EF2BFB5C" author="Bleeker" box="[873,1046,1183,1208]" pageId="16" pageNumber="317" refString="Bleeker, P. (1853) Nieuwe bijdrage tot de kennis der ichthyologische fauna van Ceram. Natuurkundig Tijdschrift Nederlandsch Indie, 3 (5), 689 - 714." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
. Furthermore, according to 
<bibRefCitation id="EFD94B6E1930AD493F32FB19EA14FB38" pageId="16" pageNumber="317" refString="Harold, A. S., Winterbottom, R., Munday, P. L. &amp; Chapman, R. W. (2008) Phylogenetic relationships of Indo-pacific coral gobies of the genus Gobiodon (Teleostei: Gobiidae), based on morphological and molecular data. Bulletin of Marine Science, 82, 119 - 136." type="journal article">
Harold 
<emphasis id="B93CEA8D1930AD493AEBFB43EBF6FB38" box="[151,203,1220,1244]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. (2008)
</bibRefCitation>
, 
<taxonomicName id="4C484D1C1930AD493B3DFB42EADEFB38" box="[321,483,1220,1244]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD493B3DFB42EADEFB38" box="[321,483,1220,1244]" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
does not have a deep curved groove on isthmus, as it is present in 
<taxonomicName id="4C484D1C1930AD493E85FB42EE76FB38" box="[1273,1355,1220,1244]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493E85FB42EE76FB38" box="[1273,1355,1220,1244]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
. Other nominal species with an entirely black colouration (at least when adult) include 
<taxonomicName id="4C484D1C1930AD493E59FB6EEE49FAE4" authority="Bleeker 1856" authorityName="Bleeker" authorityYear="1856" box="[1061,1396,1255,1280]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="heterospilos">
<emphasis id="B93CEA8D1930AD493E59FB6EEFEFFAE4" box="[1061,1234,1256,1280]" italics="true" pageId="16" pageNumber="317">G. heterospilos</emphasis>
Bleeker 1856
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493FFDFB6EEAE0FAC0" authority="Fowler 1944" authorityName="Fowler" authorityYear="1944" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="spilophthalmus">
<emphasis id="B93CEA8D1930AD493FFDFB6EEA7FFAC0" italics="true" pageId="16" pageNumber="317">G. spilophthalmus</emphasis>
Fowler 1944
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493B96FA8AE81AFAC1" authority="Smith 1959" authorityName="Smith" authorityYear="1959" box="[490,807,1292,1317]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="albolineatus">
<emphasis id="B93CEA8D1930AD493B96FA8AE9A4FAC0" box="[490,665,1292,1316]" italics="true" pageId="16" pageNumber="317">G. albolineatus</emphasis>
Smith 1959
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493948FA8AEFC6FAC0" authority="Sawada and Arai 1972" authorityName="Sawada and Arai" authorityYear="1972" box="[820,1275,1292,1317]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="albofasciatus">
<emphasis id="B93CEA8D1930AD493948FA8AE8D0FAC0" box="[820,1005,1292,1316]" italics="true" pageId="16" pageNumber="317">G. albofasciatus</emphasis>
Sawada and Arai 1972
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493F74FA8AE93CFAAC" authority="Harold and Winterbottom 1995" authorityName="Harold and Winterbottom" authorityYear="1995" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="acicularis">
<emphasis id="B93CEA8D1930AD493F74FA8AEEA1FAC0" box="[1288,1436,1292,1316]" italics="true" pageId="16" pageNumber="317">G. acicularis</emphasis>
Harold and Winterbottom 1995
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493872FAB6EF7FFAAC" authority="Suzuki, Yanao and Senou 2012" authorityName="Suzuki, Yanao and Senou" authorityYear="2012" box="[526,1090,1327,1352]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="winterbottomi">
<emphasis id="B93CEA8D1930AD493872FAB6E9F2FAAC" box="[526,719,1328,1352]" italics="true" pageId="16" pageNumber="317">G. winterbottomi</emphasis>
Suzuki, Yanao and 
<bibRefCitation id="EFD94B6E1930AD4939C7FAB6EF7FFAAC" author="Senou" box="[955,1090,1328,1352]" pageId="16" pageNumber="317" refString="Suzuki, T., Yano, K. &amp; Senou, H. (2012) Gobiodon winterbottomi, a New Goby (Actinopterygii: Perciformes: Gobiidae) from Iriomote-jima Island, the Ryukyu Islands, Japan. Bulletin of the National Museum of Nature and Science s, Series A, Supplement 6, 59 - 65." type="book chapter" year="2012">Senou 2012</bibRefCitation>
</taxonomicName>
and a potentially new species from the 
<collectingCountry id="F35F760F1930AD493A82FAD2EA5BFA89" box="[254,358,1364,1389]" name="Maldives" pageId="16" pageNumber="317">Maldives</collectingCountry>
(
<bibRefCitation id="EFD94B6E1930AD493B09FAD2E97CFA88" author="Herler" box="[373,577,1364,1389]" pageId="16" pageNumber="317" refString="Herler, J., Koblmueller, S. &amp; Sturmbauer, C. (2009) Phylogenetic relationships of coral-associated gobies (Teleostei, Gobiidae) from the Red Sea based on mitochondrial DNA data. Marine Biology, 156, 725 - 739. http: // dx. doi. org / 10.1007 / s 00227 - 008 - 1124 - 7" type="journal article" year="2009">
Herler 
<emphasis id="B93CEA8D1930AD493BBAFAD3EACAFA88" box="[454,503,1364,1388]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2009
</bibRefCitation>
; as 
<emphasis id="B93CEA8D1930AD493810FAD2E9BDFA8F" box="[620,640,1364,1387]" italics="true" pageId="16" pageNumber="317">G</emphasis>
. sp. 4). Apart from the possibility that some of the four former species may be synonymous with each other, most of these species represent a genetical clade (tested for 
<taxonomicName id="4C484D1C1930AD493E9CFAFEEE4FFA74" box="[1248,1394,1400,1424]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="acicularis">
<emphasis id="B93CEA8D1930AD493E9CFAFEEE4FFA74" box="[1248,1394,1400,1424]" italics="true" pageId="16" pageNumber="317">G. acicularis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C484D1C1930AD493FFDFAFEEA28FA50" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B93CEA8D1930AD493FFDFAFEEA28FA50" italics="true" pageId="16" pageNumber="317">G. ceramensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1930AD493B31FA1AE926FA50" box="[333,539,1436,1460]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="spilophthalmus">
<emphasis id="B93CEA8D1930AD493B31FA1AE926FA50" box="[333,539,1436,1460]" italics="true" pageId="16" pageNumber="317">G. spilophthalmus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFD94B6E1930AD49383BFA1AE825FA50" author="Harold" box="[583,792,1436,1461]" pageId="16" pageNumber="317" refString="Harold, A. S., Winterbottom, R., Munday, P. L. &amp; Chapman, R. W. (2008) Phylogenetic relationships of Indo-pacific coral gobies of the genus Gobiodon (Teleostei: Gobiidae), based on morphological and molecular data. Bulletin of Marine Science, 82, 119 - 136." type="journal article" year="2008">
Harold 
<emphasis id="B93CEA8D1930AD4938E0FA1BE9F3FA50" box="[668,718,1436,1460]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2008
</bibRefCitation>
; 
<bibRefCitation id="EFD94B6E1930AD493959FA1AE8CDFA50" author="Herler" box="[805,1008,1436,1461]" pageId="16" pageNumber="317" refString="Herler, J., Koblmueller, S. &amp; Sturmbauer, C. (2009) Phylogenetic relationships of coral-associated gobies (Teleostei, Gobiidae) from the Red Sea based on mitochondrial DNA data. Marine Biology, 156, 725 - 739. http: // dx. doi. org / 10.1007 / s 00227 - 008 - 1124 - 7" type="journal article" year="2009">
Herler 
<emphasis id="B93CEA8D1930AD493908FA1BE89BFA50" box="[884,934,1436,1460]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFD94B6E1930AD493980FA1AEFDDFA50" author="Duchene" box="[1020,1248,1436,1461]" pageId="16" pageNumber="317" refString="Duchene, D., Klanten, S. O., Munday, P. L., Herler, J. &amp; van Herwerden, L. (2013) Phylogenetic evidence for recent diversification of obligate coral-dwelling gobies compared with their host corals. Molecular Phylogenetics and Evolution, 69: 123 - 132. http: // dx. doi. org / 10.1016 / j. ympev. 2013.04.033" type="journal article" year="2013">
Duchene 
<emphasis id="B93CEA8D1930AD493E14FA1BEFA7FA50" box="[1128,1178,1436,1460]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2013
</bibRefCitation>
and herein (
<figureCitation id="13732A1A1930AD493F12FA1AEB8EFA3C" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="16" pageNumber="317">Fig. 11</figureCitation>
)), which is distant from 
<taxonomicName id="4C484D1C1930AD493B84FA46E96CFA3C" box="[504,593,1472,1496]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493B84FA46E96CFA3C" box="[504,593,1472,1496]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
and its genetically closest relatives 
<taxonomicName id="4C484D1C1930AD493E61FA46EFFCFA3C" box="[1053,1217,1472,1496]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="fuscoruber">
<emphasis id="B93CEA8D1930AD493E61FA46EFFCFA3C" box="[1053,1217,1472,1496]" italics="true" pageId="16" pageNumber="317">G. fuscoruber</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1930AD493F70FA46EEA9FA3C" box="[1292,1428,1472,1496]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="axillaris">
<emphasis id="B93CEA8D1930AD493F70FA46EEA9FA3C" box="[1292,1428,1472,1496]" italics="true" pageId="16" pageNumber="317">G. axillaris</emphasis>
</taxonomicName>
. Morphologically, 
<taxonomicName id="4C484D1C1930AD493B1EFA62EA8EFA18" box="[354,435,1508,1532]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493B1EFA62EA8EFA18" box="[354,435,1508,1532]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
differs from most members of the other genetic clade in that it is entirely black already when juvenile, and by a rounded first dorsal fin, shaped by a short first fin spine. By contrast, the first dorsal spine is the longest in several species of the other genetic clade. This results in a rather rectangular or even triangular first dorsal fin shape. Also typical for 
<taxonomicName id="4C484D1C1930AD49384FF9D6E9B4F98C" box="[563,649,1616,1640]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD49384FF9D6E9B4F98C" box="[563,649,1616,1640]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
is the presence of a groove between interopercle and isthmus: a comparison with other species having such a groove is given in the remarks section of 
<taxonomicName id="4C484D1C1930AD493EEBF9F2EE0BF968" box="[1175,1334,1652,1676]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1930AD493EEBF9F2EE0BF968" box="[1175,1334,1652,1676]" italics="true" pageId="16" pageNumber="317">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61930AD493F3CF9F3EEA1F968" box="[1344,1436,1653,1676]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
below. 
<taxonomicName id="4C484D1C1930AD493A90F91EEAAFF954" box="[236,402,1688,1712]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493A90F91EEAAFF954" box="[236,402,1688,1712]" italics="true" pageId="16" pageNumber="317">Gobiodon ater</emphasis>
</taxonomicName>
is genetically closely related to 
<taxonomicName id="4C484D1C1930AD493974F91EE8B7F954" box="[776,906,1688,1712]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="axillaris">
<emphasis id="B93CEA8D1930AD493974F91EE8B7F954" box="[776,906,1688,1712]" italics="true" pageId="16" pageNumber="317">G. axillaris</emphasis>
</taxonomicName>
(
<figureCitation id="13732A1A1930AD4939E0F911E8CDF954" box="[924,1008,1687,1712]" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="16" pageNumber="317">Fig. 11</figureCitation>
, 
<bibRefCitation id="EFD94B6E1930AD493982F911EFD7F954" author="Duchene" box="[1022,1258,1687,1712]" pageId="16" pageNumber="317" refString="Duchene, D., Klanten, S. O., Munday, P. L., Herler, J. &amp; van Herwerden, L. (2013) Phylogenetic evidence for recent diversification of obligate coral-dwelling gobies compared with their host corals. Molecular Phylogenetics and Evolution, 69: 123 - 132. http: // dx. doi. org / 10.1016 / j. ympev. 2013.04.033" type="journal article" year="2013">
Duchene 
<emphasis id="B93CEA8D1930AD493E17F91FEFA2F954" box="[1131,1183,1688,1712]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2013
</bibRefCitation>
), although they have very different life colouration, and to 
<taxonomicName id="4C484D1C1930AD49380BF93AE82FF930" box="[631,786,1724,1748]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1930AD49380BF93AE82FF930" box="[631,786,1724,1748]" italics="true" pageId="16" pageNumber="317">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61930AD493966F93BE850F930" box="[794,877,1725,1748]" pageId="16" pageNumber="317" rank="species">sp. nov</taxonomicNameLabel>
, to which it is phenotypically more similar. These species are in a clade together with another undescribed black species from the Indian Ocean (
<taxonomicName id="4C484D1C1930AD493E9FF966EE6EF91C" box="[1251,1363,1760,1784]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D1930AD493E9FF966EE6EF91C" box="[1251,1363,1760,1784]" italics="true" pageId="16" pageNumber="317">Gobiodon</emphasis>
</taxonomicName>
sp. 4, 
<bibRefCitation id="EFD94B6E1930AD493AEBF882EA53F8F8" author="Herler" box="[151,366,1796,1821]" pageId="16" pageNumber="317" refString="Herler, J., Koblmueller, S. &amp; Sturmbauer, C. (2009) Phylogenetic relationships of coral-associated gobies (Teleostei, Gobiidae) from the Red Sea based on mitochondrial DNA data. Marine Biology, 156, 725 - 739. http: // dx. doi. org / 10.1007 / s 00227 - 008 - 1124 - 7" type="journal article" year="2009">
Herler 
<emphasis id="B93CEA8D1930AD493A96F883EA1DF8F8" box="[234,288,1796,1820]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2009
</bibRefCitation>
), which has a very elongate body, and with 
<taxonomicName id="4C484D1C1930AD4939F8F882E8C9F8F8" box="[900,1012,1796,1820]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="histrio">
<emphasis id="B93CEA8D1930AD4939F8F882E8C9F8F8" box="[900,1012,1796,1820]" italics="true" pageId="16" pageNumber="317">G. histrio</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1930AD493E48F882EEA5F8F9" authority="Bleeker 1875" authorityName="Bleeker" authorityYear="1875" box="[1076,1432,1796,1821]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="erythrospilus">
<emphasis id="B93CEA8D1930AD493E48F882EFD2F8F8" box="[1076,1263,1796,1820]" italics="true" pageId="16" pageNumber="317">G. erythrospilus</emphasis>
Bleeker 1875
</taxonomicName>
. Considerable morphological similarities between 
<taxonomicName id="4C484D1C1930AD4938BEF8AEE82CF8A4" box="[706,785,1832,1856]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD4938BEF8AEE82CF8A4" box="[706,785,1832,1856]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1930AD49393BF8AEE8DFF8A4" box="[839,994,1832,1856]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1930AD49393BF8AEE8DFF8A4" box="[839,994,1832,1856]" italics="true" pageId="16" pageNumber="317">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61930AD493995F8AEEF7CF8DB" box="[1001,1089,1832,1855]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
have also been revealed by the geometric morphometric analysis, in which the former in particular overlaps with western Pacific specimens of the latter (
<figureCitation id="13732A1A1930AD493A9AF8E9EA02F86C" box="[230,319,1903,1928]" captionStart="FIGURE 10" captionStartId="23.[151,250,1629,1651]" captionTargetBox="[224,1369,736,1597]" captionTargetId="figure@23.[209,1384,724,1615]" captionTargetPageId="23" captionText="FIGURE 10. First two principal components of a PCA of Procrustes coordinates of 15 landmarks and 6 semi-landmarks put on 111 specimens of Gobiodon. The holotype (MNHN A- 4015) of G. unicolor (Castelnau, 1873) is marked by a red circle. RS = Red Sea. For species abbreviations see Table 1." httpUri="https://zenodo.org/record/249080/files/figure.png" pageId="16" pageNumber="317">Fig. 10</figureCitation>
). MANOVA, however, confirmed significant shape differences between all three populations. Species discrimination was also supported by the PCA of body proportions of the Red Sea 
<typeStatus id="54F3883D1930AD493EE0F813EFF1F849" box="[1180,1228,1941,1965]" pageId="16" pageNumber="317">type</typeStatus>
specimens (adults only): 
<taxonomicName id="4C484D1C1930AD493A9FF83FEA09F834" box="[227,308,1977,2000]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1930AD493A9FF83FEA09F834" box="[227,308,1977,2000]" italics="true" pageId="16" pageNumber="317">G. ater</emphasis>
</taxonomicName>
mainly differs from 
<taxonomicName id="4C484D1C1930AD493858F83FE9FFF834" box="[548,706,1976,2000]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1930AD493858F83FE9FFF834" box="[548,706,1976,2000]" italics="true" pageId="16" pageNumber="317">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61930AD4938B6F83FE818F834" box="[714,805,1977,2000]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
by its longer abdomen (V—A: 30.6–34.3 versus 25.0– 29.8% SL) and lower posterior body depth (D2—A: 29.5–32.3 versus 33.6–37.2% SL). They can further be distinguished by life colour and maximum size. Inferred from this morphological comparison and from a previous, more elaborate genetic analysis (
<bibRefCitation id="EFD94B6E1931AD483876FF3AE9EBFF30" author="Herler" box="[522,726,188,213]" pageId="17" pageNumber="318" refString="Herler, J., Koblmueller, S. &amp; Sturmbauer, C. (2009) Phylogenetic relationships of coral-associated gobies (Teleostei, Gobiidae) from the Red Sea based on mitochondrial DNA data. Marine Biology, 156, 725 - 739. http: // dx. doi. org / 10.1007 / s 00227 - 008 - 1124 - 7" type="journal article" year="2009">
Herler 
<emphasis id="B93CEA8D1931AD483827FF3BE9B1FF30" box="[603,652,188,212]" italics="true" pageId="17" pageNumber="318">et al</emphasis>
. 2009
</bibRefCitation>
) that included sequences of 
<emphasis id="B93CEA8D1931AD483E66FF3AEFA0FF30" box="[1050,1181,188,212]" italics="true" pageId="17" pageNumber="318">cytochrome</emphasis>
b, it is evident that 
<taxonomicName id="4C484D1C1931AD483FFDFF3AEBF9FF1C" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="ater">
<emphasis id="B93CEA8D1931AD483FFDFF3AEBF9FF1C" italics="true" pageId="17" pageNumber="318">G. ater</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C1931AD483A80FF66EAA5FF1C" box="[252,408,224,248]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1931AD483A80FF66EAA5FF1C" box="[252,408,224,248]" italics="true" pageId="17" pageNumber="318">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61931AD483BDCFF66EAC4FF13" box="[416,505,224,247]" pageId="17" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
are very closely related but distinct species. The genetic distance between the two species calculated herein exceeds the level of 2% accepted for species delineation (
<bibRefCitation id="EFD94B6E1931AD483E49FE82EE3BFEF8" author="Harold" box="[1077,1286,260,285]" pageId="17" pageNumber="318" refString="Harold, A. S., Winterbottom, R., Munday, P. L. &amp; Chapman, R. W. (2008) Phylogenetic relationships of Indo-pacific coral gobies of the genus Gobiodon (Teleostei: Gobiidae), based on morphological and molecular data. Bulletin of Marine Science, 82, 119 - 136." type="journal article" year="2008">
Harold 
<emphasis id="B93CEA8D1931AD483EF7FE83EF81FEF8" box="[1163,1212,260,284]" italics="true" pageId="17" pageNumber="318">et al</emphasis>
. 2008
</bibRefCitation>
). The genetic distance to 
<taxonomicName id="4C484D1C1931AD483B5EFEAEE9D2FEA4" authority="Harold et al. (2008)" authorityName="Harold et al." authorityYear="2008" box="[290,751,295,320]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B93CEA8D1931AD483B5EFEAEEA94FEA4" box="[290,425,296,320]" italics="true" pageId="17" pageNumber="318">G. unicolor</emphasis>
sensu 
<bibRefCitation id="EFD94B6E1931AD48387CFEA1E9D2FEA4" box="[512,751,295,320]" pageId="17" pageNumber="318" refString="Harold, A. S., Winterbottom, R., Munday, P. L. &amp; Chapman, R. W. (2008) Phylogenetic relationships of Indo-pacific coral gobies of the genus Gobiodon (Teleostei: Gobiidae), based on morphological and molecular data. Bulletin of Marine Science, 82, 119 - 136." type="journal article">
Harold 
<emphasis id="B93CEA8D1931AD483826FEAFE9ACFEA4" box="[602,657,296,320]" italics="true" pageId="17" pageNumber="318">et al</emphasis>
. (2008)
</bibRefCitation>
</taxonomicName>
from the Great Barrier Reef, being synonymous with 
<taxonomicName id="4C484D1C1931AD483FFDFEAEEA2DFE80" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="fuscoruber" status="sp. nov.">
<emphasis id="B93CEA8D1931AD483FFDFEAEEA2DFE80" italics="true" pageId="17" pageNumber="318">G. fuscoruber</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61931AD483B6BFECBEA4DFE80" box="[279,368,333,356]" pageId="17" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
(see below for details), is even greater (0.032).
</paragraph>
</subSubSection>
</treatment>
</document>