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<document ID-DOI="10.37520/fi.2022.009" ID-ISSN="2533-4069" ID-Zenodo-Dep="7167867" approvalRequired="108" approvalRequired_for_taxonomicNames="33" approvalRequired_for_textStreams="71" approvalRequired_for_treatments="4" checkinTime="1665263286735" checkinUser="felipe" docAuthor="Denk, Thomas, Sami, Marco, Teodoridis, Vasilis &amp; Martinetto, Edoardo" docDate="2022" docId="03B187ADFFB8FFF7CC9EFACCFA8A4962" docLanguage="en" docName="FossilImprint.78.1.217-262.pdf" docOrigin="Fossil Imprint 78 (1)" docSource="http://dx.doi.org/10.37520/fi.2022.009" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.2:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="2" docTitle="Tsuga chiarugii TONGIORGI 1936" docType="treatment" docVersion="2" lastPageNumber="221" masterDocId="FF88FFD5FFBCFFF3CC67FFC0FF854812" masterDocTitle="The Late Early Pleistocene Flora Of Oriolo, Faenza (Italy): Assembly Of The Modern Forest Biome" masterLastPageNumber="262" masterPageNumber="217" pageNumber="221" updateTime="1665275026667" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title>The Late Early Pleistocene Flora Of Oriolo, Faenza (Italy): Assembly Of The Modern Forest Biome</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Denk, Thomas</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sami, Marco</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Teodoridis, Vasilis</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinetto, Edoardo</mods:namePart>
</mods:name>
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<mods:title>Fossil Imprint</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-26</mods:number>
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<mods:detail type="volume">
<mods:number>78</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:url>http://dx.doi.org/10.37520/fi.2022.009</mods:url>
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<mods:identifier type="DOI">10.37520/fi.2022.009</mods:identifier>
<mods:identifier type="ISSN">2533-4069</mods:identifier>
<mods:identifier type="Zenodo-Dep">7167867</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03B187ADFFB8FFF7CC9EFACCFA8A4962" httpUri="http://treatment.plazi.org/id/03B187ADFFB8FFF7CC9EFACCFA8A4962" lastPageNumber="221" pageId="4" pageNumber="221">
<subSubSection box="[249,646,1291,1316]" pageId="4" pageNumber="221" type="nomenclature">
<paragraph blockId="4.[249,646,1291,1316]" box="[249,646,1291,1316]" pageId="4" pageNumber="221">
<heading box="[249,646,1291,1316]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="221" reason="2">
<taxonomicName authority="TONGIORGI, 1936" authorityName="TONGIORGI" authorityYear="1936" box="[249,646,1291,1316]" class="Pinopsida" family="Pinaceae" genus="Tsuga" isUncertain="true" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="species" species="chiarugii">
<emphasis bold="true" box="[249,646,1291,1316]" pageId="4" pageNumber="221">
<emphasis bold="true" box="[249,315,1292,1315]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
cf.
<emphasis bold="true" box="[355,456,1291,1315]" italics="true" pageId="4" pageNumber="221">chiarugii</emphasis>
<bibRefCitation author="Tongiorgi, E." box="[463,581,1292,1316]" pageId="4" pageNumber="221" pagination="785 - 830" refId="ref33729" refString="Tongiorgi, E. (1936): Ricerche sulla vegetazione dell'Etruria marittima. V: Documenti per la storia della vegetazione della Toscana e del Lazio. - Nuovo Giornale Botanico Italiano, Nuova serie, 43 (4): 785 - 830. https: // doi. org / 10.1080 / 11263503609438744" type="journal article" year="1936">TONGIORGI</bibRefCitation>
, 1936
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="221" type="description">
<paragraph blockId="4.[373,523,1323,1347]" box="[373,523,1323,1347]" pageId="4" pageNumber="221">
<figureCitation box="[373,523,1323,1347]" captionStart="Text-fig" captionStartId="5.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-85@5.[125,1440,151,1950]" captionTargetPageId="5" captionText="Text-fig. 2. a, b: Pinus aff. peuce cones. a: Oriolo, MSF 643. b: Tebano, MSF 1021. c: Abies aff. alba cone scale, Oriolo MSF 988. d, e: Tsuga chiarugii cones. d: Oriolo MSF 638. e: Oriolo MSF 979. f, h: Bambusa lugdunensis leaves and leafy axis. f: Oriolo MSF 937. h: Oriolo MSF 651. g: Modern leaf of Yushania for comparison (NMNS Cleared Leaf Database specimen U1347). i: Phragmites sp. Oriolo MSF n.n. Scale bars 50 mm (a, b, h, i), 10 mm (cg)." figureDoi="http://doi.org/10.5281/zenodo.7167871" httpUri="https://zenodo.org/record/7167871/files/figure.png" pageId="4" pageNumber="221">Text-fig. 2d, e</figureCitation>
</paragraph>
<paragraph blockId="4.[128,767,1370,1509]" pageId="4" pageNumber="221">
1936
<taxonomicName authority="TONGIORGI" authorityName="TONGIORGI" authorityYear="1936" box="[204,470,1370,1391]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="species" species="chiarugii">
<emphasis box="[204,360,1370,1391]" italics="true" pageId="4" pageNumber="221">Tsuga chiarugii</emphasis>
TONGIORGI
</taxonomicName>
, p. 800, pl. 11, figs 14, 7, pl. 12, figs 510.
</paragraph>
<paragraph blockId="4.[128,767,1370,1509]" pageId="4" pageNumber="221">
2014b
<taxonomicName authority="TONGIORGI" authorityName="TONGIORGI" authorityYear="1936" box="[204,460,1428,1450]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="species" species="chiarugii">
<emphasis box="[204,355,1428,1450]" italics="true" pageId="4" pageNumber="221">Tsuga chiarugii</emphasis>
TONGIORGI
</taxonomicName>
; Pini et al., p. 303, fig. 11.1.10, P, Q.
</paragraph>
<paragraph blockId="4.[128,767,1370,1509]" box="[128,734,1487,1509]" pageId="4" pageNumber="221">
2015
<taxonomicName authority="TONGIORGI" authorityName="TONGIORGI" authorityYear="1936" box="[204,489,1487,1509]" class="Pinopsida" family="Pinaceae" genus="Tsuga" isUncertain="true" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="species" species="chiarugii">
<emphasis box="[204,260,1488,1509]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
cf.
<emphasis box="[297,385,1487,1508]" italics="true" pageId="4" pageNumber="221">chiarugii</emphasis>
TONGIORGI
</taxonomicName>
; Martinetto et al., p. 155.
</paragraph>
<paragraph blockId="4.[166,603,1533,1557]" box="[166,603,1533,1557]" pageId="4" pageNumber="221">M a t e r i a l. Oriolo MSF 638, 976, 979.</paragraph>
<paragraph blockId="4.[129,768,1580,1764]" pageId="4" pageNumber="221">
D e s c r i p t i o n. Two cone impressions accompanied by reddish iron minerals originate from the typical leafbearing sediments; another specimen is a 3-dimensionally preserved cast within a globose concrection originating from another layer of the Oriolo section. Cones are
<quantity box="[635,711,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="221" unit="mm" value="28.0">28 mm</quantity>
long and
<quantity box="[174,247,1740,1764]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="221" unit="mm" value="11.0">11 mm</quantity>
wide.
</paragraph>
<paragraph blockId="4.[129,768,1787,2100]" lastBlockId="4.[843,1482,152,368]" pageId="4" pageNumber="221">
R e m a r k s. The two cone impressions are compatible with the characters shown by mummified cones of
<taxonomicName authorityName="TONGIORGI" authorityYear="1936" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="species" species="chiarugii">
<emphasis italics="true" pageId="4" pageNumber="221">Tsuga chiarugii</emphasis>
</taxonomicName>
from Ghirlanda, a deposit of possible Early Pleistocene age in Central
<collectingCountry box="[417,466,1883,1907]" name="Italy" pageId="4" pageNumber="221">Italy</collectingCountry>
(
<bibRefCitation author="Tongiorgi, E." box="[480,651,1883,1907]" pageId="4" pageNumber="221" pagination="785 - 830" refId="ref33729" refString="Tongiorgi, E. (1936): Ricerche sulla vegetazione dell'Etruria marittima. V: Documenti per la storia della vegetazione della Toscana e del Lazio. - Nuovo Giornale Botanico Italiano, Nuova serie, 43 (4): 785 - 830. https: // doi. org / 10.1080 / 11263503609438744" type="journal article" year="1936">Tongiorgi 1936</bibRefCitation>
), a species that has a mass occurrence in the Northern Italian site of Steggio (
<bibRefCitation author="Ghiotto, P." box="[221,365,1947,1972]" pageId="4" pageNumber="221" pagination="3 - 99" refId="ref28529" refString="Ghiotto, P. (2010): La carpoflora del bacino lacustre villafranchiano di Steggio (Treviso, Prealpi orientali). - Bollettino del Museo regionale di Scienze naturali di Torino, 27: 3 - 99." type="journal article" year="2010">Ghiotto 2010</bibRefCitation>
,
<bibRefCitation author="Pini, R. &amp; Bertini, A. &amp; Martinetto, E. &amp; Vassio, E." box="[373,546,1947,1971]" pageId="4" pageNumber="221" pagination="290 - 307" refId="ref31903" refString="Pini, R., Bertini, A., Martinetto, E., Vassio, E. (2014 b): The Pleistocene flora of northern Italy. - In: Kustatscher, E., Roghi, G., Bertini, A., Miola, A. (eds), Palaeobotany of Italy. Publication of the Museum of Nature South Tyrol no. 9. Bozen, Museum of Nature South Tyrol, pp. 290 - 307." type="book chapter" year="2014">Pini et al. 2014b</bibRefCitation>
).
<taxonomicName authorityName="CARRIERE" authorityYear="1855" baseAuthorityName="ENDL." box="[561,624,1948,1971]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis box="[561,624,1948,1971]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
</taxonomicName>
was common and diverse during the Early and Middle Pleistocene of southern Europe (
<bibRefCitation author="Magri, D. &amp; Di Rita, F. &amp; Aranbarri, J. &amp; Fletcher, W. &amp; Gonzalez-Samperiz, P." box="[317,503,2012,2036]" pageId="4" pageNumber="221" pagination="23 - 55" refId="ref30301" refString="Magri, D., Di Rita, F., Aranbarri, J., Fletcher, W., Gonzalez-Samperiz, P. (2017): Quaternary disappearance of tree taxa from Southern Europe: timing and trends. - Quaternary Science Review, 163: 23 - 55. https: // doi. org / 10.1016 / j. quascirev. 2017.02.014" type="journal article" year="2017">Magri et al. 2017</bibRefCitation>
,
<bibRefCitation author="Denk, T. &amp; Guner, H. T. &amp; Bouchal, J. M. &amp; Kallanxhi, M. - E." box="[513,692,2012,2036]" pageId="4" pageNumber="221" pagination="283 - 305" refId="ref27953" refString="Denk, T., Guner, H. T., Bouchal, J. M., Kallanxhi, M. - E. (2021): The Pleistocene flora of Bezhan, southeast Albania: early appearance of extant tree species. - Historical Biology, 33: 283 - 305. https: // doi. org / 10.1080 / 08912963.2019.1615061" type="journal article" year="2021">Denk et al. 2021</bibRefCitation>
). From the older Calabrian site of Bezhan in
<collectingCountry box="[545,632,2044,2068]" name="Albania" pageId="4" pageNumber="221">Albania</collectingCountry>
,
<bibRefCitation author="Denk, T. &amp; Guner, H. T. &amp; Bouchal, J. M. &amp; Kallanxhi, M. - E." pageId="4" pageNumber="221" pagination="283 - 305" refId="ref27953" refString="Denk, T., Guner, H. T., Bouchal, J. M., Kallanxhi, M. - E. (2021): The Pleistocene flora of Bezhan, southeast Albania: early appearance of extant tree species. - Historical Biology, 33: 283 - 305. https: // doi. org / 10.1080 / 08912963.2019.1615061" type="journal article" year="2021">Denk et al. (2021)</bibRefCitation>
reported three pollen taxa of
<taxonomicName authorityName="CARRIERE" authorityYear="1855" baseAuthorityName="ENDL." box="[520,583,2076,2099]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis box="[520,583,2076,2099]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
</taxonomicName>
with affinities to modern Chinese, Japanese, and North American species. According to
<bibRefCitation author="Magri, D. &amp; Di Rita, F. &amp; Aranbarri, J. &amp; Fletcher, W. &amp; Gonzalez-Samperiz, P." box="[995,1207,184,208]" pageId="4" pageNumber="221" pagination="23 - 55" refId="ref30301" refString="Magri, D., Di Rita, F., Aranbarri, J., Fletcher, W., Gonzalez-Samperiz, P. (2017): Quaternary disappearance of tree taxa from Southern Europe: timing and trends. - Quaternary Science Review, 163: 23 - 55. https: // doi. org / 10.1016 / j. quascirev. 2017.02.014" type="journal article" year="2017">Magri et al. (2017)</bibRefCitation>
,
<taxonomicName authorityName="CARRIERE" authorityYear="1855" baseAuthorityName="ENDL." box="[1221,1284,184,207]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis box="[1221,1284,184,207]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
</taxonomicName>
disappeared from Northern
<collectingCountry box="[950,999,216,240]" name="Italy" pageId="4" pageNumber="221">Italy</collectingCountry>
(north of 44°) during the latest Calabrian, while it persisted slightly longer in Central
<collectingCountry box="[1315,1366,248,272]" name="Italy" pageId="4" pageNumber="221">Italy</collectingCountry>
.
<bibRefCitation author="Pini, R. &amp; Castellano, L. &amp; Ravazzi, C. &amp; Raineri, L. &amp; Bona, F. &amp; Sala, B." pageId="4" pageNumber="221" pagination="211 - 212" refId="ref31990" refString="Pini, R., Castellano, L., Ravazzi, C., Raineri, L., Bona, F., Sala, B. (2014 a): First combined findings of in-situ fossil wood and pollen of Parrotia persica and cf. Eucommia sp. in late Early Pleistocene deposits from the northern Apennine fringe (N Italy). - In: Abstract volume 9 th EPPC, Padua, pp. 211 - 212." type="book chapter" year="2014">Pini et al. (2014a: 299)</bibRefCitation>
stated that pollen of
<taxonomicName authorityName="CARRIERE" authorityYear="1855" baseAuthorityName="ENDL." box="[1214,1277,280,303]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis box="[1214,1277,280,303]" italics="true" pageId="4" pageNumber="221">Tsuga</emphasis>
</taxonomicName>
is “almost absent” in Northern
<collectingCountry box="[977,1026,312,336]" name="Italy" pageId="4" pageNumber="221">Italy</collectingCountry>
after 0.87 Ma; hence, sporadic records in the earliest Chibanian cannot be excluded.
</paragraph>
</subSubSection>
</treatment>
</document>