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<document id="B3D5B48429DFC235E080D1F9C4FADAE3" ID-CLB-Dataset="25460" ID-DOI="10.11646/zootaxa.4671.1.10" ID-GBIF-Dataset="f3187ddd-f79a-43d2-9692-e101210783bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="3450086" ID-ZooBank="B4441448-ED8F-4690-A2F1-286E911025C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568898282055" checkinUser="plazi" docAuthor="Smaga, Christopher R., Ttito, Alex &amp; Catenazzi, Alessandro" docDate="2019" docId="03DE0124FFDCFF83FF4AFB27FDD7FE60" docLanguage="en" docName="zootaxa.4671.1.10.pdf" docOrigin="Zootaxa 4671 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcanumophis problematicus Smaga, Ttito &amp; Catenazzi, 2019, new combination" docType="treatment" docVersion="9" lastPageNumber="133" masterDocId="FFE7795CFFDFFF86FFDDFFD2FFD3FFEB" masterDocTitle="Arcanumophis, a new genus and generic allocation for Erythrolamprus problematicus (Myers 1986), Xenodontinae (Colubridae) from the Cordillera de Carabaya, southern Peru" masterLastPageNumber="138" masterPageNumber="129" pageNumber="132" updateTime="1732568123160" updateUser="ExternalLinkService">
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<mods:title id="0F646465D94A87830E1B3AAE1EC62880">Arcanumophis, a new genus and generic allocation for Erythrolamprus problematicus (Myers 1986), Xenodontinae (Colubridae) from the Cordillera de Carabaya, southern Peru</mods:title>
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<mods:namePart id="FDD28D8565BA10F0971224CE0742CF69">Smaga, Christopher R.</mods:namePart>
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<mods:namePart id="B34C8A01C173BC70EFA00EB245C8E070">Ttito, Alex</mods:namePart>
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<mods:namePart id="0C102F1129A3A6B45D0CA8E23F2C6614">Catenazzi, Alessandro</mods:namePart>
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<mods:title id="997AC423D8BC31404666833B790BD396">Zootaxa</mods:title>
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<mods:date id="8A1EA4948BC79C73BEB4686B9F694ADE">2019</mods:date>
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<treatment id="03DE0124FFDCFF83FF4AFB27FDD7FE60" ID-GBIF-Taxon="159565316" LSID="urn:lsid:plazi:treatment:03DE0124FFDCFF83FF4AFB27FDD7FE60" httpUri="http://treatment.plazi.org/id/03DE0124FFDCFF83FF4AFB27FDD7FE60" lastPageId="5" lastPageNumber="133" pageId="3" pageNumber="132" scope_family="Colubridae">
<subSubSection id="C36DE3B9FFDCFF85FF4AFB27FD09FAE4" box="[151,730,1269,1296]" pageId="3" pageNumber="132" type="nomenclature">
<paragraph id="8BC8B032FFDCFF85FF4AFB27FD09FAE4" blockId="3.[151,730,1269,1331]" box="[151,730,1269,1296]" pageId="3" pageNumber="132">
<heading id="D080075EFFDCFF85FF4AFB27FD09FAE4" bold="true" box="[151,730,1269,1296]" fontSize="11" level="1" pageId="3" pageNumber="132" reason="1">
<emphasis id="B9036C20FFDCFF85FF4AFB27FD09FAE4" bold="true" box="[151,730,1269,1296]" pageId="3" pageNumber="132">
<taxonomicName id="4C77CBB1FFDCFF85FF4AFB27FE2EFAFB" ID-CoL="8H6CV" authorityName="Smaga &amp; Ttito &amp; Catenazzi" authorityYear="2019" baseAuthorityName="Smaga &amp; Ttito &amp; Catenazzi" baseAuthorityYear="1986" box="[151,509,1269,1296]" class="Reptilia" family="Colubridae" genus="Arcanumophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="132" phylum="Chordata" rank="species" species="problematicus" status="comb. nov.">
<emphasis id="B9036C20FFDCFF85FF4AFB27FE2EFAFB" bold="true" box="[151,509,1269,1296]" italics="true" pageId="3" pageNumber="132">Arcanumophis problematicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230D15BFFDCFF85FDD9FB27FD09FAE4" box="[516,730,1269,1295]" pageId="3" pageNumber="132" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DE3B9FFDCFF85FF4AFACBFEC1FAD8" box="[151,274,1305,1331]" pageId="3" pageNumber="132" type="description">
<paragraph id="8BC8B032FFDCFF85FF4AFACBFEC1FAD8" blockId="3.[151,730,1269,1331]" box="[151,274,1305,1331]" pageId="3" pageNumber="132">
(
<figureCitation id="134CACB7FFDCFF85FF42FACBFED9FAD8" box="[159,266,1305,1331]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,880,905]" captionStartId-1="4.[151,250,1816,1841]" captionStartId-2="5.[151,250,989,1014]" captionTargetBox-0="[162,1425,288,842]" captionTargetBox-1="[151,1436,956,1797]" captionTargetBox-2="[151,1403,409,960]" captionTargetId-0="figure@4.[151,1436,301,861]" captionTargetId-1="figure@4.[151,1436,956,1797]" captionTargetId-2="figure@5.[151,1436,409,969]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Full body of Arcanumophis problematicus CORBIDI 18731 showing dorsal and ventral coloration patterns (pre- served specimen). Photographs by A. Catenazzi." captionText-1="FIGURE 3. Coloration in life of Arcanumophis problematicus (CORBIDI 18731) from Santo Domingo, Department of Puno, Peru. Photographs by A. Catenazzi." captionText-2="FIGURE 4. Head of Arcanumophis problematicus (CORBIDI 18731), showing horizontal crease on rostrum. Photographs by A. Catenazzi." figureDoi-0="http://doi.org/10.5281/zenodo.3450090" figureDoi-1="http://doi.org/10.5281/zenodo.3450092" figureDoi-2="http://doi.org/10.5281/zenodo.3450094" httpUri-0="https://zenodo.org/record/3450090/files/figure.png" httpUri-1="https://zenodo.org/record/3450092/files/figure.png" httpUri-2="https://zenodo.org/record/3450094/files/figure.png" pageId="3" pageNumber="132">Figs. 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36DE3B9FFDCFF85FF4AFAB3FEDCF99C" pageId="3" pageNumber="132" type="materials_examined">
<paragraph id="8BC8B032FFDCFF85FF4AFAB3FD28FA03" blockId="3.[151,1437,1376,2015]" pageId="3" pageNumber="132">
<materialsCitation id="3B1FBA6FFFDCFF85FF4AFAB3FD28FA03" ID-GBIF-Occurrence="2416712447" collectingDate="22 November- 20 December" collectionCode="FMNH" county="Hilda Hempl Heller" elevation="1520" latitude="-14.21667" location="Sandia" longLatPrecision="1" longitude="-69.16667" municipality="San Juan del Oro" pageId="3" pageNumber="132" specimenCode="FMNH 64733" specimenCount="1" stateProvince="Puno" typeStatus="holotype">
<emphasis id="B9036C20FFDCFF85FF4AFAB3FED8FA90" bold="true" box="[151,267,1377,1403]" pageId="3" pageNumber="132">
<typeStatus id="54CC0E90FFDCFF85FF4AFAB3FED5FA90" box="[151,262,1377,1403]" pageId="3" pageNumber="132" type="holotype">Holotype</typeStatus>
.
</emphasis>
adult male, total length
<quantity id="4C8F1DD7FFDCFF85FDF1FAB2FD57FA90" box="[556,644,1376,1403]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="132" unit="mm" value="275.0">275mm</quantity>
, tail length
<quantity id="4C8F1DD7FFDCFF85FCCFFAB2FC8FFA90" box="[786,860,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="132" unit="mm" value="51.0">51mm</quantity>
, tail 18% of total length,
<specimenCode id="DBD11849FFDCFF85FB51FAB3FAFEFA90" box="[1164,1325,1377,1403]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="132" type="Museum">FMNH 64733</specimenCode>
collected between
<date id="FFC996F2FFDCFF85FF21FA57FDEFFA74" box="[252,572,1413,1439]" pageId="3" pageNumber="132">
<collectingDate id="EF8D6F1AFFDCFF85FF21FA57FDEFFA74" box="[252,572,1413,1439]" pageId="3" pageNumber="132">22 November20 December</collectingDate>
</date>
of 1950 by
<collectingCounty id="62A9C8BEFFDCFF85FD1AFA57FC74FA74" box="[711,935,1413,1439]" pageId="3" pageNumber="132">Hilda Hempl Heller</collectingCounty>
at
<quantity id="4C8F1DD7FFDCFF85FC16FA56FBF0FA74" box="[971,1059,1412,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="132" unit="m" value="1520.0">
<elevation id="005A5701FFDCFF85FC16FA56FBF0FA74" box="[971,1059,1412,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="132" unit="m" value="1520.0">1520 m</elevation>
</quantity>
, near
<collectingMunicipality id="6BAC2A48FFDCFF85FBB5FA57FAFDFA74" box="[1128,1326,1413,1439]" pageId="3" pageNumber="132">San Juan del Oro</collectingMunicipality>
, about
<quantity id="4C8F1DD7FFDCFF85FA5DFA57FF68FA28" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="132" unit="km" value="30.0">30 km</quantity>
ENE of the town of
<location id="8EA8E6E9FFDCFF85FE77FA7BFE2AFA28" LSID="urn:lsid:plazi:treatment:03DE0124FFDCFF83FF4AFB27FDD7FE60:8EA8E6E9FFDCFF85FE77FA7BFE2AFA28" box="[426,505,1449,1475]" county="Hilda Hempl Heller" latitude="-14.21667" longLatPrecision="1" longitude="-69.16667" municipality="San Juan del Oro" name="Sandia" pageId="3" pageNumber="132" stateProvince="Puno">Sandia</location>
,
<location id="8EA8E6E9FFDCFF85FDD8FA7BFD0CFA28" LSID="urn:lsid:plazi:treatment:03DE0124FFDCFF83FF4AFB27FDD7FE60:8EA8E6E9FFDCFF85FDD8FA7BFD0CFA28" box="[517,735,1449,1475]" county="Hilda Hempl Heller" latitude="-14.21667" longLatPrecision="1" longitude="-69.16667" municipality="San Juan del Oro" name="Province of Sandia" pageId="3" pageNumber="132" stateProvince="Puno">Province of Sandia</location>
,
<location id="8EA8E6E9FFDCFF85FD36FA7BFCBCFA28" LSID="urn:lsid:plazi:treatment:03DE0124FFDCFF83FF4AFB27FDD7FE60:8EA8E6E9FFDCFF85FD36FA7BFCBCFA28" box="[747,879,1449,1475]" county="Hilda Hempl Heller" latitude="-14.21667" longLatPrecision="1" longitude="-69.16667" municipality="San Juan del Oro" name="Department" pageId="3" pageNumber="132" stateProvince="Puno">Department</location>
of
<collectingRegion id="49B37ED0FFDCFF85FC4BFA7BFC01FA28" box="[918,978,1449,1475]" country="Peru" name="Puno" pageId="3" pageNumber="132">Puno</collectingRegion>
, on the west bank of the
<location id="8EA8E6E9FFDCFF85FB2DFA7BFA4FFA28" LSID="urn:lsid:plazi:treatment:03DE0124FFDCFF83FF4AFB27FDD7FE60:8EA8E6E9FFDCFF85FB2DFA7BFA4FFA28" box="[1264,1436,1449,1475]" county="Hilda Hempl Heller" latitude="-14.21667" longLatPrecision="1" longitude="-69.16667" municipality="San Juan del Oro" name="Rio Tambopata" pageId="3" pageNumber="132" stateProvince="Puno">Río Tambopata</location>
(approximated coordinates
<geoCoordinate id="EE43D6F5FFDCFF85FE17FA1FFD83FA0C" box="[458,592,1485,1511]" degrees="14.21667" direction="south" orientation="latitude" pageId="3" pageNumber="132" precision="1" value="-14.21667">14.21667°S</geoCoordinate>
,
<geoCoordinate id="EE43D6F5FFDCFF85FD86FA1FFD3CFA03" box="[603,751,1485,1512]" degrees="69.16667" direction="west" orientation="longitude" pageId="3" pageNumber="132" precision="1" value="-69.16667">69.16667°W</geoCoordinate>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC8B032FFDCFF85FF1AFA22FEDCF99C" blockId="3.[151,1437,1376,2015]" pageId="3" pageNumber="132">
<emphasis id="B9036C20FFDCFF85FF1AFA22FE63F9E0" bold="true" box="[199,432,1520,1547]" pageId="3" pageNumber="132">Referred specimen.</emphasis>
adult female, total length
<quantity id="4C8F1DD7FFDCFF85FD01FA23FCE6F9E0" box="[732,821,1521,1547]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="132" unit="mm" value="210.0">210mm</quantity>
, tail length
<quantity id="4C8F1DD7FFDCFF85FC66FA23FBD5F9E0" box="[955,1030,1521,1547]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="132" unit="mm" value="40.0">40mm</quantity>
, tail 19% of total length,
<collectionCode id="ED6628F7FFDCFF85FAF4FA23FA4FF9E0" box="[1321,1436,1521,1547]" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="3" pageNumber="132">CORBIDI</collectionCode>
18731 collected near the abandoned mine of Santo Domingo at
<quantity id="4C8F1DD7FFDCFF85FCB7F9C7FC13F9C4" box="[874,960,1557,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="3" pageNumber="132" unit="m" value="1960.0">1960 m</quantity>
a.s.l. (
<geoCoordinate id="EE43D6F5FFDCFF85FBD1F9C7FB47F9C4" box="[1036,1172,1557,1583]" degrees="13.83202" direction="south" orientation="latitude" pageId="3" pageNumber="132" precision="1" value="-13.83202">13.83202°S</geoCoordinate>
,
<geoCoordinate id="EE43D6F5FFDCFF85FB7DF9C6FAE2F9C4" box="[1184,1329,1556,1583]" degrees="69.63705" direction="west" orientation="longitude" pageId="3" pageNumber="132" precision="1" value="-69.63705">69.63705°W</geoCoordinate>
; GPS coordinates, WGS84), Province of Carabaya, Department of
<collectingRegion id="49B37ED0FFDCFF85FCF5F9EBFCB1F9B8" box="[808,866,1593,1619]" country="Peru" name="Puno" pageId="3" pageNumber="132">Puno</collectingRegion>
on
<date id="FFC996F2FFDCFF85FC51F9EBFBC7F9B8" box="[908,1044,1593,1619]" pageId="3" pageNumber="132" value="2016-06-08">8 June 2016</date>
by A. Catenazzi, A. Ttito and J. C. Jahuanchi.
</paragraph>
</subSubSection>
<subSubSection id="C36DE3B9FFDCFF83FF1AF953FDD7FE60" lastPageId="5" lastPageNumber="134" pageId="3" pageNumber="132" type="description">
<paragraph id="8BC8B032FFDCFF82FF1AF953FAA0FECB" blockId="3.[151,1437,1376,2015]" lastBlockId="4.[151,1437,153,288]" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132">
<emphasis id="B9036C20FFDCFF85FF1AF953FD73F970" bold="true" box="[199,672,1664,1691]" pageId="3" pageNumber="132">
<emphasis id="B9036C20FFDCFF85FF1AF953FEB8F970" bold="true" box="[199,363,1664,1691]" italics="true" pageId="3" pageNumber="132">Description of</emphasis>
CORBIDI 18731 (
<figureCitation id="134CACB7FFDCFF85FD96F952FD42F970" box="[587,657,1664,1691]" captionStart="FIGURE 4" captionStartId="5.[151,250,989,1014]" captionTargetBox="[151,1403,409,960]" captionTargetId="figure@5.[151,1436,409,969]" captionTargetPageId="5" captionText="FIGURE 4. Head of Arcanumophis problematicus (CORBIDI 18731), showing horizontal crease on rostrum. Photographs by A. Catenazzi." figureDoi="http://doi.org/10.5281/zenodo.3450094" httpUri="https://zenodo.org/record/3450094/files/figure.png" pageId="3" pageNumber="132">Fig. 4</figureCitation>
).
</emphasis>
Small, slender snake with a slender head and no distinct neck, head narrowing at snout; pupil rounded, eyes diameter almost as long as eye-nostril distance; rostral crescent shaped and twice as wide as high, visible from above, wider in the center, with a horizontal crease at lower 2/3 of the rostral scale, extending all the way across, curving with the shape of the scale; loreal present, as long as wide; eight supralabials, 2
<superScript id="7C021D7AFFDCFF85FEE4F8C2FE9AF8F5" attach="left" box="[313,329,1808,1822]" fontSize="6" pageId="3" pageNumber="132">nd</superScript>
contacting loreal; 3
<superScript id="7C021D7AFFDCFF85FDF4F8C2FD95F8C0" attach="none" box="[553,582,1808,1835]" fontSize="6" pageId="3" pageNumber="132">rd</superScript>
5
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contacting eye; eight infralabials, 1
<superScript id="7C021D7AFFDCFF85FC2CF8C2FBD8F8C0" attach="none" box="[1009,1035,1808,1835]" fontSize="6" pageId="3" pageNumber="132">st</superScript>
4
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contacting anterior genial, 4
<superScript id="7C021D7AFFDCFF85FABAF8C2FA50F8C0" attach="none" box="[1383,1411,1808,1835]" fontSize="6" pageId="3" pageNumber="132">th</superScript>
5
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contacting posterior genial; two internasals, roughly trapezoidal, wider posteriorly and narrowing toward the rostral; frontal roughly pentagonal, twice as long as wide and twice as long as prefrontals; supraocular kidney shaped, slightly shorter than frontal, narrowing anteriorly; parietals paired, slightly longer than frontal and one-and-a-half times as long as wide; nasal twice as long as high, contacting 1
<superScript id="7C021D7AFFDCFF85FCBCF872FCBFF845" attach="left" box="[865,876,1952,1966]" fontSize="6" pageId="3" pageNumber="132">st</superScript>
and 2
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supralabial; one preocular two-and-a-half times as high as long, longer dorsally and shortening toward supralabials; two postoculars, upper postocular dis- tinctly larger than lower; temporals 1+2; anterior temporal large, two-and-a-half times as long as high, in contact with 6
<superScript id="7C021D7AFFDBFF82FF3DFF6EFF3EFF21" attach="left" box="[224,237,188,202]" fontSize="6" pageId="4" pageNumber="133">th</superScript>
and 7
<superScript id="7C021D7AFFDBFF82FEE8FF6EFE91FF21" attach="left" box="[309,322,188,202]" fontSize="6" pageId="4" pageNumber="133">th</superScript>
supralabials; two upper posterior temporals three times as long as lower posterior temporal; four genials; anterior pair twice as long as wide; posterior pair three time as long as wide; dorsal scales smooth, 19/17/17; ventrals 137, anal plate divided; subcaudals 39 pairs. For details on hemipenial morphology, see
<bibRefCitation id="EFE6CDC3FFDBFF82FB0FFED7FABDFECB" author="Myers, C. W." box="[1234,1390,261,288]" pageId="4" pageNumber="133" pagination="1 - 12" refId="ref4173" refString="Myers, C. W. (1986) An enigmatic new snake from the Peruvian Andes, with notes on the Xenodontini (Colubridae: Xenodontinae). American Museum Novitates, 2853, 1 - 12." type="journal article" year="1986">Myers (1986)</bibRefCitation>
.
</paragraph>
<caption id="DF08E0BAFFDBFF82FF4AFCA2FD51FC46" ID-DOI="http://doi.org/10.5281/zenodo.3450090" ID-Zenodo-Dep="3450090" httpUri="https://zenodo.org/record/3450090/files/figure.png" pageId="4" pageNumber="133" startId="4.[151,250,880,905]" targetBox="[162,1425,288,842]" targetPageId="4">
<paragraph id="8BC8B032FFDBFF82FF4AFCA2FD51FC46" blockId="4.[151,1436,880,941]" pageId="4" pageNumber="133">
<emphasis id="B9036C20FFDBFF82FF4AFCA2FEC7FC62" bold="true" box="[151,276,880,905]" pageId="4" pageNumber="133">FIGURE 2.</emphasis>
Full body of
<taxonomicName id="4C77CBB1FFDBFF82FE7FFCA3FD1FFC62" authorityName="Smaga &amp; Ttito &amp; Catenazzi" authorityYear="2019" baseAuthorityName="Smaga &amp; Ttito &amp; Catenazzi" baseAuthorityYear="1986" box="[418,716,881,905]" class="Reptilia" family="Colubridae" genus="Arcanumophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="133" phylum="Chordata" rank="species" species="problematicus">
<emphasis id="B9036C20FFDBFF82FE7FFCA3FD1FFC62" box="[418,716,881,905]" italics="true" pageId="4" pageNumber="133">Arcanumophis problematicus</emphasis>
</taxonomicName>
CORBIDI 18731 showing dorsal and ventral coloration patterns (pre- served specimen). Photographs by A. Catenazzi.
</paragraph>
</caption>
<caption id="DF08E0BAFFDBFF82FF4AF8CAFDD3F8BE" ID-DOI="http://doi.org/10.5281/zenodo.3450092" ID-Zenodo-Dep="3450092" httpUri="https://zenodo.org/record/3450092/files/figure.png" pageId="4" pageNumber="133" startId="4.[151,250,1816,1841]" targetBox="[151,1436,956,1797]" targetPageId="4">
<paragraph id="8BC8B032FFDBFF82FF4AF8CAFDD3F8BE" blockId="4.[151,1436,1816,1877]" pageId="4" pageNumber="133">
<emphasis id="B9036C20FFDBFF82FF4AF8CAFEC0F8DA" bold="true" box="[151,275,1816,1841]" pageId="4" pageNumber="133">FIGURE 3.</emphasis>
Coloration in life of
<taxonomicName id="4C77CBB1FFDBFF82FE31F8CBFCC5F8DA" authorityName="Smaga &amp; Ttito &amp; Catenazzi" authorityYear="2019" baseAuthorityName="Smaga &amp; Ttito &amp; Catenazzi" baseAuthorityYear="1986" box="[492,790,1817,1841]" class="Reptilia" family="Colubridae" genus="Arcanumophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="133" phylum="Chordata" rank="species" species="problematicus">
<emphasis id="B9036C20FFDBFF82FE31F8CBFCC5F8DA" box="[492,790,1817,1841]" italics="true" pageId="4" pageNumber="133">Arcanumophis problematicus</emphasis>
</taxonomicName>
(CORBIDI 18731) from Santo Domingo, Department of Puno, Peru. Photographs by A. Catenazzi.
</paragraph>
</caption>
<paragraph id="8BC8B032FFDBFF83FF1AF850FDD7FE60" blockId="4.[151,1436,1922,2022]" lastBlockId="5.[151,1437,152,395]" lastPageId="5" lastPageNumber="134" pageId="4" pageNumber="133">
<emphasis id="B9036C20FFDBFF82FF1AF850FDE6F876" bold="true" box="[199,565,1922,1950]" pageId="4" pageNumber="133">
<emphasis id="B9036C20FFDBFF82FF1AF850FE7BF875" bold="true" box="[199,424,1922,1950]" italics="true" pageId="4" pageNumber="133">Coloration patterns</emphasis>
(
<figureCitation id="134CACB7FFDBFF82FE6AF850FDF4F876" box="[439,551,1922,1949]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,880,905]" captionStartId-1="4.[151,250,1816,1841]" captionStartId-2="5.[151,250,989,1014]" captionTargetBox-0="[162,1425,288,842]" captionTargetBox-1="[151,1436,956,1797]" captionTargetBox-2="[151,1403,409,960]" captionTargetId-0="figure@4.[151,1436,301,861]" captionTargetId-1="figure@4.[151,1436,956,1797]" captionTargetId-2="figure@5.[151,1436,409,969]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Full body of Arcanumophis problematicus CORBIDI 18731 showing dorsal and ventral coloration patterns (pre- served specimen). Photographs by A. Catenazzi." captionText-1="FIGURE 3. Coloration in life of Arcanumophis problematicus (CORBIDI 18731) from Santo Domingo, Department of Puno, Peru. Photographs by A. Catenazzi." captionText-2="FIGURE 4. Head of Arcanumophis problematicus (CORBIDI 18731), showing horizontal crease on rostrum. Photographs by A. Catenazzi." figureDoi-0="http://doi.org/10.5281/zenodo.3450090" figureDoi-1="http://doi.org/10.5281/zenodo.3450092" figureDoi-2="http://doi.org/10.5281/zenodo.3450094" httpUri-0="https://zenodo.org/record/3450090/files/figure.png" httpUri-1="https://zenodo.org/record/3450092/files/figure.png" httpUri-2="https://zenodo.org/record/3450094/files/figure.png" pageId="4" pageNumber="133">Figs. 24</figureCitation>
):
</emphasis>
Dorsal color dark brown, fading to a dark olive on flanks; head sides exhibiting two yellowish-white stripes running from rostral through lower margin of eye and ending at mouth corner; two small and cream white circular nape blotches separated from parietal tips by one dorsal scale; a cream, elongated blotch, three times as long as high, present on each side of neck; supralabials 13 dark, turning pale at 45, and retuning dark at 68; infralabials pale with slight black blotching; two paravertebral cream white stripes originating slightly dorsally and posteriorly to cream neck blotch; paravertebral stripes mostly restricted to 6
<superScript id="7C021D7AFFDAFF83FB02FF32FB3FFF05" attach="left" box="[1247,1260,224,238]" fontSize="6" pageId="5" pageNumber="134">th</superScript>
dorsal row, descending to 4
<superScript id="7C021D7AFFDAFF83FEF6FED6FEEBFEF9" attach="left" box="[299,312,260,274]" fontSize="6" pageId="5" pageNumber="134">th</superScript>
dorsal row near the tail; venter cream white, ventral surface of head slightly darker than body venter. In life, the head and flanks exuded bright yellow stripes (
<figureCitation id="134CACB7FFDAFF83FCFDFEFBFCB6FEA8" box="[800,869,297,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1816,1841]" captionTargetBox="[151,1436,956,1797]" captionTargetId="figure@4.[151,1436,956,1797]" captionTargetPageId="4" captionText="FIGURE 3. Coloration in life of Arcanumophis problematicus (CORBIDI 18731) from Santo Domingo, Department of Puno, Peru. Photographs by A. Catenazzi." figureDoi="http://doi.org/10.5281/zenodo.3450092" httpUri="https://zenodo.org/record/3450092/files/figure.png" pageId="5" pageNumber="134">Fig. 3</figureCitation>
), which faded to white after preservation (
<figureCitation id="134CACB7FFDAFF83FA94FEFBFA5CFEA8" box="[1353,1423,297,323]" captionStart="FIGURE 2" captionStartId="4.[151,250,880,905]" captionTargetBox="[162,1425,288,842]" captionTargetId="figure@4.[151,1436,301,861]" captionTargetPageId="4" captionText="FIGURE 2. Full body of Arcanumophis problematicus CORBIDI 18731 showing dorsal and ventral coloration patterns (pre- served specimen). Photographs by A. Catenazzi." figureDoi="http://doi.org/10.5281/zenodo.3450090" httpUri="https://zenodo.org/record/3450090/files/figure.png" pageId="5" pageNumber="134">Fig. 2</figureCitation>
). The nuchal spots, which were a less bright yellow than the stripes, retained similar coloration after preservative, with only a slight loss of yellow.
</paragraph>
</subSubSection>
</treatment>
</document>