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<document id="3646F83F11C409334005589481246CEF" ID-DOI="10.11646/zootaxa.3716.3.7" ID-GBIF-Dataset="c3d8f250-18be-4e39-b2fa-937f217c0021" ID-ISSN="1175-5326" ID-Zenodo-Dep="221814" ID-ZooBank="D8B9E93B-2EC9-4DF8-B2F7-40F39AB0A65B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460682344186" checkinUser="plazi" docAuthor="Geginat, Gernot" docDate="2013" docId="038D87C5FFC3F61D60D7FAC730DA3BD8" docLanguage="en" docName="zt03716p459.pdf" docOrigin="Zootaxa 3716 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechosiella scotti subsp. swartbergensis Geginat, 2013, subspec. nov." docType="treatment" docVersion="7" lastPageNumber="457" masterDocId="FFB4FFBDFFC8F60D6040FFFB31193A53" masterDocTitle="Taxonomic revision of the African genus Trechosiella Jeannel (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="459" masterPageNumber="441" pageNumber="452" updateTime="1698350580333" updateUser="plazi">
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<mods:title id="CA341D77B2DAEDA3660E627F27FBB05F">Taxonomic revision of the African genus Trechosiella Jeannel (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="CCA55C0BFA950111C4ABCB88D814EB97">Geginat, Gernot</mods:namePart>
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<treatment id="038D87C5FFC3F61D60D7FAC730DA3BD8" ID-DOI="http://doi.org/10.5281/zenodo.6164138" ID-GBIF-Taxon="119580465" ID-Zenodo-Dep="6164138" LSID="urn:lsid:plazi:treatment:038D87C5FFC3F61D60D7FAC730DA3BD8" httpUri="http://treatment.plazi.org/id/038D87C5FFC3F61D60D7FAC730DA3BD8" lastPageId="16" lastPageNumber="457" pageId="11" pageNumber="452">
<subSubSection id="C33E6558FFC3F60660D7FAC730223F24" pageId="11" pageNumber="452" type="nomenclature">
<paragraph id="8B9B36D3FFC3F60660D7FAC733C13F05" blockId="11.[151,728,1340,1399]" box="[151,728,1340,1366]" pageId="11" pageNumber="452">
<heading id="D0D381BFFFC3F60660D7FAC733C13F05" bold="true" box="[151,728,1340,1366]" fontSize="11" level="1" pageId="11" pageNumber="452" reason="1">
<emphasis id="B950EAC1FFC3F60660D7FAC733C13F05" bold="true" box="[151,728,1340,1366]" pageId="11" pageNumber="452">
<taxonomicName id="4C244D50FFC3F60660D7FAC733343F05" ID-CoL="5F8PL" box="[151,557,1340,1366]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="scotti" status="subsp. nov." subSpecies="swartbergensis">
<emphasis id="B950EAC1FFC3F60660D7FAC733343F05" bold="true" box="[151,557,1340,1366]" italics="true" pageId="11" pageNumber="452">Trechosiella scotti swartbergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC3F6066274FAC733C13F05" box="[564,728,1340,1366]" pageId="11" pageNumber="452" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B36D3FFC3F60660D7FAA530223F24" blockId="11.[151,728,1340,1399]" box="[151,315,1374,1399]" pageId="11" pageNumber="452">
<figureCitation id="131F2A56FFC3F60660D7FAA531FA3F24" box="[151,227,1374,1399]" captionStart="FIGURE 5" captionStartId="12.[151,250,1832,1854]" captionTargetBox="[163,1418,210,1793]" captionTargetId="figure@12.[151,1436,193,1810]" captionTargetPageId="12" captionText="FIGURE 5. Trechosiella scotti swartbergensis subspec. nov., HT ♂ from Swartberg, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221819/files/figure.png" pageId="11" pageNumber="452">Figs. 5</figureCitation>
AC, 9
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC3F60660D7FA5D357A3C54" pageId="11" pageNumber="452" type="materials_examined">
<paragraph id="8B9B36D3FFC3F60660D7FA5D357A3C54" blockId="11.[151,1436,1446,1981]" pageId="11" pageNumber="452">
<emphasis id="B950EAC1FFC3F60660D7FA5D32643FEC" bold="true" box="[151,893,1446,1471]" pageId="11" pageNumber="452">
<typeStatus id="549F8871FFC3F60660D7FA5D31CB3FEC" box="[151,210,1446,1471]" pageId="11" pageNumber="452">Type</typeStatus>
material. [
<collectingCountry id="F3337643FFC3F606611AFA5D33203FEC" box="[346,569,1446,1471]" name="South Africa" pageId="11" pageNumber="452">SOUTH AFRICA</collectingCountry>
, Western Cape Province]:
</emphasis>
HT ♂, labelled S.Africa, Cape, Swartb[er]g., Meiringspoort. cent[ral]., shorewashing,
<geoCoordinate id="EE105014FFC3F6066220FA3133B73FB0" box="[608,686,1482,1507]" direction="south" orientation="latitude" pageId="11" pageNumber="452" precision="555" value="-33.25">33.25S</geoCoordinate>
/
<geoCoordinate id="EE105014FFC3F60662F7FA3132133FB0" box="[695,778,1482,1507]" direction="east" orientation="longitude" pageId="11" pageNumber="452" precision="555" value="22.33">22.33E</geoCoordinate>
,
<date id="FF9A1013FFC3F6066354FA30329F3FB0" box="[788,902,1483,1507]" pageId="11" pageNumber="452" value="1993-01-11">1.11.1993</date>
, E-Y: 2925, leg. Endrödy-Younga' (TMSA). PT
<specimenCount id="9D22FD5AFFC3F60660D7FA1431D83C55" box="[151,193,1519,1543]" pageId="11" pageNumber="452" type="generic">1 ♂</specimenCount>
with the same locality data (GGC). The aedeagi of the HT and PT were examined.
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC3F6016087F9E934423D93" lastPageId="12" lastPageNumber="453" pageId="11" pageNumber="452" type="description">
<paragraph id="8B9B36D3FFC3F6066087F9E933FC3DEF" blockId="11.[151,1436,1446,1981]" pageId="11" pageNumber="452">
<emphasis id="B950EAC1FFC3F6066087F9E930403C78" bold="true" box="[199,345,1554,1579]" pageId="11" pageNumber="452">Description.</emphasis>
Body length
<quantity id="4CDC9B36FFC3F60661BAF9E833BD3C79" box="[506,676,1555,1579]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.27" metricValueMin="3.1" pageId="11" pageNumber="452" unit="mm" value="3.185" valueMax="3.27" valueMin="3.1">3.103.27 mm</quantity>
. Body and appendages light brown to yellow, legs lighter yellow (
<figureCitation id="131F2A56FFC3F60660DFF9CD31FC3C1C" box="[159,229,1590,1615]" captionStart="FIGURE 5" captionStartId="12.[151,250,1832,1854]" captionTargetBox="[163,1418,210,1793]" captionTargetId="figure@12.[151,1436,193,1810]" captionTargetPageId="12" captionText="FIGURE 5. Trechosiella scotti swartbergensis subspec. nov., HT ♂ from Swartberg, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221819/files/figure.png" pageId="11" pageNumber="452">Fig. 5</figureCitation>
A). Eyes small, not protuberant; temple as long as diameter of eye; temporal lobe strongly developed, smooth. Frontal furrows and antennal length as in
<taxonomicName id="4C244D50FFC3F60662AFF9A7328D3C20" box="[751,916,1628,1651]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="scotti">
<emphasis id="B950EAC1FFC3F60662AFF9A7328D3C20" box="[751,916,1628,1651]" italics="true" pageId="11" pageNumber="452">T. scotti scotti</emphasis>
</taxonomicName>
. Ligula not examined. Pronotum cordiform, weakly transverse (length
<quantity id="4CDC9B36FFC3F6066183F98533733CC5" box="[451,618,1662,1687]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.0" metricValueMin="6.8" pageId="11" pageNumber="452" unit="mm" value="0.69" valueMax="0.7" valueMin="0.68">0.680.70 mm</quantity>
, width
<quantity id="4CDC9B36FFC3F6066280F984327E3CC5" box="[704,871,1663,1687]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.299999999999999" metricValueMin="8.0" pageId="11" pageNumber="452" unit="mm" value="0.815" valueMax="0.83" valueMin="0.8">0.800.83 mm</quantity>
, average length/width ratio 0.85,
<emphasis id="B950EAC1FFC3F60664A7F98435EC3CC5" box="[1255,1269,1663,1686]" italics="true" pageId="11" pageNumber="452">n</emphasis>
= 2); pronotal base, angles and impressions as in
<taxonomicName id="4C244D50FFC3F606625CF95F33AE3CE8" box="[540,695,1700,1723]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="scotti">
<emphasis id="B950EAC1FFC3F606625CF95F33AE3CE8" box="[540,695,1700,1723]" italics="true" pageId="11" pageNumber="452">T. scotti scotti</emphasis>
</taxonomicName>
. Shape of elytra similar to
<taxonomicName id="4C244D50FFC3F60663A6F95F35983CE8" box="[998,1153,1700,1723]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="scotti">
<emphasis id="B950EAC1FFC3F60663A6F95F35983CE8" box="[998,1153,1700,1723]" italics="true" pageId="11" pageNumber="452">T. scotti scotti</emphasis>
</taxonomicName>
; elytral length
<quantity id="4CDC9B36FFC3F606656CF95831DC3C8D" metricMagnitude="-3" metricUnit="m" metricValue="1.745" metricValueMax="1.79" metricValueMin="1.7" pageId="11" pageNumber="452" unit="mm" value="1.745" valueMax="1.79" valueMin="1.7">1.701.79 mm</quantity>
, width
<quantity id="4CDC9B36FFC3F606615AF93C30D93C8D" box="[282,448,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.2" metricValueMin="1.17" pageId="11" pageNumber="452" unit="mm" value="1.185" valueMax="1.2" valueMin="1.17">1.171.20 mm</quantity>
, average length/width ratio 1.47,
<emphasis id="B950EAC1FFC3F606637DF93C32523C8D" box="[829,843,1735,1758]" italics="true" pageId="11" pageNumber="452">n</emphasis>
= 2. Scutellar and elytral striae as in
<taxonomicName id="4C244D50FFC3F60664B7F93C348A3C8D" box="[1271,1427,1735,1758]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="scotti">
<emphasis id="B950EAC1FFC3F60664B7F93C348A3C8D" box="[1271,1427,1735,1758]" italics="true" pageId="11" pageNumber="452">T. scotti scotti</emphasis>
</taxonomicName>
. Intervals on elytral disc convex, not flattened. Recurrent sutural stria fairly strongly impressed on apex, confluent with the converging 5th and 7th elytral striae. Setiferous punctures of third stria inserted at anterior 1/5 and at middle of 3rd stria, respectively. Microsculpture of elytra weakly transverse, length/width ratio of mesh approximately 1:1 to 1:2 (
<figureCitation id="131F2A56FFC3F60660A6F8A730313D26" box="[230,296,1884,1909]" captionStart="FIGURE 5" captionStartId="12.[151,250,1832,1854]" captionTargetBox="[163,1418,210,1793]" captionTargetId="figure@12.[151,1436,193,1810]" captionTargetPageId="12" captionText="FIGURE 5. Trechosiella scotti swartbergensis subspec. nov., HT ♂ from Swartberg, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221819/files/figure.png" pageId="11" pageNumber="452">Fig. 5</figureCitation>
C). Brachypterous. Aedeagus length (in dorsal view)
<quantity id="4CDC9B36FFC3F60663C0F8A732FD3D27" box="[896,996,1884,1909]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="452" unit="mm" value="0.64">0.64 mm</quantity>
(
<emphasis id="B950EAC1FFC3F60663B3F8A635183D27" box="[1011,1025,1885,1908]" italics="true" pageId="11" pageNumber="452">n</emphasis>
= 2) (
<figureCitation id="131F2A56FFC3F6066404F8A7359F3D26" box="[1092,1158,1884,1909]" captionStart="FIGURE 5" captionStartId="12.[151,250,1832,1854]" captionTargetBox="[163,1418,210,1793]" captionTargetId="figure@12.[151,1436,193,1810]" captionTargetPageId="12" captionText="FIGURE 5. Trechosiella scotti swartbergensis subspec. nov., HT ♂ from Swartberg, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221819/files/figure.png" pageId="11" pageNumber="452">Fig. 5</figureCitation>
B); distal half of median lobe bulbous, very weakly bent ventrad, almost straight, apex blunt; internal sac with loop-like structure; styles broadly rounded, each bearing 57 long apical setae.
</paragraph>
<caption id="DF5B665BFFC4F60160D7F8D334623D0F" httpUri="https://zenodo.org/record/221819/files/figure.png" pageId="12" pageNumber="453" targetBox="[163,1418,210,1793]" targetPageId="12">
<paragraph id="8B9B36D3FFC4F60160D7F8D334623D0F" blockId="12.[151,1436,1832,1884]" pageId="12" pageNumber="453">
<emphasis id="B950EAC1FFC4F60160D7F8D3300A3D6E" bold="true" box="[151,275,1832,1854]" pageId="12" pageNumber="453">FIGURE 5.</emphasis>
<taxonomicName id="4C244D50FFC4F601615AF8D3336A3D6D" box="[282,627,1832,1854]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="scotti" status="subsp. nov." subSpecies="swartbergensis">
<emphasis id="B950EAC1FFC4F601615AF8D3336A3D6D" box="[282,627,1832,1854]" italics="true" pageId="12" pageNumber="453">Trechosiella scotti swartbergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC4F601623BF8D3321F3D6D" box="[635,774,1832,1854]" pageId="12" pageNumber="453" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
, HT ♂ from Swartberg, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C).
</paragraph>
</caption>
<paragraph id="8B9B36D3FFC4F6016087F87F34423D93" blockId="12.[151,1436,1924,2021]" pageId="12" pageNumber="453">
<emphasis id="B950EAC1FFC4F6016087F87F30D73DCE" bold="true" box="[199,462,1924,1949]" pageId="12" pageNumber="453">Differential diagnosis.</emphasis>
Smaller and more lightly pigmented than
<taxonomicName id="4C244D50FFC4F60163EDF87E351C3DCF" box="[941,1029,1925,1948]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="453" phylum="Arthropoda" rank="species" sensu="stricto" species="scotti">
<emphasis id="B950EAC1FFC4F60163EDF87E351C3DCF" box="[941,1029,1925,1948]" italics="true" pageId="12" pageNumber="453">T. scotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC4F601644EF87E35583DCF" box="[1038,1089,1925,1948]" pageId="12" pageNumber="453" sensu="stricto">s.str.</taxonomicNameLabel>
and
<taxonomicName id="4C244D50FFC4F6016439F87E34453DCF" box="[1145,1372,1925,1948]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="macroptera">
<emphasis id="B950EAC1FFC4F6016439F87E34453DCF" box="[1145,1372,1925,1948]" italics="true" pageId="12" pageNumber="453">T. scotti macroptera</emphasis>
</taxonomicName>
, with smaller and less protuberant eyes, and with the distal part of the median lobe of the aedeagus almost straight.
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC4F6006087F83730223BDB" lastPageId="13" lastPageNumber="454" pageId="12" pageNumber="453" type="discussion">
<paragraph id="8B9B36D3FFC4F6006087F83732173AAB" blockId="12.[151,1436,1924,2021]" lastBlockId="13.[151,1436,151,248]" lastPageId="13" lastPageNumber="454" pageId="12" pageNumber="453">
<emphasis id="B950EAC1FFC4F6016087F83730223DB6" bold="true" box="[199,315,1996,2021]" pageId="12" pageNumber="453">Remarks.</emphasis>
Of this subspecies, only the two
<typeStatus id="549F8871FFC4F60162F3F83633FA3DB6" box="[691,739,1997,2021]" pageId="12" pageNumber="453">type</typeStatus>
specimens from the Swartberg are known.
<taxonomicName id="4C244D50FFC4F600648EF83730263AE3" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" lastPageId="13" lastPageNumber="454" order="Coleoptera" pageId="12" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="swartbergensis">
<emphasis id="B950EAC1FFC4F600648EF83730263AE3" italics="true" lastPageId="13" lastPageNumber="454" pageId="12" pageNumber="453">Trechosiella scotti swartbergensis</emphasis>
</taxonomicName>
strongly resembles
<taxonomicName id="4C244D50FFC5F600627CFF6233F33AE3" box="[572,746,153,176]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="scotti">
<emphasis id="B950EAC1FFC5F600627CFF6233F33AE3" box="[572,746,153,176]" italics="true" pageId="13" pageNumber="454">T. scotti scotti</emphasis>
</taxonomicName>
, but shows some adaptations to endogeic life, such as depigmentation and some reduction of the eyes. Like many specimens of the other two subspecies, the specimens of
<taxonomicName id="4C244D50FFC5F60060F5FF1A30A43AAB" box="[181,445,223,248]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="scotti" subSpecies="swartbergensis">
<emphasis id="B950EAC1FFC5F60060F5FF1A30A43AAB" box="[181,445,223,248]" italics="true" pageId="13" pageNumber="454">T. scotti swartbergensis</emphasis>
</taxonomicName>
were found by shorewashing.
</paragraph>
<paragraph id="8B9B36D3FFC5F60060D7FEB730223BDB" blockId="13.[151,589,332,392]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F60060D7FEB733543B36" bold="true" box="[151,589,332,357]" pageId="13" pageNumber="454">
<taxonomicName id="4C244D50FFC5F60060D7FEB730CD3B36" box="[151,468,332,357]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="oligophthalma" status="sp. nov.">
<emphasis id="B950EAC1FFC5F60060D7FEB730CD3B36" bold="true" box="[151,468,332,357]" italics="true" pageId="13" pageNumber="454">Trechosiella oligophthalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC5F600619BFEB733543B36" box="[475,589,332,357]" pageId="13" pageNumber="454" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="131F2A56FFC5F60060D7FE9431FA3BDB" box="[151,227,367,392]" captionStart="FIGURE 6" captionStartId="14.[151,250,1970,1992]" captionTargetBox="[152,1433,212,1969]" captionTargetId="figure@14.[151,1436,193,1970]" captionTargetPageId="14" captionText="FIGURE 6. Trechosiella oligophthalma spec. nov., HT ♂ from Black Mt., Lesotho. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221820/files/figure.png" pageId="13" pageNumber="454">Figs. 6</figureCitation>
AC, 9
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC5F60060D7FE4C353B3BA7" pageId="13" pageNumber="454" type="materials_examined">
<paragraph id="8B9B36D3FFC5F60060D7FE4C353B3BA7" blockId="13.[151,1436,439,1409]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F60060D7FE4C30EC3B83" bold="true" box="[151,501,439,464]" pageId="13" pageNumber="454">
<typeStatus id="549F8871FFC5F60060D7FE4C31CB3B83" box="[151,210,439,464]" pageId="13" pageNumber="454">Type</typeStatus>
material. [
<collectingCountry id="F3337643FFC5F6006117FE4C30F03B83" box="[343,489,439,464]" name="Lesotho" pageId="13" pageNumber="454">LESOTHO</collectingCountry>
]:
</emphasis>
HT ♂, labelled S. Africa,
<collectingCountry id="F3337643FFC5F6006375FE4C328B3B83" box="[821,914,439,464]" name="Lesotho" pageId="13" pageNumber="454">Lesotho</collectingCountry>
, Drakensb[er]g., Black M[oun]t[ain].,
<geoCoordinate id="EE105014FFC5F600650DFE4334853B83" box="[1357,1436,440,464]" direction="south" orientation="latitude" pageId="13" pageNumber="454" precision="555" value="-29.31">29.31S</geoCoordinate>
<geoCoordinate id="EE105014FFC5F60060D7FE2731F03BA7" box="[151,233,476,500]" direction="east" orientation="longitude" pageId="13" pageNumber="454" precision="555" value="29.12">29.12E</geoCoordinate>
,
<date id="FF9A1013FFC5F60060B5FE2730423BA6" box="[245,347,476,501]" pageId="13" pageNumber="454" value="1976-03-09">9.3.1976</date>
; E-Y: 1060, from under stones, leg. Endrödy-Younga' (TMSA).
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC5F6006087FE04350A3E9E" pageId="13" pageNumber="454" type="description">
<paragraph id="8B9B36D3FFC5F6006087FE0433323E33" blockId="13.[151,1436,439,1409]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F6006087FE043040384B" bold="true" box="[199,345,511,536]" pageId="13" pageNumber="454">Description.</emphasis>
Body shape and size as in
<taxonomicName id="4C244D50FFC5F60062C3FDFA33F5384B" box="[643,748,511,536]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="laetula">
<emphasis id="B950EAC1FFC5F60062C3FDFA33F5384B" box="[643,748,511,536]" italics="true" pageId="13" pageNumber="454">T. laetula</emphasis>
</taxonomicName>
; body length
<quantity id="4CDC9B36FFC5F60063C5FDFB32F03844" box="[901,1001,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="454" unit="mm" value="3.3">3.30 mm</quantity>
(
<emphasis id="B950EAC1FFC5F60063B8FDFA351F384B" box="[1016,1030,513,536]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 1). Color of body and appendages similar to that of
<taxonomicName id="4C244D50FFC5F6006122FDDE30D7386E" box="[354,462,548,573]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="laetula">
<emphasis id="B950EAC1FFC5F6006122FDDE30D7386E" box="[354,462,548,573]" italics="true" pageId="13" pageNumber="454">T. laetula</emphasis>
</taxonomicName>
, body yellowish brown, legs more yellowish (
<figureCitation id="131F2A56FFC5F60063A7FDDF3535386E" box="[999,1068,548,573]" captionStart="FIGURE 6" captionStartId="14.[151,250,1970,1992]" captionTargetBox="[152,1433,212,1969]" captionTargetId="figure@14.[151,1436,193,1970]" captionTargetPageId="14" captionText="FIGURE 6. Trechosiella oligophthalma spec. nov., HT ♂ from Black Mt., Lesotho. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221820/files/figure.png" pageId="13" pageNumber="454">Fig. 6</figureCitation>
A). Eyes small, not protruding; temple twice as long as diameter of eye; temporal lobe well developed, smooth and without any wrinkles. Frontal furrows converging anteriad in basal half of frons, diverging thereafter towards anterior margin of clypeus, distinctly continued on clypeus towards anterior margin. Antenna reaching to between anterior and posterior setae on elytral disc. Anterior margin of labrum concave, almost triangular. Ligula not examined. Pronotum weakly cordiform, transverse (length
<quantity id="4CDC9B36FFC5F60061A2FD23335E38BC" box="[482,583,728,752]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="454" unit="mm" value="0.7">0.70 mm</quantity>
, width
<quantity id="4CDC9B36FFC5F60062D8FD2333E738BC" box="[664,766,728,752]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="454" unit="mm" value="0.9">0.90 mm</quantity>
, length/width ratio 0.78,
<emphasis id="B950EAC1FFC5F6006455FD22353A38A3" box="[1045,1059,729,752]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 1), maximum width at the level of anterior 1/3, posterior angles rectangular, not protruding. Pronotal base laterally slightly slanted anteriad, basal impressions shallow, not divided into separate central and lateral impressions. Elytral length
<quantity id="4CDC9B36FFC5F60064F0FCDB340E3964" box="[1200,1303,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="454" unit="mm" value="1.9">1.90 mm</quantity>
, width
<quantity id="4CDC9B36FFC5F600652BFCDB31DC390F" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="13" pageNumber="454" unit="mm" value="1.31">1.31 mm</quantity>
, length/width ratio 1.58 (
<emphasis id="B950EAC1FFC5F60061A6FCBE30ED390F" box="[486,500,837,860]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 1), humeri rounded. Scutellar stria shallow, ending well before level of anterior setiferous puncture on elytral disc, continued by 1st elytral stria. Striae 24 effaced, yet still discernible, striae 57 totally effaced, indiscernible, 8th stria strongly impressed; recurrent sutural stria totally effaced on apex. Setiferous punctures of third stria inserted at anterior 1/4 and behind middle of 3rd stria, respectively. Microsculpture of elytra very fine, transverse, with length/width ratio of mesh approximately 1:5 to 1:10 (
<figureCitation id="131F2A56FFC5F600647BFC27359939A6" box="[1083,1152,988,1013]" captionStart="FIGURE 6" captionStartId="14.[151,250,1970,1992]" captionTargetBox="[152,1433,212,1969]" captionTargetId="figure@14.[151,1436,193,1970]" captionTargetPageId="14" captionText="FIGURE 6. Trechosiella oligophthalma spec. nov., HT ♂ from Black Mt., Lesotho. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221820/files/figure.png" pageId="13" pageNumber="454">Fig. 6</figureCitation>
C). Apterous. Length of aedeagus in dorsal view
<quantity id="4CDC9B36FFC5F60061F1FBFB330E3E4B" box="[433,535,1024,1049]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="454" unit="mm" value="0.75">0.75 mm</quantity>
(
<emphasis id="B950EAC1FFC5F6006268FBFA332F3E4B" box="[552,566,1025,1048]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 1) (
<figureCitation id="131F2A56FFC5F600623FFBFB33DA3E4A" box="[639,707,1024,1049]" captionStart="FIGURE 6" captionStartId="14.[151,250,1970,1992]" captionTargetBox="[152,1433,212,1969]" captionTargetId="figure@14.[151,1436,193,1970]" captionTargetPageId="14" captionText="FIGURE 6. Trechosiella oligophthalma spec. nov., HT ♂ from Black Mt., Lesotho. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221820/files/figure.png" pageId="13" pageNumber="454">Fig. 6</figureCitation>
B). Median lobe long and straight, apex elongate, apex slightly bent ventrad, terminating in rounded tip; internal sac with triangular sclerotized internal piece; styles slightly tapered, each bearing 5 apical setae.
</paragraph>
<paragraph id="8B9B36D3FFC5F6006087FB97350A3E9E" blockId="13.[151,1436,439,1409]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F6006087FB9730CB3ED6" bold="true" box="[199,466,1132,1157]" pageId="13" pageNumber="454">Differential diagnosis.</emphasis>
<taxonomicName id="4C244D50FFC5F600619DFB97320F3ED6" box="[477,790,1132,1157]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="oligophthalma">
<emphasis id="B950EAC1FFC5F600619DFB97320F3ED6" box="[477,790,1132,1157]" italics="true" pageId="13" pageNumber="454">Trechosiella oligophthalma</emphasis>
</taxonomicName>
differs from all the other species of this genus by the strong reduction of its eyes, its depigmentation, the effaced elytral striae and the morphology of the aedeagus, having a long, straight median lobe and a single, triangular endophallic sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC5F6006087FB2330723C42" pageId="13" pageNumber="454" type="discussion">
<paragraph id="8B9B36D3FFC5F6006087FB2335FD3FD2" blockId="13.[151,1436,439,1409]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F6006087FB2330223EA2" bold="true" box="[199,315,1240,1265]" pageId="13" pageNumber="454">Remarks.</emphasis>
An apterous species known from a single specimen discovered under a stone on Black Mountain in the Drakensberg Mountain Range.
<taxonomicName id="4C244D50FFC5F6006268FB0732473F46" box="[552,862,1276,1301]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="oligophthalma">
<emphasis id="B950EAC1FFC5F6006268FB0732473F46" box="[552,862,1276,1301]" italics="true" pageId="13" pageNumber="454">Trechosiella oligophthalma</emphasis>
</taxonomicName>
shows advanced reduction of the eyes and a high degree of depigmentation, but the body and appendages have shapes similar to those in
<taxonomicName id="4C244D50FFC5F600643FFADB34113F6A" box="[1151,1288,1312,1337]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAC1FFC5F600643FFADB34113F6A" box="[1151,1288,1312,1337]" italics="true" pageId="13" pageNumber="454">Trechosiella</emphasis>
</taxonomicName>
species with fully developed eyes. Judging by the morphology of the aedeagus and the elytral microsculpture,
<taxonomicName id="4C244D50FFC5F6006495FABE34823F0E" box="[1237,1435,1348,1373]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="oligophthtalma">
<emphasis id="B950EAC1FFC5F6006495FABE34823F0E" box="[1237,1435,1348,1373]" italics="true" pageId="13" pageNumber="454">T. oligophthtalma</emphasis>
</taxonomicName>
is probably more closely related to
<taxonomicName id="4C244D50FFC5F6006262FA9233923FD2" box="[546,651,1384,1409]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="laetula">
<emphasis id="B950EAC1FFC5F6006262FA9233923FD2" box="[546,651,1384,1409]" italics="true" pageId="13" pageNumber="454">T. laetula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C244D50FFC5F6006282FA92322E3FD2" box="[706,823,1384,1409]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="katicola">
<emphasis id="B950EAC1FFC5F6006282FA92322E3FD2" box="[706,823,1384,1409]" italics="true" pageId="13" pageNumber="454">T. katicola</emphasis>
</taxonomicName>
than to the other species of the genus.
</paragraph>
<paragraph id="8B9B36D3FFC5F60060D7FA2F30723C42" blockId="13.[151,620,1492,1553]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F60060D7FA2F33753FBE" bold="true" box="[151,620,1492,1517]" pageId="13" pageNumber="454">
<taxonomicName id="4C244D50FFC5F60060D7FA2F30EA3FBE" box="[151,499,1492,1517]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="endroedyyoungai" status="sp. nov.">
<emphasis id="B950EAC1FFC5F60060D7FA2F30EA3FBE" bold="true" box="[151,499,1492,1517]" italics="true" pageId="13" pageNumber="454">Trechosiella endroedyyoungai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC5F60061BAFA2F33753FBE" box="[506,620,1492,1517]" pageId="13" pageNumber="454" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="131F2A56FFC5F60060D7FA0331C53C43" box="[151,220,1528,1553]" captionStart="FIGURE 7" captionStartId="15.[151,250,1575,1597]" captionTargetBox="[153,1412,205,1557]" captionTargetId="figure@15.[151,1436,193,1568]" captionTargetPageId="15" captionText="FIGURE 7. Trechosiella endroedyyoungai spec. nov., HT ♂ from Vanrhyn's Pass, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221821/files/figure.png" pageId="13" pageNumber="454">Figs 7</figureCitation>
AC, 8 A, 9
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC5F60060D7F9BB35C53CF2" pageId="13" pageNumber="454" type="materials_examined">
<paragraph id="8B9B36D3FFC5F60060D7F9BB35C53CF2" blockId="13.[151,1436,1600,2024]" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F60060D7F9BB328E3C0A" bold="true" box="[151,919,1600,1625]" pageId="13" pageNumber="454">
<typeStatus id="549F8871FFC5F60060D7F9BB31CB3C0A" box="[151,210,1600,1625]" pageId="13" pageNumber="454">Type</typeStatus>
material. [
<collectingCountry id="F3337643FFC5F6006123F9BB335F3C0A" box="[355,582,1600,1625]" name="South Africa" pageId="13" pageNumber="454">SOUTH AFRICA</collectingCountry>
, Western Cape Province]:
</emphasis>
HT ♂, labelled R.S.Africa, S. W. Cape, Vanrhyns Pass, from road gutter, 3123S/1902E,
<date id="FF9A1013FFC5F600628BF99E32243C2E" box="[715,829,1637,1661]" pageId="13" pageNumber="454" value="1983-04-11">4.11.1983</date>
, E-Y: 2048, leg. Endrödy-Younga' (TMSA). PTs:
<specimenCount id="9D22FD5AFFC5F600652BF99E348E3C2F" box="[1387,1431,1637,1661]" pageId="13" pageNumber="454" type="generic">2 ♂</specimenCount>
,
<specimenCount id="9D22FD5AFFC5F60060D7F97231D83CF3" box="[151,193,1673,1696]" pageId="13" pageNumber="454" type="generic">7 ♂</specimenCount>
with the same locality data (TMSA, GGC). The aedeagi of the HT and 2 PTs were examined.
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFC5F61D6087F95735E53A84" lastPageId="16" lastPageNumber="457" pageId="13" pageNumber="454" type="description">
<paragraph id="8B9B36D3FFC5F6026087F957356A3DF4" blockId="13.[151,1436,1600,2024]" lastBlockId="15.[151,1436,1672,2035]" lastPageId="15" lastPageNumber="456" pageId="13" pageNumber="454">
<emphasis id="B950EAC1FFC5F6006087F95730403C96" bold="true" box="[199,345,1708,1733]" pageId="13" pageNumber="454">Description.</emphasis>
Size and shape similar to
<taxonomicName id="4C244D50FFC5F60062CFF95632373C96" box="[655,814,1708,1733]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B950EAC1FFC5F60062CFF95632373C96" box="[655,814,1708,1733]" italics="true" pageId="13" pageNumber="454">T. basilewskyi</emphasis>
</taxonomicName>
, average body length
<quantity id="4CDC9B36FFC5F6006472F95635813C97" box="[1074,1176,1709,1733]" metricMagnitude="-3" metricUnit="m" metricValue="4.84" pageId="13" pageNumber="454" unit="mm" value="4.84">4.84 mm</quantity>
(
<emphasis id="B950EAC1FFC5F60064EBF95635A03C97" box="[1195,1209,1709,1732]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 9). Body rufousbrown, legs lighter brown to yellow (
<figureCitation id="131F2A56FFC5F600620BF92B33893CBB" box="[587,656,1744,1769]" captionStart="FIGURE 7" captionStartId="15.[151,250,1575,1597]" captionTargetBox="[153,1412,205,1557]" captionTargetId="figure@15.[151,1436,193,1568]" captionTargetPageId="15" captionText="FIGURE 7. Trechosiella endroedyyoungai spec. nov., HT ♂ from Vanrhyn's Pass, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221821/files/figure.png" pageId="13" pageNumber="454">Fig. 7</figureCitation>
A). Eyes not protruding, temples half as long as diameter of eyes; temporal lobe well developed, smooth. Frontal furrows as in
<taxonomicName id="4C244D50FFC5F6006311F90E32A53D5E" box="[849,956,1780,1805]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="454" phylum="Arthropoda" rank="species" species="laetula">
<emphasis id="B950EAC1FFC5F6006311F90E32A53D5E" box="[849,956,1780,1805]" italics="true" pageId="13" pageNumber="454">T. laetula</emphasis>
</taxonomicName>
. Antenna reaching anterior seta on elytral disc. Anterior margin of labrum very weakly concave. Ligula rectangular, apex flat, not rounded (
<figureCitation id="131F2A56FFC5F6006563F8E334733D62" box="[1315,1386,1816,1841]" captionStart="FIGURE 8" captionStartId="16.[151,250,1964,1986]" captionTargetBox="[195,1411,487,1934]" captionTargetId="figure@16.[151,1436,450,1942]" captionTargetPageId="16" captionText="FIGURE 8. Ligulae and labial palps (dorsal views) of T. endroedyyoungai spec. nov. (A), T. laetula (B), T. scotti s. str. (C), and T. katicola (D)." httpUri="https://zenodo.org/record/221822/files/figure.png" pageId="13" pageNumber="454">Fig. 8</figureCitation>
A); anterior margin bearing two long central setae and 2 shorter lateral setae on each side, 3rd lateral seta absent, possible vestigial insertion point present at middle of lateral margin of ligula. Pronotum weakly cordiform, transverse (average length
<quantity id="4CDC9B36FFC5F600618AF87C332B3DCC" box="[458,562,1927,1951]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="454" unit="mm" value="0.9">0.90 mm</quantity>
, average width
<quantity id="4CDC9B36FFC5F60062A9F87C32483DCC" box="[745,849,1927,1951]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="13" pageNumber="454" unit="mm" value="1.31">1.31 mm</quantity>
, average length/width ratio 0.69,
<emphasis id="B950EAC1FFC5F6006493F87335F83DCC" box="[1235,1249,1928,1951]" italics="true" pageId="13" pageNumber="454">n</emphasis>
= 9), maximum width at level of anterior 1/3; posterior angles rectangular, not protruding; pronotal base straight, basal impressions shallow, not divided into separate central and lateral impressions. Average elytral length
<quantity id="4CDC9B36FFC5F60064C6F83435F53DB4" box="[1158,1260,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.87" pageId="13" pageNumber="454" unit="mm" value="2.87">2.87 mm</quantity>
, average width
<quantity id="4CDC9B36FFC7F60260D7F972301D3CF3" box="[151,260,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="15" pageNumber="456" unit="mm" value="1.87">1.87 mm</quantity>
, average length/width ratio 1.53 (
<emphasis id="B950EAC1FFC7F60262E3F97133A83CF2" box="[675,689,1674,1697]" italics="true" pageId="15" pageNumber="456">n</emphasis>
= 9), humeri well developed, rounded. Scutellar stria deeply impressed, ending before level of anterior setiferous puncture on elytral disc. Elytral striae 18 well discernible; 8th stria strongly impressed, 3rd and 4th striae confluent in the anterior 1/4 of elytral disc, anterior of anterior setiferous puncture; recurrent sutural stria weak on apex, confluent with the converging 5th, 6th and 7th striae. A single (sometimes duplicated) setiferous puncture inserted at anterior 1/4 of 3rd stria. Microsculpture of elytra coarse, length/width ratio of mesh approximately 1:2 to 1:3 (
<figureCitation id="131F2A56FFC7F60262B4F8BC32213D0C" box="[756,824,1863,1888]" captionStart="FIGURE 7" captionStartId="15.[151,250,1575,1597]" captionTargetBox="[153,1412,205,1557]" captionTargetId="figure@15.[151,1436,193,1568]" captionTargetPageId="15" captionText="FIGURE 7. Trechosiella endroedyyoungai spec. nov., HT ♂ from Vanrhyn's Pass, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221821/files/figure.png" pageId="15" pageNumber="456">Fig. 7</figureCitation>
C). Brachypterous. Aedeagus with average length (in dorsal view)
<quantity id="4CDC9B36FFC7F602616CF89730883DD7" box="[300,401,1900,1925]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="456" unit="mm" value="0.75">0.75 mm</quantity>
(
<emphasis id="B950EAC1FFC7F60261E3F89630A83DD7" box="[419,433,1901,1924]" italics="true" pageId="15" pageNumber="456">n</emphasis>
= 3) (
<figureCitation id="131F2A56FFC7F60261B9F89733243DD7" box="[505,573,1900,1925]" captionStart="FIGURE 7" captionStartId="15.[151,250,1575,1597]" captionTargetBox="[153,1412,205,1557]" captionTargetId="figure@15.[151,1436,193,1568]" captionTargetPageId="15" captionText="FIGURE 7. Trechosiella endroedyyoungai spec. nov., HT ♂ from Vanrhyn's Pass, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C)." httpUri="https://zenodo.org/record/221821/files/figure.png" pageId="15" pageNumber="456">Fig. 7</figureCitation>
B). Median lobe slightly curved ventrad, apex tapered; internal sac without prominent sclerotized internal armature; styles tapered, each bearing 2 long apical setae.
</paragraph>
<caption id="DF5B665BFFC6F60360D7F84935833DB5" httpUri="https://zenodo.org/record/221820/files/figure.png" pageId="14" pageNumber="455" targetBox="[152,1433,212,1969]" targetPageId="14">
<paragraph id="8B9B36D3FFC6F60360D7F84935833DB5" blockId="14.[151,1435,1970,2022]" pageId="14" pageNumber="455">
<emphasis id="B950EAC1FFC6F60360D7F849300C3D94" bold="true" box="[151,277,1970,1992]" pageId="14" pageNumber="455">FIGURE 6.</emphasis>
<taxonomicName id="4C244D50FFC6F603615EF849332E3D9B" box="[286,567,1970,1992]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="455" phylum="Arthropoda" rank="species" species="oligophthalma" status="sp. nov.">
<emphasis id="B950EAC1FFC6F603615EF849332E3D9B" box="[286,567,1970,1992]" italics="true" pageId="14" pageNumber="455">Trechosiella oligophthalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC6F6036200F84833B33D9B" box="[576,682,1971,1992]" pageId="14" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
, HT ♂ from Black Mt., Lesotho. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C).
</paragraph>
</caption>
<caption id="DF5B665BFFC7F60260D7F9DC34623C0F" httpUri="https://zenodo.org/record/221821/files/figure.png" pageId="15" pageNumber="456" targetBox="[153,1412,205,1557]" targetPageId="15">
<paragraph id="8B9B36D3FFC7F60260D7F9DC34623C0F" blockId="15.[151,1436,1575,1628]" pageId="15" pageNumber="456">
<emphasis id="B950EAC1FFC7F60260D7F9DC300C3C6F" bold="true" box="[151,277,1575,1597]" pageId="15" pageNumber="456">FIGURE 7.</emphasis>
<taxonomicName id="4C244D50FFC7F602615EF9DC334B3C6E" box="[286,594,1575,1597]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="456" phylum="Arthropoda" rank="species" species="endroedyyoungai" status="sp. nov.">
<emphasis id="B950EAC1FFC7F602615EF9DC334B3C6E" box="[286,594,1575,1597]" italics="true" pageId="15" pageNumber="456">Trechosiella endroedyyoungai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357BAFFC7F602621AF9D333DD3C6E" box="[602,708,1576,1597]" pageId="15" pageNumber="456" rank="species">spec. nov.</taxonomicNameLabel>
, HT ♂ from Vanrhyn's Pass, Western Cape Province. General habitus (A), aedeagus (lateral view) (B), microsculpture of elytra between the anterior and posterior discal setiferous punctures (C).
</paragraph>
</caption>
<paragraph id="8B9B36D3FFC7F61D6087F84F35E53A84" blockId="15.[151,1436,1672,2035]" lastBlockId="16.[151,1436,151,395]" lastPageId="16" lastPageNumber="457" pageId="15" pageNumber="456">
<emphasis id="B950EAC1FFC7F6026087F84F30D63D9E" bold="true" box="[199,463,1972,1997]" pageId="15" pageNumber="456">Differential diagnosis.</emphasis>
<taxonomicName id="4C244D50FFC7F6026197F84E33AC3D9E" box="[471,693,1972,1997]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="456" phylum="Arthropoda" rank="species" species="endroedyyoungai">
<emphasis id="B950EAC1FFC7F6026197F84E33AC3D9E" box="[471,693,1972,1997]" italics="true" pageId="15" pageNumber="456">T. endroedyyoungai</emphasis>
</taxonomicName>
is different from all other
<taxonomicName id="4C244D50FFC7F60263A6F84F35693D9E" box="[998,1136,1972,1997]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAC1FFC7F60263A6F84F35693D9E" box="[998,1136,1972,1997]" italics="true" pageId="15" pageNumber="456">Trechosiella</emphasis>
</taxonomicName>
species by the striation of the elytral disc, characterized by the confluence of the 3rd and 4th striae in the anterior quarter of the disc; and by the absence of the posterior setiferous puncture of the 3rd stria. The species is also characterized by an almost rectangular ligula and by the tapered styles of the aedeagus, each bearing only two long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C33E6558FFD8F61D6087FF1930DA3BD8" pageId="16" pageNumber="457" type="discussion">
<paragraph id="8B9B36D3FFD8F61D6087FF1930DA3BD8" blockId="16.[151,1436,151,395]" pageId="16" pageNumber="457">
<emphasis id="B950EAC1FFD8F61D6087FF1930223AA8" bold="true" box="[199,315,226,251]" pageId="16" pageNumber="457">Remarks.</emphasis>
<taxonomicName id="4C244D50FFD8F61D6103FF1F33393AA8" box="[323,544,226,251]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="457" phylum="Arthropoda" rank="species" species="endroedyyoungai">
<emphasis id="B950EAC1FFD8F61D6103FF1F33393AA8" box="[323,544,226,251]" italics="true" pageId="16" pageNumber="457">T. endroedyyoungai</emphasis>
</taxonomicName>
is a brachypterous species known from only the
<typeStatus id="549F8871FFD8F61D640DFF1835643AA8" box="[1101,1149,227,251]" pageId="16" pageNumber="457">type</typeStatus>
series, discovered by the late Dr. Endrödy-Younga in Vanrhyns Pass on the border of the Western and Northern Cape Provinces. The presence of a flightless
<taxonomicName id="4C244D50FFD8F61D61DEFED133313B10" box="[414,552,298,323]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAC1FFD8F61D61DEFED133313B10" box="[414,552,298,323]" italics="true" pageId="16" pageNumber="457">Trechosiella</emphasis>
</taxonomicName>
in this semi-arid area is unexpected as all other known
<taxonomicName id="4C244D50FFD8F61D64D2FED134023B10" box="[1170,1307,298,323]" class="Insecta" family="Carabidae" genus="Trechosiella" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAC1FFD8F61D64D2FED134023B10" box="[1170,1307,298,323]" italics="true" pageId="16" pageNumber="457">Trechosiella</emphasis>
</taxonomicName>
species are humicolous or ripicolous. Considering the unique morphologies of the ligula and the aedeagus, this species seems quite isolated in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>