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<document id="70A34FB139C403C2809CC8175AF528A5" ID-DOI="10.5281/zenodo.177498" ID-GBIF-Dataset="f5ce3ded-95df-45a1-b0f9-f4944458e064" ID-ISSN="1175-5326" ID-Zenodo-Dep="177498" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459863126866" checkinUser="plazi" docAuthor="Cumberlidge, Neil, Marijnissen, Saskia A. E. &amp; Thompson, Jonelle" docDate="2007" docId="038D87FCF311FFCBFF3EFDE16C2BFD59" docLanguage="en" docName="zt01524p068.pdf" docOrigin="Zootaxa 1524" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydrothelphusa vencesi Cumberlidge, Marijnissen &amp; Thompson, 2007, n. sp." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFB4FF84F310FFCDFFA9FE76694FFFCB" masterDocTitle="Hydrothelphusa vencesi, a new species of freshwater crab (Brachyura: Potamoidea: Potamonautidae) from southeastern Madagascar." masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698226250642" updateUser="plazi">
<mods:mods id="284E1205272618A1FBA5F15FB5091375" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C4E9CFC0873A0D9571FF47D50E3F1CAC">Hydrothelphusa vencesi, a new species of freshwater crab (Brachyura: Potamoidea: Potamonautidae) from southeastern Madagascar.</mods:title>
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<mods:name id="C430F6FC111434F5DADE56A3C1BE8FC7" type="personal">
<mods:role id="0D08C131303EEDFAF0D7780BB8E7C69F">
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<mods:namePart id="7F10E2AABA5AE309EA6DF40DAD997D71">Cumberlidge, Neil</mods:namePart>
</mods:name>
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<mods:role id="79A79B2BD75EA880342E344062694479">
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<mods:namePart id="053F9CCB2A5ED6C471B9A803C0BDE1C9">Marijnissen, Saskia A. E.</mods:namePart>
</mods:name>
<mods:name id="4B836B760287617E3F35A10306D3E2A0" type="personal">
<mods:role id="4868F96A784044F183F45ADC8F643BDF">
<mods:roleTerm id="A5E4E9594F3C33F3CCC3F6B4431A985A">Author</mods:roleTerm>
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<mods:namePart id="040304A8563B43C329357725C5DEBA70">Thompson, Jonelle</mods:namePart>
</mods:name>
<mods:typeOfResource id="5D08202632886B324E48E57F7378D05A">text</mods:typeOfResource>
<mods:relatedItem id="B43EDC1ABB4CB2703B800F3E7DD0BB5E" type="host">
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<mods:title id="336CAEC719A22E5A39DAD63F77B96CD0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="040509782DB51B6FBD34A4EE8CCA7754">
<mods:date id="FBAD1CECE4FADEAC09AEB13667D09D6F">2007</mods:date>
<mods:detail id="4508CD52F98FB0DFD95A5EB981DE3132" type="volume">
<mods:number id="646948FDAAF09CD81D4649E5801C53C4">1524</mods:number>
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<mods:start id="78A472ED22457AFB6C5A0BFDECBD3DBE">61</mods:start>
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<mods:classification id="432036BFD616E925EBB235ED376E4958">journal article</mods:classification>
<mods:identifier id="D9058ED0F58EA181C5406D4D90AFC15A" type="DOI">10.5281/zenodo.177498</mods:identifier>
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<mods:identifier id="688E834327455945561FF1ED393D2488" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4E47BE79F4828B692EEF4DDBF13C398B" type="Zenodo-Dep">177498</mods:identifier>
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<treatment id="038D87FCF311FFCBFF3EFDE16C2BFD59" ID-DOI="http://doi.org/10.5281/zenodo.6251924" ID-GBIF-Taxon="119368816" ID-Zenodo-Dep="6251924" LSID="urn:lsid:plazi:treatment:038D87FCF311FFCBFF3EFDE16C2BFD59" httpUri="http://treatment.plazi.org/id/038D87FCF311FFCBFF3EFDE16C2BFD59" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62">
<subSubSection id="C33E6561F311FFCCFF3EFDE16869FC1F" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8B9B36EAF311FFCCFF3EFDE16B4CFC7A" blockId="1.[151,515,919,980]" box="[151,515,919,945]" pageId="1" pageNumber="62">
<heading id="D0D38186F311FFCCFF3EFDE16B4CFC7A" bold="true" box="[151,515,919,945]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="B950EAF8F311FFCCFF3EFDE16B4CFC7A" bold="true" box="[151,515,919,945]" pageId="1" pageNumber="62">
<taxonomicName id="4C244D69F311FFCCFF3EFDE168F6FC7A" box="[151,441,919,945]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="vencesi" status="sp. nov.">
<emphasis id="B950EAF8F311FFCCFF3EFDE168F6FC7A" bold="true" box="[151,441,919,945]" italics="true" pageId="1" pageNumber="62">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635783F311FFCCFE68FDE16B4CFC7A" box="[449,515,919,945]" pageId="1" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B36EAF311FFCCFF3EFDCC6869FC1F" blockId="1.[151,515,919,980]" box="[151,294,954,980]" pageId="1" pageNumber="62">
(
<figureCitation id="131F2A6FF311FFCCFF36FDCC6852FC1F" box="[159,285,954,980]" captionStart-0="FIGURES 1 9" captionStart-1="FIGURES 10 15" captionStart-2="FIGURES 16 17" captionStartId-0="3.[151,269,1843,1867]" captionStartId-1="4.[151,269,1545,1569]" captionStartId-2="5.[153,272,1151,1175]" captionTargetBox-0="[189,1410,219,1795]" captionTargetBox-1="[191,1406,216,1504]" captionTargetBox-2="[469,1117,197,1126]" captionTargetId-0="figure@3.[151,1436,197,1817]" captionTargetId-1="figure@4.[164,1424,194,1521]" captionTargetId-2="figure@5.[468,1118,194,1127]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." captionText-1="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." captionText-2="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri-0="https://zenodo.org/record/177499/files/figure.png" httpUri-1="https://zenodo.org/record/177500/files/figure.png" httpUri-2="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">Figs. 117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F311FFCCFF3EFA7C6A70FADF" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="8B9B36EAF311FFCCFF3EFA7C6CD9FB57" blockId="1.[151,1436,1034,2020]" pageId="1" pageNumber="62">
<emphasis id="B950EAF8F311FFCCFF3EFA7C6BF7FBEF" bold="true" box="[151,696,1034,1060]" pageId="1" pageNumber="62">
<typeStatus id="549F8848F311FFCCFF3EFA7C699BFBEF" box="[151,212,1034,1060]" pageId="1" pageNumber="62">Type</typeStatus>
material examined:
<collectingCountry id="F333767AF311FFCCFE7EFA7C6BFCFBEF" box="[471,691,1034,1060]" name="Madagascar" pageId="1" pageNumber="62">MADAGASCAR</collectingCountry>
:
</emphasis>
adult male,
<typeStatus id="549F8848F311FFCCFCE3FA7C6AFFFBEF" box="[842,944,1034,1060]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
(designated here, cw 52, cl 40.5, ch 19.5, fw
<quantity id="4CDC9B0FF311FFCCFF17FA446864FB87" box="[190,299,1074,1100]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="62" unit="mm" value="16.5">16.5 mm</quantity>
), Fianarantsoa Province: region of Vondrozo near village of Vevembe (
<geoCoordinate id="EE10502DF311FFCCFBDCFA446DB5FB87" box="[1141,1274,1074,1100]" direction="south" orientation="latitude" pageId="1" pageNumber="62" precision="15" value="-22.795555">22º47'44&quot;S</geoCoordinate>
,
<geoCoordinate id="EE10502DF311FFCCFAAEFA446CC1FB87" box="[1287,1422,1074,1100]" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="15" value="47.188614">47º11'19&quot;E</geoCoordinate>
), Ramanara River, tributary of Sahampindra River, Mananara River drainage basin,
<quantity id="4CDC9B0FF311FFCCFB30FA2C6DB0FBBF" box="[1177,1279,1114,1140]" metricMagnitude="4" metricUnit="m" metricValue="6.659999999999999" pageId="1" pageNumber="62" unit="km" value="66.6">66.6 km</quantity>
inland from Farafangana, in a forested area in the mountains of southeastern
<collectingCountry id="F333767AF311FFCCFC2BFAF46D40FB57" box="[898,1039,1154,1180]" name="Madagascar" pageId="1" pageNumber="62">Madagascar</collectingCountry>
(M. Vences) (
<collectionCode id="ED35AE2FF311FFCCFB1AFAF46DB9FB57" box="[1203,1270,1154,1180]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="62">ZMA</collectionCode>
De. 205976).
</paragraph>
<paragraph id="8B9B36EAF311FFCCFF6FFADC6A70FADF" blockId="1.[151,1436,1034,2020]" pageId="1" pageNumber="62">
<emphasis id="B950EAF8F311FFCCFF6FFADC6A75FB0F" bold="true" box="[198,826,1194,1220]" pageId="1" pageNumber="62">
Additional material examined:
<collectingCountry id="F333767AF311FFCCFDF3FADC6A7AFB0F" box="[602,821,1194,1220]" name="Madagascar" pageId="1" pageNumber="62">MADAGASCAR</collectingCountry>
:
</emphasis>
<collectionCode id="ED35AE2FF311FFCCFCEFFADC6AD4FB0F" LSID="urn:lsid:biocol.org:col:34795" box="[838,923,1194,1220]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="62">FMNH</collectionCode>
5730, Sahavatov River, basin of Manampatrana River, Andringita Camp II,
<quantity id="4CDC9B0FF311FFCCFD97FAA46BD9FB27" box="[574,662,1234,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="1" pageNumber="62" unit="m" value="1390.0">1,390m</quantity>
asl (
<date id="FF9A102AF311FFCCFD79FAA46A18FB27" box="[720,855,1234,1260]" pageId="1" pageNumber="62" value="1993-11-20">20-Nov-93</date>
);
<collectionCode id="ED35AE2FF311FFCCFCC5FAA46A8EFB27" LSID="urn:lsid:biocol.org:col:34795" box="[876,961,1234,1260]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="62">FMNH</collectionCode>
5748, Sahambano River, Mananara-sud River drainage basin,
<collectingCountry id="F333767AF311FFCCFE31FA8C6B68FADF" box="[408,551,1274,1300]" name="Madagascar" pageId="1" pageNumber="62">Madagascar</collectingCountry>
,
<quantity id="4CDC9B0FF311FFCCFD9AFA8C6B32FADF" box="[563,637,1274,1300]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="62" unit="m" value="570.0">570 m</quantity>
asl,
<date id="FF9A102AF311FFCCFD1AFA8C6A74FADF" box="[691,827,1274,1300]" pageId="1" pageNumber="62" value="1995-05-25">25-May-95</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F311FFCCFF6FFB546CD9FA17" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="8B9B36EAF311FFCCFF6FFB546CD9FA17" blockId="1.[151,1436,1034,2020]" pageId="1" pageNumber="62">
<emphasis id="B950EAF8F311FFCCFF6FFB546809FAF7" bold="true" box="[198,326,1314,1340]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
Terminal article of G1 long (ratio of length of terminal article to subterminal segment 0.3), straight-sided cone, bent outward at 45° to longitudinal axis of gonopod, evenly-tapering to pointed tip; broad, diamond-shaped dorsal membrane at segment junction. Palm of propodus of major cheliped enlarged, dactylus slightly arched, enclosing long narrow interspace; propodus, and dactylus of major cheliped serrated, lined by small teeth interspersed by larger rounded teeth, with fused cluster of several small low teeth, tips pointed.
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F311FFCFFF6FFB9C6DA6FB81" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="description">
<paragraph id="8B9B36EAF311FFCFFF6FFB9C68C4FC19" blockId="1.[151,1436,1034,2020]" lastBlockId="2.[151,1437,152,1978]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
<emphasis id="B950EAF8F311FFCCFF6FFB9C68BCF9CF" bold="true" box="[198,499,1514,1540]" pageId="1" pageNumber="62">
Description of
<typeStatus id="549F8848F311FFCCFE29FB9C68A1F9CF" box="[384,494,1514,1540]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace moderately wide (cw/fw 3.2) moderately high (ch/fw 1.2); anterolateral, posterolateral regions of carapace distinctly granular, with short carinae (
<figureCitation id="131F2A6FF311FFCCFB94F8646DDEF9E7" box="[1085,1169,1554,1580]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFB08F8646D8DF9E7" box="[1185,1218,1554,1580]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16</figureCitation>
). Epigastric crests raised, anterior margins straight, positioned anterior to postorbital crests; mid-groove between epigastric crests broad, shallow; distinct gap between epigastric, postorbital crests; postorbital crests ending well before meeting anterolateral margins (
<figureCitation id="131F2A6FF311FFCCFDAAF8FC6B3CF96F" box="[515,627,1674,1700]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1, 2</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFDD6F8FC6BEFF96F" box="[639,672,1674,1700]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16</figureCitation>
). Semi-circular groove shallow, faint; urogastric, cardiac grooves shallow; cervical grooves long, ending before meeting postorbital crests (
<figureCitation id="131F2A6FF311FFCCFBAEF8C46D15F907" box="[1031,1114,1714,1740]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFBC2F8C46DC3F907" box="[1131,1164,1714,1740]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16</figureCitation>
). Front horizontal (not deflexed) leaving antennulular fossae exposed; upper edge of frontal margin granular, distinctly indented, lower edge V-shaped, broadest in middle, narrowest at sides; frontal margin of carapace approximately onethird cw (0.32) (
<figureCitation id="131F2A6FF311FFCCFEF3F95C6881F88F" box="[346,462,1834,1860]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1, 2</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFE75F95C6B66F88F" box="[476,553,1834,1860]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16, 17</figureCitation>
). Exorbital, epibranchial teeth large, pointed, directed forward, outer margins granular. Anterolateral margin between exorbital, epibranchial teeth lacking intermediate tooth (
<figureCitation id="131F2A6FF311FFCCFAEAF92469E7F85F" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1, 2</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFF1EF90C6849F85F" box="[183,262,1914,1940]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16, 17</figureCitation>
). Anterolateral margin posterior to epibranchial tooth raised, with small teeth grading into heavy granules (
<figureCitation id="131F2A6FF311FFCCFEA7F9D4682BF877" box="[270,356,1954,1980]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFEDFF9D468D8F877" box="[374,407,1954,1980]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">16</figureCitation>
). Suborbital margin raised, heavily granulated (
<figureCitation id="131F2A6FF311FFCCFC76F9D46D78F877" box="[991,1079,1954,1980]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFBEEF9D46D28F877" box="[1095,1127,1954,1980]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">17</figureCitation>
). Suborbital, pterygomial regions of carapace sidewall heavily granulated, subhepatic region with carinae (
<figureCitation id="131F2A6FF311FFCCFBEFF9BC6DD6F82F" box="[1094,1177,1994,2020]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
,
<figureCitation id="131F2A6FF311FFCCFB0CF9BC6D8AF82F" box="[1189,1221,1994,2020]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="1" pageNumber="62">17</figureCitation>
). Mandibular palp two-segmented, bilobed, with medium-sized anterior process (about one-third size of terminal segment). Ischium of third maxilliped with deep vertical sulcus (
<figureCitation id="131F2A6FF312FFCFFD5DFEB66A01FF11" box="[756,846,192,218]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="2" pageNumber="63">Fig. 17</figureCitation>
), exopod of third maxilliped with long flagellum (
<figureCitation id="131F2A6FF312FFCFFF36FE9E69A7FEC9" box="[159,232,232,258]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 7</figureCitation>
). Sternal sulci s2/s3, s3/s4 completely crossing sternum, s3/s4 slightly V-shaped, not meeting anterior margin of sterno-abdominal cavity on sternite 4 (
<figureCitation id="131F2A6FF312FFCFFD71FF666A50FEE1" box="[728,799,272,298]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
). Episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 all smooth lacking visible groove (
<figureCitation id="131F2A6FF312FFCFFE06FF4E68B8FE99" box="[431,503,312,338]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
). Palm of propodus of major (left) cheliped enlarged, dactylus slightly arched, enclosing long rectangular interspace; fingers of major cheliped lined by small even teeth interspersed by large rounded teeth, propodus with fused cluster of three or four small low teeth (
<figureCitation id="131F2A6FF312FFCFFBDBFFFE6DA6FE69" box="[1138,1257,392,418]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Figs. 3, 4</figureCitation>
). Walking legs (p2p5) of medium size (ratio of length of merus of p5 to cw 0.400.45) (
<figureCitation id="131F2A6FF312FFCFFC5CFFC66D04FE01" box="[1013,1099,432,458]" captionStart="FIGURES 16 17" captionStartId="5.[153,272,1151,1175]" captionTargetBox="[469,1117,197,1126]" captionTargetId="figure@5.[468,1118,194,1127]" captionTargetPageId="5" captionText="FIGURES 16 17. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view." httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="2" pageNumber="63">Fig. 16</figureCitation>
). Anterior inferior margin of ischium of pereiopod 1 granular; medial inferior margin of merus of pereiopod 1 heavily granulated, lateral inferior margin with several small teeth; distal meral tooth reduced to granule; superior surface of merus roughly granulated (
<figureCitation id="131F2A6FF312FFCFFE2DFC5E6883FD89" box="[388,460,552,578]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 6</figureCitation>
). Inner margin of carpus of cheliped with two large, pointed carpal teeth, first tooth spine-like, second tooth one-third size of first, margin behind second tooth with small tooth (
<figureCitation id="131F2A6FF312FFCFFB55FC266C08FDA1" box="[1276,1351,592,618]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 5</figureCitation>
). Male abdomen long, slim, triangular (a3 widest, a6 shorter than width of distal margin of a6), telson (a7) bellshaped with indented sides. Terminal article of G1 long (ratio of length of terminal article to subterminal segment 0.3), straight-sided cone, bent outward at 45° to longitudinal axis of gonopod, evenly-tapering to pointed tip (
<figureCitation id="131F2A6FF312FFCFFF6FFC866808FCC1" box="[198,327,752,778]" captionStart-0="FIGURES 1 9" captionStart-1="FIGURES 10 15" captionStartId-0="3.[151,269,1843,1867]" captionStartId-1="4.[151,269,1545,1569]" captionTargetBox-0="[189,1410,219,1795]" captionTargetBox-1="[191,1406,216,1504]" captionTargetId-0="figure@3.[151,1436,197,1817]" captionTargetId-1="figure@4.[164,1424,194,1521]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." captionText-1="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri-0="https://zenodo.org/record/177499/files/figure.png" httpUri-1="https://zenodo.org/record/177500/files/figure.png" pageId="2" pageNumber="63">Figs. 915</figureCitation>
); no visible groove between terminal article and subterminal segment on ventral side (
<figureCitation id="131F2A6FF312FFCFFAEAFC866CD8FCC1" box="[1347,1431,752,778]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Figs. 9</figureCitation>
,
<figureCitation id="131F2A6FF312FFCFFF3EFD6E69F8FCF9" box="[151,183,792,818]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="2" pageNumber="63">10</figureCitation>
); dorsal side with broad, diamond-shaped dorsal membrane at segment junction (
<figureCitation id="131F2A6FF312FFCFFBC7FD6E6DB3FCF9" box="[1134,1276,792,818]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="2" pageNumber="63">Figs. 11, 14</figureCitation>
). Ventral side of subterminal segment of G1 with long lateral flap folded inward; medial part of dorsal face of subterminal segment of G1 flat with distinct disto-medial triangular shoulder, and large rounded baso-lateral projection (
<figureCitation id="131F2A6FF312FFCFFF36FDE669B8FC61" box="[159,247,912,938]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="2" pageNumber="63">Fig. 10</figureCitation>
). G2 longer than G1; terminal article of G2 long flagellum, ratio length terminal article to subterminal segment 0.8 (
<figureCitation id="131F2A6FF312FFCFFE9CFDCE6833FC19" box="[309,380,952,978]" captionStart="FIGURES 1 9" captionStartId="3.[151,269,1843,1867]" captionTargetBox="[189,1410,219,1795]" captionTargetId="figure@3.[151,1436,197,1817]" captionTargetPageId="3" captionText="FIGURES 1 9. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (3 7)." httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="2" pageNumber="63">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B9B36EAF312FFCFFF6EFDA96D06FC31" blockId="2.[151,1437,152,1978]" box="[199,1097,991,1018]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EFDA9684FFC32" bold="true" box="[199,256,991,1017]" pageId="2" pageNumber="63">Size.</emphasis>
The largest specimen known is the
<typeStatus id="549F8848F312FFCFFD0DFD966A42FC31" box="[676,781,992,1018]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
, an adult male, cw
<quantity id="4CDC9B0FF312FFCFFC46FD966D09FC31" box="[1007,1094,992,1018]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="63" unit="mm" value="52.0">52 mm</quantity>
.
</paragraph>
<paragraph id="8B9B36EAF312FFCFFF6EFA716DA6FB81" blockId="2.[151,1437,152,1978]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EFA71685CFBEA" bold="true" box="[199,275,1031,1057]" pageId="2" pageNumber="63">Color.</emphasis>
In life, carapace purple-brown, lacking stains or spots, tips of chelipeds and pereiopods lighter shade of brown, granules on suborbital and pterygostomial regions of carapace sidewall beige.
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F312FFCFFF6EFA216DE6FB51" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="8B9B36EAF312FFCFFF6EFA216DE6FB51" blockId="2.[151,1437,152,1978]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EFA216822FBBA" bold="true" box="[199,365,1111,1137]" pageId="2" pageNumber="63">
<typeStatus id="549F8848F312FFCFFF6EFA21684BFBBA" box="[199,260,1111,1137]" pageId="2" pageNumber="63">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F333767AF312FFCFFEDCFA2E6B4AFBB9" box="[373,517,1112,1138]" name="Madagascar" pageId="2" pageNumber="63">Madagascar</collectingCountry>
, Vevembe Forest (
<geoCoordinate id="EE10502DF312FFCFFD49FA2E6A29FBB9" box="[736,870,1112,1138]" direction="south" orientation="latitude" pageId="2" pageNumber="63" precision="15" value="-22.795555">22º47'44&quot;S</geoCoordinate>
,
<geoCoordinate id="EE10502DF312FFCFFCDAFA2E6AB5FBB9" box="[883,1018,1112,1138]" direction="east" orientation="longitude" pageId="2" pageNumber="63" precision="15" value="47.188614">47º11'19&quot;E</geoCoordinate>
),
<quantity id="4CDC9B0FF312FFCFFBB9FA2E6D3EFBB9" box="[1040,1137,1112,1138]" metricMagnitude="4" metricUnit="m" metricValue="6.659999999999999" pageId="2" pageNumber="63" unit="km" value="66.6">66.6 km</quantity>
inland from Farafangana in Fianarantsoa Province in a forested area in the mountains of southeastern
<collectingCountry id="F333767AF312FFCFFBBCFAF66DEAFB51" box="[1045,1189,1152,1178]" name="Madagascar" pageId="2" pageNumber="63">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F312FFCFFF6EFAD16C10FAA9" pageId="2" pageNumber="63" type="distribution">
<paragraph id="8B9B36EAF312FFCFFF6EFAD16C10FAA9" blockId="2.[151,1437,152,1978]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EFAD16828FB0A" bold="true" box="[199,359,1191,1217]" pageId="2" pageNumber="63">Distribution.</emphasis>
Endemic to
<collectingCountry id="F333767AF312FFCFFDABFADE6BDDFB09" box="[514,658,1192,1218]" name="Madagascar" pageId="2" pageNumber="63">Madagascar</collectingCountry>
.
<taxonomicName id="4C244D69F312FFCFFD36FADE6AF5FB0A" box="[671,954,1192,1217]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F312FFCFFD36FADE6AF5FB0A" box="[671,954,1192,1217]" italics="true" pageId="2" pageNumber="63">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
is known from three localities in southeastern
<collectingCountry id="F333767AF312FFCFFF5BFAA668CDFB21" box="[242,386,1232,1258]" name="Madagascar" pageId="2" pageNumber="63">Madagascar</collectingCountry>
, two of them in the Mananara River drainage basin. The
<typeStatus id="549F8848F312FFCFFB85FAA66D10FB21" box="[1068,1119,1232,1258]" pageId="2" pageNumber="63">type</typeStatus>
locality is in the Vevembe Forest near the village of Vevembe in the region of Vondrozo in Fianarantsoa Province. This species is also found in the Sahavatov River which is part of the drainage basin of the Manampatrana River (Andringita Camp II,
<quantity id="4CDC9B0FF312FFCFFEAEFB3E6828FAA9" box="[263,359,1352,1378]" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="2" pageNumber="63" unit="m" value="1390.0">1,390 m</quantity>
asl), and at
<quantity id="4CDC9B0FF312FFCFFE5CFB3E6B70FAA9" box="[501,575,1352,1378]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="63" unit="m" value="570.0">570 m</quantity>
asl in the Sahambano River in the basin of the Mananara-sud River.
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F312FFCFFF6EFB196D7EF8A1" pageId="2" pageNumber="63" type="discussion">
<paragraph id="8B9B36EAF312FFCFFF6EFB196D7EF8A1" blockId="2.[151,1437,152,1978]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EFB19680FFA42" bold="true" box="[199,320,1391,1417]" pageId="2" pageNumber="63">Remarks.</emphasis>
The following morphological characters justify including
<taxonomicName id="4C244D69F312FFCFFC44FB066D28FA42" box="[1005,1127,1392,1417]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F312FFCFFC44FB066D28FA42" box="[1005,1127,1392,1417]" italics="true" pageId="2" pageNumber="63">H. vencesi</emphasis>
</taxonomicName>
in
<taxonomicName id="4C244D69F312FFCFFB25FB066C07FA42" box="[1164,1352,1392,1417]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF8F312FFCFFB25FB066C07FA42" box="[1164,1352,1392,1417]" italics="true" pageId="2" pageNumber="63">Hydrothelphusa</emphasis>
</taxonomicName>
(Cumberlidge &amp; Sternberg 2002: 47;
<bibRefCitation id="EFB54B1BF312FFCFFDA6FBEE6B9CFA79" author="Thompson" box="[527,723,1432,1458]" pageId="2" pageNumber="68" refString="Thompson, J. 2005. A molecular phylogenetic analysis of the Malagasy freshwater crab genus Hydrothelphusa H. Milne Edwards, 1872, with biogeographical implications for the group. M. S. Thesis, Department of Biology, Northern Michigan University, Marquette, Michigan MI, USA." type="book chapter" year="2005">Thompson 2005</bibRefCitation>
). The carapace outline is transversely oval with wide frontal and posterior margins (each approximately one-third carapace width); the mandibular palp is two-segmented with a lobe-like, medium-sized anterior process (about one-third the size of the terminal segment) at the junction between the segments; the ischium of the third maxilliped has a deep vertical sulcus, and the exopod of the third maxilliped has a long flagellum; sternal sulcus s2/s3 is deep and horizontal, and sternal sulcus s3/s4 is shallow and V-shaped; episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 are smooth and not visible; walking legs (p2p5) are neither shortened nor elongated, with the ratio of the length of the merus of p5 to the cw of 0.40 0.45; the male abdomen is long, slim, and triangular, with a bell-shaped telson whose sides are indented; the terminal article of G1 is long (with a ratio of the length of the terminal article to the subterminal segment of 0.3); the subterminal segment of G1 has a distinct disto-lateral triangular shoulder-like projection at the junction of the segments; and G2 is longer than G1 with a terminal article that has a long flagellum (the ratio of the length of the terminal article to the subterminal segment is approximately 0.8).
</paragraph>
</subSubSection>
<subSubSection id="C33E6561F312FFC9FF6EF9016DBBF852" lastPageId="4" lastPageNumber="65" pageId="2" pageNumber="63" type="etymology">
<paragraph id="8B9B36EAF312FFCFFF6EF9016C76F871" blockId="2.[151,1437,152,1978]" pageId="2" pageNumber="63">
<emphasis id="B950EAF8F312FFCFFF6EF901681CF85A" bold="true" box="[199,339,1911,1937]" pageId="2" pageNumber="63">Etymology.</emphasis>
The species is named for Prof. Dr. Miguel Vences in recognition of his important contributions to our knowledge of the diversity, evolution, and natural history of Madgascars endemic fauna.
</paragraph>
<caption id="DF5B6662F313FFCEFF3EF9456A11F81E" httpUri="https://zenodo.org/record/177499/files/figure.png" pageId="3" pageNumber="64" targetBox="[189,1410,219,1795]" targetPageId="3">
<paragraph id="8B9B36EAF313FFCEFF3EF9456A11F81E" blockId="3.[151,1436,1843,2005]" pageId="3" pageNumber="64">
<emphasis id="B950EAF8F313FFCEFF3EF945680AF880" bold="true" box="[151,325,1843,1867]" pageId="3" pageNumber="64">FIGURES 19.</emphasis>
<taxonomicName id="4C244D69F313FFCEFEE6F9456B1DF881" box="[335,594,1843,1866]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphsua" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="vencesi" status="sp. nov.">
<emphasis id="B950EAF8F313FFCEFEE6F9456B1DF881" box="[335,594,1843,1866]" italics="true" pageId="3" pageNumber="64">Hydrothelphsua vencesi</emphasis>
</taxonomicName>
<emphasis id="B950EAF8F313FFCEFDF5F9456BD4F880" bold="true" box="[604,667,1843,1867]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="A2635783F313FFCEFDF5F9456BD4F880" box="[604,667,1843,1867]" pageId="3" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 1, carapace and eyes, dorsal view; 2, cephalothorax, carapace and eyes, frontal view; 3, right cheliped, frontal view; 4, left cheliped, frontal view; 5, carpus and merus of right cheliped, dorsal view; 6, merus of right cheliped, inferior view; 7, left third maxilliped; 8, sternum and abdomen; 9, abdomen pulled back to show first and second gonopods. Scale = 13 mm (1,2, 8, 9); 7 mm (37).
</paragraph>
</caption>
<caption id="DF5B6662F314FFC9FF3EF87F6CD3F942" httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="4" pageNumber="65" targetBox="[191,1406,216,1504]" targetPageId="4">
<paragraph id="8B9B36EAF314FFC9FF3EF87F6CD3F942" blockId="4.[151,1436,1545,1673]" pageId="4" pageNumber="65">
<emphasis id="B950EAF8F314FFC9FF3EF87F6810F9EA" bold="true" box="[151,351,1545,1569]" pageId="4" pageNumber="65">FIGURES 1015.</emphasis>
<taxonomicName id="4C244D69F314FFC9FECFF87C6B27F9EA" box="[358,616,1546,1569]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphsua" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="vencesi" status="sp. nov.">
<emphasis id="B950EAF8F314FFC9FECFF87C6B27F9EA" box="[358,616,1546,1569]" italics="true" pageId="4" pageNumber="65">Hydrothelphsua vencesi</emphasis>
</taxonomicName>
<emphasis id="B950EAF8F314FFC9FDD9F87F6BE2F9EA" bold="true" box="[624,685,1545,1569]" pageId="4" pageNumber="65">
<taxonomicNameLabel id="A2635783F314FFC9FDD9F87F6BE2F9EA" box="[624,685,1545,1569]" pageId="4" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G1, ventral view; 11, right G1, dorsal view; 12, terminal article of right G1, dorsal view; 13, detail of tip of terminal article of right G1, dorsal view; 14, detail of tip of terminal article of right G1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (1215).
</paragraph>
</caption>
<paragraph id="8B9B36EAF314FFC9FF6EF8C16DBBF852" blockId="4.[151,1436,1719,1945]" pageId="4" pageNumber="65">
<emphasis id="B950EAF8F314FFC9FF6EF8C1683BF91A" bold="true" box="[199,372,1719,1745]" pageId="4" pageNumber="65">Comparisons.</emphasis>
<taxonomicName id="4C244D69F314FFC9FED4F8C16BD7F91B" box="[381,664,1719,1744]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F314FFC9FED4F8C16BD7F91B" box="[381,664,1719,1744]" italics="true" pageId="4" pageNumber="65">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
can be distinguished from other species of
<emphasis id="B950EAF8F314FFC9FB0AF8C16C11F91B" box="[1187,1374,1719,1744]" italics="true" pageId="4" pageNumber="65">Hydrothelphsua</emphasis>
(and indeed from all other
<collectingCountry id="F333767AF314FFC9FE31F8A96B45F932" box="[408,522,1759,1785]" name="Madagascar" pageId="4" pageNumber="65">Malagasy</collectingCountry>
brachyurans) by examination of G1 morphology (
<figureCitation id="131F2A6FF314FFC9FBF6F8A96DA0F932" box="[1119,1263,1759,1785]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="4" pageNumber="65">Figs. 1015</figureCitation>
). The terminal article of G1 of
<taxonomicName id="4C244D69F314FFC9FEF2F9716898F8EB" box="[347,471,1799,1824]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F314FFC9FEF2F9716898F8EB" box="[347,471,1799,1824]" italics="true" pageId="4" pageNumber="65">H. vencesi</emphasis>
</taxonomicName>
is a long straight-sided cone that tapers evenly to a pointed tip and lacks a pronounced raised mid section; in addition there is a broad diamond-shaped dorsal membrane at the segment junction on its dorsal side. These characters are unique to
<taxonomicName id="4C244D69F314FFC9FCE6F9216A85F8BB" box="[847,970,1879,1904]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F314FFC9FCE6F9216A85F8BB" box="[847,970,1879,1904]" italics="true" pageId="4" pageNumber="65">H. vencesi</emphasis>
</taxonomicName>
and set this taxon apart from the other species assigned to this genus (see below) (Cumberlidge &amp; Sternberg 2002: 4859, fig. 9AQ).
</paragraph>
</subSubSection>
<caption id="DF5B6662F315FFC8FF30FA096C11FB72" httpUri="https://zenodo.org/record/177501/files/figure.png" pageId="5" pageNumber="66" targetBox="[469,1117,197,1126]" targetPageId="5">
<paragraph id="8B9B36EAF315FFC8FF30FA096C11FB72" blockId="5.[151,1436,1151,1209]" pageId="5" pageNumber="66">
<emphasis id="B950EAF8F315FFC8FF30FA09682FFB5C" bold="true" box="[153,352,1151,1175]" pageId="5" pageNumber="66">FIGURES 1617.</emphasis>
<taxonomicName id="4C244D69F315FFC8FEC1FAF66B25FB5C" box="[360,618,1152,1175]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphsua" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi" status="sp. nov.">
<emphasis id="B950EAF8F315FFC8FEC1FAF66B25FB5C" box="[360,618,1152,1175]" italics="true" pageId="5" pageNumber="66">Hydrothelphsua vencesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635783F315FFC8FDD8FA096BE1FB5C" box="[625,686,1151,1175]" pageId="5" pageNumber="66" rank="species">
<emphasis id="B950EAF8F315FFC8FDD8FA096BE1FB5C" bold="true" box="[625,686,1151,1175]" pageId="5" pageNumber="66">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 16, whole animal dorsal view; 17, detail of eyes and mouthparts, frontal view.
</paragraph>
</caption>
<subSubSection id="C33E6561F315FFCBFF6EFA9B6C2BFD59" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="description">
<paragraph id="8B9B36EAF315FFC8FF6EFA9B6C38F9D4" blockId="5.[151,1437,1261,2007]" pageId="5" pageNumber="66">
<taxonomicName id="4C244D69F315FFC8FF6EFA9B68ABFACD" box="[199,484,1261,1286]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FF6EFA9B68ABFACD" box="[199,484,1261,1286]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C244D69F315FFC8FCBAFA9B6D40FACD" box="[787,1039,1261,1286]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B950EAF8F315FFC8FCBAFA9B6D40FACD" box="[787,1039,1261,1286]" italics="true" pageId="5" pageNumber="66">H. madagascariensis</emphasis>
</taxonomicName>
because these two large species share the following characters. The frontal margin, front behind the margin, and the suborbital, subhepatic, and pterygostomial regions of the carapace sidewall are all distinctly granulated, the exorbital and epibranchial teeth are both large and pointed, and the anterolateral margin behind the epibranchial tooth is either heavily granulated or toothed; the suborbital margin, the ischium of pereiopod 1, and the inferior margins of merus of pereiopod 1 are also granulated.
<taxonomicName id="4C244D69F315FFC8FD38FBC36AE3FA05" box="[657,940,1461,1486]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FD38FBC36AE3FA05" box="[657,940,1461,1486]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D69F315FFC8FBE4FBC36C08FA05" box="[1101,1351,1461,1486]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B950EAF8F315FFC8FBE4FBC36C08FA05" box="[1101,1351,1461,1486]" italics="true" pageId="5" pageNumber="66">H. madagascariensis</emphasis>
</taxonomicName>
in that the G1 terminal article of
<taxonomicName id="4C244D69F315FFC8FE67FBAB6B06FA3D" box="[462,585,1501,1526]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FE67FBAB6B06FA3D" box="[462,585,1501,1526]" italics="true" pageId="5" pageNumber="66">H. vencesi</emphasis>
</taxonomicName>
is a long straight-sided evenly-tapering cone that lacks the pronounced raised mid-section seen in
<taxonomicName id="4C244D69F315FFC8FE66F8736B89F9D5" box="[463,710,1541,1566]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B950EAF8F315FFC8FE66F8736B89F9D5" box="[463,710,1541,1566]" italics="true" pageId="5" pageNumber="66">H. madagascariensis</emphasis>
</taxonomicName>
(
<figureCitation id="131F2A6FF315FFC8FD7EF8736A29F9D4" box="[727,870,1541,1567]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="5" pageNumber="66">Figs. 1015</figureCitation>
; Cumberlidge &amp; Sternberg 2002: figs. 9EI).
</paragraph>
<paragraph id="8B9B36EAF315FFC8FF6EF85B6DDDF84C" blockId="5.[151,1437,1261,2007]" pageId="5" pageNumber="66">
<taxonomicName id="4C244D69F315FFC8FF6EF85B68ACF98D" box="[199,483,1581,1606]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FF6EF85B68ACF98D" box="[199,483,1581,1606]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
shares the following characters with
<taxonomicName id="4C244D69F315FFC8FC02F85B6D32F98D" box="[939,1149,1581,1606]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B950EAF8F315FFC8FC02F85B6D32F98D" box="[939,1149,1581,1606]" italics="true" pageId="5" pageNumber="66">H. bombetokensis</emphasis>
</taxonomicName>
. The exorbital and epibranchial teeth are both large and pointed and the first carpal tooth on the carpus of pereiopod 1 is a long, slen- der spine.
<taxonomicName id="4C244D69F315FFC8FEB2F80B6B75F95D" box="[283,570,1661,1686]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FEB2F80B6B75F95D" box="[283,570,1661,1686]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C244D69F315FFC8FC3CF80B6D24F95D" box="[917,1131,1661,1686]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B950EAF8F315FFC8FC3CF80B6D24F95D" box="[917,1131,1661,1686]" italics="true" pageId="5" pageNumber="66">H. bombetokensis</emphasis>
</taxonomicName>
as follows. The inferior margins of the merus of pereiopod 1 of
<taxonomicName id="4C244D69F315FFC8FDDBF8D36A0AF975" box="[626,837,1701,1726]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B950EAF8F315FFC8FDDBF8D36A0AF975" box="[626,837,1701,1726]" italics="true" pageId="5" pageNumber="66">H. bombetokensis</emphasis>
</taxonomicName>
are granular (whereas these margins in
<taxonomicName id="4C244D69F315FFC8FA89F8D36CD3F975" box="[1312,1436,1701,1726]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FA89F8D36CD3F975" box="[1312,1436,1701,1726]" italics="true" pageId="5" pageNumber="66">H. vencesi</emphasis>
</taxonomicName>
have several large and pointed teeth); the anterior carapace of
<taxonomicName id="4C244D69F315FFC8FCC4F8BB6D72F92D" box="[877,1085,1741,1766]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B950EAF8F315FFC8FCC4F8BB6D72F92D" box="[877,1085,1741,1766]" italics="true" pageId="5" pageNumber="66">H. bombetokensis</emphasis>
</taxonomicName>
is smooth (whereas that of
<taxonomicName id="4C244D69F315FFC8FAD6F8BB69A2F8C5" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FAD6F8BB69A2F8C5" italics="true" pageId="5" pageNumber="66">H. vencesi</emphasis>
</taxonomicName>
is heavily granulated in the anterolateral corners and the frontal region); and the G1 terminal article of
<taxonomicName id="4C244D69F315FFC8FF3EF96B6827F8FD" box="[151,360,1821,1846]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B950EAF8F315FFC8FF3EF96B6827F8FD" box="[151,360,1821,1846]" italics="true" pageId="5" pageNumber="66">H. bombetokensis</emphasis>
</taxonomicName>
is slim and distinctly raised in the mid-section (the result of a higher medial fold) ending in an upcurved tip (Cumberlidge &amp; Sternberg 2002: figs. 9JM), whereas that of
<emphasis id="B950EAF8F315FFC8FBE3F9336DF4F895" box="[1098,1211,1861,1886]" italics="true" pageId="5" pageNumber="66">P. v e n c e s i</emphasis>
is a long straightsided evenly-tapering cone that lacks the pronounced raised mid-section (
<figureCitation id="131F2A6FF315FFC8FC5CF91B6DCBF84C" box="[1013,1156,1901,1927]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="5" pageNumber="66">Figs. 1015</figureCitation>
).
</paragraph>
<paragraph id="8B9B36EAF315FFCBFF6EF9E36DCFFEB1" blockId="5.[151,1437,1261,2007]" lastBlockId="6.[151,1436,152,658]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<taxonomicName id="4C244D69F315FFC8FF6EF9E368AEF865" box="[199,481,1941,1966]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FF6EF9E368AEF865" box="[199,481,1941,1966]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C244D69F315FFC8FD28F9E36BA8F865" box="[641,743,1941,1966]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F315FFC8FD28F9E36BA8F865" box="[641,743,1941,1966]" italics="true" pageId="5" pageNumber="66">H. agilis</emphasis>
</taxonomicName>
in that the frontal margin of both of these large species is noticeably indented in the middle.
<taxonomicName id="4C244D69F315FFC8FD82F9CB6A0CF81D" box="[555,835,1981,2006]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F315FFC8FD82F9CB6A0CF81D" box="[555,835,1981,2006]" italics="true" pageId="5" pageNumber="66">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C244D69F315FFC8FBD7F9CB6DADF81D" box="[1150,1250,1981,2006]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F315FFC8FBD7F9CB6DADF81D" box="[1150,1250,1981,2006]" italics="true" pageId="5" pageNumber="66">H. agilis</emphasis>
</taxonomicName>
as follows. The frontal margin is distinctly toothed in
<taxonomicName id="4C244D69F316FFCBFDFCFEEE6BF4FF7A" box="[597,699,152,177]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F316FFCBFDFCFEEE6BF4FF7A" box="[597,699,152,177]" italics="true" pageId="6" pageNumber="67">H. agilis</emphasis>
</taxonomicName>
but granular in
<taxonomicName id="4C244D69F316FFCBFCD1FEEE6ABDFF7A" box="[888,1010,152,177]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFCD1FEEE6ABDFF7A" box="[888,1010,152,177]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
; the inferior margin of the merus of pereiopod 1 of
<taxonomicName id="4C244D69F316FFCBFEE1FEB668E2FF12" box="[328,429,192,217]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F316FFCBFEE1FEB668E2FF12" box="[328,429,192,217]" italics="true" pageId="6" pageNumber="67">H. agilis</emphasis>
</taxonomicName>
has several large and pointed teeth, while these margins in
<taxonomicName id="4C244D69F316FFCBFBCBFEB66D93FF12" box="[1122,1244,192,217]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFBCBFEB66D93FF12" box="[1122,1244,192,217]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
are granular but not toothed; the suborbital margin and the p1 ischium of
<taxonomicName id="4C244D69F316FFCBFCE9FE9E6AE8FECA" box="[832,935,232,257]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F316FFCBFCE9FE9E6AE8FECA" box="[832,935,232,257]" italics="true" pageId="6" pageNumber="67">H. agilis</emphasis>
</taxonomicName>
are both distinctly toothed, whereas these structures are granular in
<taxonomicName id="4C244D69F316FFCBFE65FF666B06FEE2" box="[460,585,272,297]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFE65FF666B06FEE2" box="[460,585,272,297]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
; and the terminal article of G1 of
<taxonomicName id="4C244D69F316FFCBFC44FF666D1BFEE2" box="[1005,1108,272,297]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B950EAF8F316FFCBFC44FF666D1BFEE2" box="[1005,1108,272,297]" italics="true" pageId="6" pageNumber="67">H. agilis</emphasis>
</taxonomicName>
is slim and evenly proportioned along its entire length, ending in an upcurved tip (Cumberlidge &amp; Sternberg 2002: figs. 9AD), whereas that of
<emphasis id="B950EAF8F316FFCBFEF8FF16688BFEB2" box="[337,452,352,377]" italics="true" pageId="6" pageNumber="67">
P.
<taxonomicName id="4C244D69F316FFCBFEC7FF16688BFEB2" box="[366,452,352,377]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">vencesi</taxonomicName>
</emphasis>
is a long straight-sided evenly-tapering cone (
<figureCitation id="131F2A6FF316FFCBFC4AFF166D3DFEB1" box="[995,1138,352,378]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="6" pageNumber="67">Figs. 1015</figureCitation>
).
</paragraph>
<paragraph id="8B9B36EAF316FFCBFF6EFFFE6C2BFD59" blockId="6.[151,1436,152,658]" pageId="6" pageNumber="67">
<taxonomicName id="4C244D69F316FFCBFF6EFFFE68ADFE6A" box="[199,482,392,417]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFF6EFFFE68ADFE6A" box="[199,482,392,417]" italics="true" pageId="6" pageNumber="67">Hydrothelphusa vencesi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C244D69F316FFCBFC81FFFE6AE9FE6A" box="[808,934,392,417]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B950EAF8F316FFCBFC81FFFE6AE9FE6A" box="[808,934,392,417]" italics="true" pageId="6" pageNumber="67">H. goudoti</emphasis>
</taxonomicName>
as follows. The carapace of
<taxonomicName id="4C244D69F316FFCBFAA9FFFE6CCFFE6A" box="[1280,1408,392,417]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B950EAF8F316FFCBFAA9FFFE6CCFFE6A" box="[1280,1408,392,417]" italics="true" pageId="6" pageNumber="67">H. goudoti</emphasis>
</taxonomicName>
is wider (cw/fw 3.9) and more arched (ch/fw 1.6) than that of
<taxonomicName id="4C244D69F316FFCBFCCCFFC66AAEFE02" box="[869,993,432,457]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFCCCFFC66AAEFE02" box="[869,993,432,457]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
(cw/fw 3.2, ch/fw 1.2); the exorbital and epibranchial teeth of
<taxonomicName id="4C244D69F316FFCBFE6AFFAE6B0FFE3A" box="[451,576,472,497]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B950EAF8F316FFCBFE6AFFAE6B0FFE3A" box="[451,576,472,497]" italics="true" pageId="6" pageNumber="67">H. goudoti</emphasis>
</taxonomicName>
are both small and blunt, whereas these teeth in
<taxonomicName id="4C244D69F316FFCBFB28FFAE6DB3FE3A" box="[1153,1276,472,497]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFB28FFAE6DB3FE3A" box="[1153,1276,472,497]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
are large and pointed; the postfrontal crest of
<taxonomicName id="4C244D69F316FFCBFDBEFC766BD9FDD2" box="[535,662,512,537]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B950EAF8F316FFCBFDBEFC766BD9FDD2" box="[535,662,512,537]" italics="true" pageId="6" pageNumber="67">H. goudoti</emphasis>
</taxonomicName>
is incomplete (wherein epibranchial crests and postorbital crests are separate), whereas in
<taxonomicName id="4C244D69F316FFCBFE17FC5E6B77FD8A" box="[446,568,552,577]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="vencesi">
<emphasis id="B950EAF8F316FFCBFE17FC5E6B77FD8A" box="[446,568,552,577]" italics="true" pageId="6" pageNumber="67">H. vencesi</emphasis>
</taxonomicName>
the postfrontal crest is complete; and the terminal article of G1 of
<taxonomicName id="4C244D69F316FFCBFAEAFC5E698AFDA2" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B950EAF8F316FFCBFAEAFC5E698AFDA2" italics="true" pageId="6" pageNumber="67">H. goudoti</emphasis>
</taxonomicName>
is slim with a raised and rounded mesial lobe, ending in a broad straight tip (Cumberlidge &amp; Sternberg 2002: figs. 9NQ), whereas that of
<emphasis id="B950EAF8F316FFCBFD9FFC0E6BEAFD5A" box="[566,677,632,657]" italics="true" pageId="6" pageNumber="67">P. v e n c e s i</emphasis>
is a long straight-sided evenly-tapering cone (
<figureCitation id="131F2A6FF316FFCBFB61FC0E6C18FD59" box="[1224,1367,632,658]" captionStart="FIGURES 10 15" captionStartId="4.[151,269,1545,1569]" captionTargetBox="[191,1406,216,1504]" captionTargetId="figure@4.[164,1424,194,1521]" captionTargetPageId="4" captionText="FIGURES 10 15. Hydrothelphsua vencesi n. sp., adult male holotype (cw 52 mm, ZMA De. 205976) from Vevembe, Fianarantsoa Province, Madagascar. 10, right G 1, ventral view; 11, right G 1, dorsal view; 12, terminal article of right G 1, dorsal view; 13, detail of tip of terminal article of right G 1, dorsal view; 14, detail of tip of terminal article of right G 1, dorsal view; 15, right first gonopod turned to show detail of tip of terminal article. Scale = 7 mm (10, 11); 2 mm (12 15)." httpUri="https://zenodo.org/record/177500/files/figure.png" pageId="6" pageNumber="67">Figs. 1015</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>