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<document id="E822B78D33F0314B85D1DF4DB8E52617" ID-CLB-Dataset="9742" ID-DOI="10.5852/ejt.2020.720.1119" ID-GBIF-Dataset="60b5a286-c831-49ba-b499-ad61183e7f36" ID-Zenodo-Dep="4085203" ID-ZooBank="A32297C8-00D2-4A71-837A-CE49107C1F27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1602601192165" checkinUser="valdenar" docAuthor="Wesener, Thomas" docDate="2020" docId="03B187B1B716FFA1255835CBFD3E26FA" docLanguage="en" docName="ejt_720_wesener.pdf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichromatobolus Wesener 2020, gen. nov." docType="treatment" docUuid="2BC93D12-96E1-4ECC-B83D-44249A06E099" docUuidSource="ZooBank" docVersion="8" lastPageNumber="109" masterDocId="FF88FFC9B714FFA327203643FF952113" masterDocTitle="Dichromatobolus, a new genus of spirobolidan millipedes from Madagascar (Spirobolida, Pachybolidae)" masterLastPageNumber="120" masterPageNumber="107" pageNumber="109" updateTime="1698846057459" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="84EB38D76934F029942D35E06456F779" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="82F3B75F15EC4C8FC080C849A9C86D38">
<mods:title id="C1EF7202C56AEC75D4389FC72A20D4EC">Dichromatobolus, a new genus of spirobolidan millipedes from Madagascar (Spirobolida, Pachybolidae)</mods:title>
</mods:titleInfo>
<mods:name id="E05E921450855C719E7C44714A6FC749" type="personal">
<mods:role id="C7D1954EDCB7E9D863915308BC1214BF">
<mods:roleTerm id="1D7848FEA887E94832C9660A1F17679D">Author</mods:roleTerm>
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<mods:namePart id="6252610CEA90CCE318EF42B0BFE9A22E">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="ED7DC37F99C0C5D33852ECA9D67E9B78" type="LSID">urn:lsid:zoobank.org:author:86DEA7CD-988C-43EC-B9D6-C51000595B47</mods:nameIdentifier>
<mods:affiliation id="F272379D6EAFCCE70ADCFDC17A7BC198">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="744645D3CF1FD65C75DD60D3B50554EB" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5E1FBBF49E00A520E27E59A0B6466376">text</mods:typeOfResource>
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<mods:title id="C28D2A93D11F8012C4A3D082AB2824B8">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C352399771749DEABC833DBF87C30133">
<mods:date id="BD1A540291D98F96F17E1AD2769A323F">2020</mods:date>
<mods:detail id="FBB2A4191604D78D39C9BE372F550DD6" type="pubDate">
<mods:number id="AC70E765F9EEAEED3B2979BD9294DC12">2020-10-12</mods:number>
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<mods:detail id="813291CCE0BA9E34A749028B99F1AE91" type="volume">
<mods:number id="117285CCBD98AF93892811F74DCC3250">720</mods:number>
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<mods:classification id="DAEF76E0B076B1A284799DDD1DC42B32">journal article</mods:classification>
<mods:identifier id="BB9194504A82CF766FB8D8DBFB1802A6" type="CLB-Dataset">9742</mods:identifier>
<mods:identifier id="B30EBD3338892A3427DB521A1DEBA2A6" type="DOI">10.5852/ejt.2020.720.1119</mods:identifier>
<mods:identifier id="08DD3DC1931618A4FD7F1420FA18A5B7" type="GBIF-Dataset">60b5a286-c831-49ba-b499-ad61183e7f36</mods:identifier>
<mods:identifier id="3008AD518C40B734D1CCFC2B85A7A0A4" type="Zenodo-Dep">4085203</mods:identifier>
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<treatment id="03B187B1B716FFA1255835CBFD3E26FA" ID-DOI="http://doi.org/10.5281/zenodo.4324179" ID-GBIF-Taxon="168338727" ID-Zenodo-Dep="4324179" LSID="urn:lsid:zoobank.org:act:2BC93D12-96E1-4ECC-B83D-44249A06E099" httpUri="http://treatment.plazi.org/id/03B187B1B716FFA1255835CBFD3E26FA" lastPageNumber="109" pageId="2" pageNumber="109">
<subSubSection id="C302652CB716FFA1255835CBFC2922B7" box="[632,956,904,932]" pageId="2" pageNumber="109" type="nomenclature">
<paragraph id="8BA736A7B716FFA1255835CBFC2922B7" blockId="2.[366,1222,904,968]" box="[632,956,904,932]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1255835CBFC2922B7" box="[632,956,904,932]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="109" reason="2">
<taxonomicName id="4C184D24B716FFA1255835CBFCDF22B0" ID-CoL="ea79ab5d-6fff-49ee-b977-7109a8d15241" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[632,842,904,931]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA1255835CBFCDF22B0" bold="true" box="[632,842,904,931]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA1247135C9FC2922B7" box="[849,956,906,932]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1264E35EDFB5322DB" box="[366,1222,942,968]" pageId="2" pageNumber="109" type="description">
<paragraph id="8BA736A7B716FFA1264E35EDFB5322DB" blockId="2.[366,1222,904,968]" box="[366,1222,942,968]" pageId="2" pageNumber="109">
<uri id="FF893AA5B716FFA1264E35EDFB5322DB" box="[366,1222,942,968]" pageId="2" pageNumber="109">
urn:lsid:zoobank.org:act:
<uuid id="FFBE0C72B716FFA125AD35EDFB5322DB" box="[653,1222,942,968]" pageId="2" pageNumber="109">2BC93D12-96E1-4ECC-B83D-44249A06E099</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1279D35B2FCF82528" pageId="2" pageNumber="109" type="type_taxon">
<paragraph id="8BA736A7B716FFA1279D35B2FECD2518" blockId="2.[189,344,1009,1035]" box="[189,344,1009,1035]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1279D35B2FECD2518" bold="true" box="[189,344,1009,1035]" fontSize="11" level="3" pageId="2" pageNumber="109" reason="3">
<emphasis id="B96CEAB5B716FFA1279D35B2FECD2518" bold="true" box="[189,344,1009,1035]" pageId="2" pageNumber="109">
<typeStatus id="54A38805B716FFA1279D35B2FF6F2518" box="[189,250,1009,1035]" pageId="2" pageNumber="109">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736A7B716FFA1279D3262FCF82528" blockId="2.[189,877,1057,1084]" box="[189,877,1057,1084]" pageId="2" pageNumber="109">
<taxonomicName id="4C184D24B716FFA1279D3262FDB22528" ID-CoL="5a94bc9b-53c2-4fe0-8333-cc7ca7fa548e" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[189,551,1057,1083]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
<emphasis id="B96CEAB5B716FFA1279D3262FDB22528" box="[189,551,1057,1083]" italics="true" pageId="2" pageNumber="109">Dichromatobolus elephantulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA1250E3261FD4B252F" box="[558,734,1058,1084]" pageId="2" pageNumber="109" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1279D3227FEBD24A2" pageId="2" pageNumber="109" type="diagnosis">
<paragraph id="8BA736A7B716FFA1279D3227FEA3256D" blockId="2.[189,310,1124,1150]" box="[189,310,1124,1150]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1279D3227FEA3256D" bold="true" box="[189,310,1124,1150]" fontSize="11" level="3" pageId="2" pageNumber="109" reason="3">
<emphasis id="B96CEAB5B716FFA1279D3227FEA3256D" bold="true" box="[189,310,1124,1150]" pageId="2" pageNumber="109">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736A7B716FFA1279D32D6FEBD24A2" blockId="2.[189,1399,1173,1457]" pageId="2" pageNumber="109">
Massive, short, differs from all other known genera of
<taxonomicName id="4C184D24B716FFA1246932D6FC7225BC" authorityName="Cook, 1897 sensu Hoffman" authorityYear="1980" box="[841,999,1173,1199]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="family">Pachybolidae</taxonomicName>
in matUre males being only eight times longer than wide. Coloration UniqUely differing between sexes, females grey, males red (
<figureCitation id="13232A22B716FFA1222B32FAFAFD25C7" box="[1291,1384,1209,1236]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="2" pageNumber="109">Fig. 1A</figureCitation>
). Transverse ridge on mentum of gnathochilarium absent (
<figureCitation id="13232A22B716FFA1245B329DFC4A25EA" box="[891,991,1246,1273]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="2" pageNumber="109">Fig. 4A</figureCitation>
). Male legs with tarsal pads and unusually spinose tarsi (
<figureCitation id="13232A22B716FFA126F73340FDA3240E" box="[471,566,1283,1309]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="2" pageNumber="109">Fig. 4D</figureCitation>
). Shares anterior telopods with apical process of telopodite (
<figureCitation id="13232A22B716FFA123D73340FAAE240E" box="[1271,1339,1283,1309]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="2" pageNumber="109">Fig. 4</figureCitation>
E–F) with
<taxonomicName id="4C184D24B716FFA127D9336BFDF22451" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[249,615,1320,1347]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAB5B716FFA127D9336BFE252451" box="[249,432,1320,1346]" italics="true" pageId="2" pageNumber="109">Zehntnerobolus</emphasis>
Wesener, 2009
</taxonomicName>
, from which
<taxonomicName id="4C184D24B716FFA12423336BFC5A2451" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[771,975,1320,1346]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA12423336BFC5A2451" box="[771,975,1320,1346]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA124F7336AFBD72450" box="[983,1090,1321,1347]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
differs strongly in habitUs and posterior gonopods. Simple, apically swollen posterior gonopods (
<figureCitation id="13232A22B716FFA12301330EFBFF247B" box="[1057,1130,1357,1384]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="2" pageNumber="109">Fig. 5</figureCitation>
A–C) are shared with
<taxonomicName id="4C184D24B716FFA1279D3331FD83249F" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[189,534,1394,1421]" class="Diplopoda" family="Pachybolidae" genus="Corallobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAB5B716FFA1279D3331FECD249F" box="[189,344,1394,1420]" italics="true" pageId="2" pageNumber="109">Corallobolus</emphasis>
Wesener, 2009
</taxonomicName>
, from which
<taxonomicName id="4C184D24B716FFA1259D3331FC1C249F" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[701,905,1394,1420]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA1259D3331FC1C249F" box="[701,905,1394,1420]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA124B43330FB96249E" box="[916,1027,1395,1421]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
differs in habitUs and anterior telopods.
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1279D3399FDE1275A" pageId="2" pageNumber="109" type="etymology">
<paragraph id="8BA736A7B716FFA1279D3399FED124E7" blockId="2.[189,324,1498,1524]" box="[189,324,1498,1524]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1279D3399FED124E7" bold="true" box="[189,324,1498,1524]" fontSize="11" level="3" pageId="2" pageNumber="109" reason="3">
<emphasis id="B96CEAB5B716FFA1279D3399FED124E7" bold="true" box="[189,324,1498,1524]" pageId="2" pageNumber="109">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736A7B716FFA1279D3049FDE1275A" blockId="2.[189,1398,1546,1610]" pageId="2" pageNumber="109">
After the differences in the color pattern between the sexes, and ʻ-bolUsʼ, a common sUffix for genUs names of the
<taxonomicName id="4C184D24B716FFA1267A306CFE72275A" authorityName="Cook" authorityYear="1895" box="[346,487,1583,1609]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. Masculine.
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1279D3031FADF2615" pageId="2" pageNumber="109" type="description">
<paragraph id="8BA736A7B716FFA1279D3031FE24279F" blockId="2.[189,433,1650,1676]" box="[189,433,1650,1676]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1279D3031FE24279F" bold="true" box="[189,433,1650,1676]" fontSize="11" level="3" pageId="2" pageNumber="109" reason="3">
<emphasis id="B96CEAB5B716FFA1279D3031FE24279F" bold="true" box="[189,433,1650,1676]" pageId="2" pageNumber="109">Taxonomic position</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736A7B716FFA1279D30E1FADF2615" blockId="2.[189,1399,1698,1799]" pageId="2" pageNumber="109">
The posterior gonopods being connected by a sclerotized sternite as well as the absence of apodemes on the anterior gonopods (
<figureCitation id="13232A22B716FFA126DB3084FDD727F1" box="[507,578,1735,1762]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="2" pageNumber="109">Fig. 4</figureCitation>
E–F) place
<taxonomicName id="4C184D24B716FFA125EE3084FC0F27F2" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[718,922,1735,1761]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA125EE3084FC0F27F2" box="[718,922,1735,1761]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA12484308BFB8427F1" box="[932,1041,1736,1762]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
in the suborder
<taxonomicName id="4C184D24B716FFA123F83084FEC82615" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subOrder" subOrder="Trigoniulidea">Trigoniulidea Attems, 1909</taxonomicName>
, family
<taxonomicName id="4C184D24B716FFA1269E30AFFDF52615" authorityName="Cook, 1897 sensu Hoffman" authorityYear="1980" box="[446,608,1772,1798]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="family">Pachybolidae</taxonomicName>
. The closest relative of
<taxonomicName id="4C184D24B716FFA1245530AFFBD42615" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[885,1089,1772,1798]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA1245530AFFBD42615" box="[885,1089,1772,1798]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA1236930AEFB262614" box="[1097,1203,1773,1799]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C302652CB716FFA1279D3173FD3E26FA" pageId="2" pageNumber="109" type="discussion">
<paragraph id="8BA736A7B716FFA1279D3173FEBA2659" blockId="2.[189,303,1840,1866]" box="[189,303,1840,1866]" pageId="2" pageNumber="109">
<heading id="D0EF81CBB716FFA1279D3173FEBA2659" bold="true" box="[189,303,1840,1866]" fontSize="11" level="3" pageId="2" pageNumber="109" reason="3">
<emphasis id="B96CEAB5B716FFA1279D3173FEBA2659" bold="true" box="[189,303,1840,1866]" pageId="2" pageNumber="109">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736A7B716FFA1279D3123FD3E26FA" blockId="2.[189,1399,1888,2025]" pageId="2" pageNumber="109">
There is a second potential species,&gt;
<quantity id="4CE09B42B716FFA1254E3123FD582669" box="[622,717,1888,1914]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="109" unit="mm" value="100.0">100 mm</quantity>
in length, with specimens from Manongarivo, Ananambao (FMNH-INS 56254), all unfortunately female. Their wide body with short legs and grey color (all other larger
<taxonomicName id="4C184D24B716FFA1267131EAFE4F26D0" authorityName="Cook" authorityYear="1895" box="[337,474,1961,1987]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
genera on
<collectingCountry id="F30F7637B716FFA1254431EAFD6726D0" box="[612,754,1961,1987]" name="Madagascar" pageId="2" pageNumber="109">Madagascar</collectingCountry>
are black/ red, Wesener
<emphasis id="B96CEAB5B716FFA1233E31E9FBCE26D0" box="[1054,1115,1961,1987]" italics="true" pageId="2" pageNumber="109">et al.</emphasis>
2009) might indicate a relationship to
<taxonomicName id="4C184D24B716FFA1264D318DFDAC26FB" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[365,569,1998,2024]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAB5B716FFA1264D318DFDAC26FB" box="[365,569,1998,2024]" italics="true" pageId="2" pageNumber="109">Dichromatobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57CEB716FFA12561318CFD3E26FA" box="[577,683,1999,2025]" pageId="2" pageNumber="109" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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