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<document id="C959522C5E403E07B6F40F5076749385" ID-DOI="10.1590/S0031-10492009004400001" ID-ISSN="1807-0205" ID-Zenodo-Dep="13308299" IM.bibliography_requiresApprovalFor="GgImagineBatch" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1720050473348" checkinUser="felipe" docAuthor="Clarke 1, Robin O. S." docDate="2009" docId="0386F37AFFBC4532FD0B15D3FDF5FBD7" docLanguage="en" docName="PapAvZool.49.44.577-591.pdf.imf" docOrigin="Papéis Avulsos de Zoologia 49 (44)" docStyle="DocumentStyle:342E0B2A517D47F211C16A9F07059896.3:PapAvZool.2002-2013.journal_article.type2" docStyleId="342E0B2A517D47F211C16A9F07059896" docStyleName="PapAvZool.2002-2013.journal_article.type2" docStyleVersion="3" docTitle="Isthmiade ichneumoniformis Bates 1870" docType="treatment" docVersion="3" lastPageNumber="586" masterDocId="FFBF8B02FFBB453BFFED1564FFAFFFD8" masterDocTitle="Bolivian Rhinotragini II: ISTHMIADE Thomson, 1864 (Coleoptera, Cerambycidae), with two new species Robin O. S. Clarke" masterLastPageNumber="591" masterPageNumber="577" pageNumber="584" updateTime="1723494001863" updateUser="ExternalLinkService">
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<mods:title id="7BED4942CD06F98E6207539A53C679CB">Bolivian Rhinotragini II: ISTHMIADE Thomson, 1864 (Coleoptera, Cerambycidae), with two new species Robin O. S. Clarke</mods:title>
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<mods:namePart id="2E2ADBF938E3DD6B65653869708AF216">Clarke 1, Robin O. S.</mods:namePart>
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<mods:typeOfResource id="2D26FBBF7CFDE5D6C22B9E1EB2277D18">text</mods:typeOfResource>
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<mods:title id="722610966CDC9321754F54C3AF1048F2">Papéis Avulsos de Zoologia</mods:title>
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<mods:part id="5720609134069180155934FA43B4CA95">
<mods:date id="3A6137B9AA0FD38F7D4F5CEA82563FE6">2009</mods:date>
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<mods:number id="49B408B8B68F11395A7FC6DE621EA996">2009-12-31</mods:number>
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<mods:number id="18DDB0B5A4AE21CAD7518B7875AE7EEB">49</mods:number>
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<mods:number id="A64E48F1CABFFC1DF6530B37F7FD4D24">44</mods:number>
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<mods:identifier id="AE101AF732338BA388A8B1F67546E328" type="DOI">10.1590/S0031-10492009004400001</mods:identifier>
<mods:identifier id="810E647909C6E2F9EBB9750E5EE009B9" type="ISSN">1807-0205</mods:identifier>
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<treatment id="0386F37AFFBC4532FD0B15D3FDF5FBD7" ID-DOI="http://doi.org/10.5281/zenodo.13308307" ID-Zenodo-Dep="13308307" LSID="urn:lsid:plazi:treatment:0386F37AFFBC4532FD0B15D3FDF5FBD7" httpUri="http://treatment.plazi.org/id/0386F37AFFBC4532FD0B15D3FDF5FBD7" lastPageId="9" lastPageNumber="586" pageId="7" pageNumber="584">
<subSubSection id="C33511E7FFBC453CFD0B15D3FB20FF08" box="[742,1167,183,208]" pageId="7" pageNumber="584" type="nomenclature">
<paragraph id="8B90426CFFBC453CFD0B15D3FB20FF08" blockId="7.[742,1167,183,240]" box="[742,1167,183,208]" pageId="7" pageNumber="584">
<heading id="D0D8F500FFBC453CFD0B15D3FB20FF08" bold="true" box="[742,1167,183,208]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="584" reason="2">
<taxonomicName id="4C2F39EFFFBC453CFD0B15D3FB20FF08" ID-CoL="3QD6X" authority="Bates, 1870" authorityName="Bates" authorityYear="1870" box="[742,1167,183,208]" class="Insecta" family="Cerambycidae" genus="Isthmiade" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="ichneumoniformis">
<emphasis id="B95B9E7EFFBC453CFD0B15D3FB20FF08" bold="true" box="[742,1167,183,208]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD0B15D3FBA2FF17" bold="true" box="[742,1037,183,207]" italics="true" pageId="7" pageNumber="584">Isthmiade ichneumoniformis</emphasis>
<bibRefCitation id="EFBE3F9DFFBC453CFBFE15DCFB20FF08" author="Bates, H. W." box="[1043,1167,184,208]" pageId="7" pageNumber="584" pagination="391 - 444" refId="ref12254" refString="Bates, H. W. 1870. Contributions to an insect fauna of the Amazon Valley. Transactions of the Entomological Society of London, 391 - 444." type="book chapter" year="1870">Bates, 1870</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33511E7FFBC453CFC9C15BCFBABFF28" box="[881,1028,216,240]" pageId="7" pageNumber="584" type="description">
<paragraph id="8B90426CFFBC453CFC9C15BCFBABFF28" blockId="7.[742,1167,183,240]" box="[881,1028,216,240]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFC9C15BCFBABFF28" bold="true" box="[881,1028,216,240]" pageId="7" pageNumber="584">
(
<figureCitation id="13145EE9FFBC453CFC9515BCFC53FF28" box="[888,1020,216,240]" captionStart="FIGURAS 1‑5" captionStartId="3.[95,186,1800,1819]" captionTargetBox="[94,1224,175,1757]" captionTargetId="figure-3@3.[94,1228,170,1764]" captionTargetPageId="3" captionText="FIGURAS 1‑5: Bolivian species of Isthmiade Thomson, 1864. 1. Isthmiade laevicollis Tippmann, 1953, 1A female 1B male; 2. Isthmiade zamalloae sp. nov., 2A male holotype, 2B female; 3. Isthmiade ichneumoniformis Bates, 1870, 3A male, 3B female; 4. Isthmiade martinsi sp. nov., 4A male holotype; 5. Isthmiade planifrons Zajciw, 1972, 5A male, 5B female." figureDoi="http://doi.org/10.5281/zenodo.13308301" httpUri="https://zenodo.org/record/13308301/files/figure.png" pageId="7" pageNumber="584">Figs. 3A, 3B</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33511E7FFBC453CFD45147CFBDFFE68" pageId="7" pageNumber="584" type="reference_group">
<paragraph id="8B90426CFFBC453CFD45147CFBDFFE68" blockId="7.[680,1229,280,432]" pageId="7" pageNumber="584">
<treatmentCitationGroup id="AB3F6542FFBC453CFD45147CFBDFFE68" pageId="7" pageNumber="584">
<treatmentCitation id="0A8E647DFFBC453CFD45147CFB66FEE8" author="Bates, H. W." box="[680,1225,280,304]" page="326" pageId="7" pageNumber="584" year="1870">
<taxonomicName id="4C2F39EFFFBC453CFD45147CFB66FEE8" ID-CoL="3QD6X" authority="Bates, 1870: 326" authorityName="Bates" authorityPageNumber="326" authorityYear="1870" box="[680,1225,280,304]" class="Insecta" family="Cerambycidae" genus="Isthmiade" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="ichneumoniformis">
<emphasis id="B95B9E7EFFBC453CFD45147CFC74FEE8" box="[680,987,280,304]" italics="true" pageId="7" pageNumber="584">Isthmiade ichneumoniformis</emphasis>
<bibRefCitation id="EFBE3F9DFFBC453CFC10147CFB66FEE8" author="Bates, H. W." box="[1021,1225,280,304]" pageId="7" pageNumber="584" pagination="391 - 444" refId="ref12254" refString="Bates, H. W. 1870. Contributions to an insect fauna of the Amazon Valley. Transactions of the Entomological Society of London, 391 - 444." type="book chapter" year="1870">Bates, 1870: 326</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A8E647DFFBC453CFD0C145CFCE1FE88" author="Bates, H. W." box="[737,846,312,336]" page="122" pageId="7" pageNumber="584" year="1873">
<bibRefCitation id="EFBE3F9DFFBC453CFD0C145CFCE1FE88" author="Bates, H. W." box="[737,846,312,336]" pageId="7" pageNumber="584" pagination="117 - 132" refId="ref12284" refString="Bates, H. W. 1873. Notes on the Longicorn Coleoptera of Tropical America. The Annals and Magazine of Natural History, (4) 11: 117 - 132." type="journal article" year="1873">1873: 122</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EFBE3F9DFFBC453CFC9B145CFB9AFE88" author="Gounelle, E." box="[886,1077,312,336]" pageId="7" pageNumber="584" pagination="1 - 150" refId="ref12318" refString="Gounelle, E. 1911. Liste des cerambycides de la region de Jatahy, Etat de Goyaz, Bresil. Annales de la Societe Entomologique de France, 80: 1 - 150." type="journal article" year="1911">Gounelle, 1911</bibRefCitation>
a: 51;
<treatmentCitation id="0A8E647DFFBC453CFB7D145CFCE3FEA8" author="Lima, A. M. da &amp; Costa" page="105" pageId="7" pageNumber="584" year="1955">
<bibRefCitation id="EFBE3F9DFFBC453CFB7D145CFCE3FEA8" author="Lima, A. M. da &amp; Costa" pageId="7" pageNumber="584" refId="ref12458" refString="Lima, A. M. da Costa, 1955. Insetos do Brasil. Escola Nacional de Agronomia, Rio de Janeiro, v. 9, 289 p." type="book" year="1955">
<collectingRegion id="49EB8C8EFFBC453CFB7D145CFB67FE88" box="[1168,1224,312,336]" country="Peru" name="Lima" pageId="7" pageNumber="584">Lima</collectingRegion>
, 1955: 105
</bibRefCitation>
</treatmentCitation>
(biol.);
<treatmentCitation id="0A8E647DFFBC453CFC42143CFB2DFEA8" author="Zajciw, D." box="[943,1154,344,368]" page="578" pageId="7" pageNumber="584" year="1972">
<bibRefCitation id="EFBE3F9DFFBC453CFC42143CFB2DFEA8" author="Zajciw, D." box="[943,1154,344,368]" pageId="7" pageNumber="584" pagination="573 - 583" refId="ref12797" refString="Zajciw, D. 1972 b. Revisao das especies sul-americanas do genero Isthmiade Thomson, 1864 (Coleoptera, Cerambycidae, Rhinotragini). Revista Brasileira de Biologia, 32 (4): 573 - 583." type="journal article" year="1972">Zajciw, 1972b: 578</bibRefCitation>
</treatmentCitation>
; Tavakilian &amp; Peñaherrera-Leiva, 2005: 43, figs. 15, 28a-c (lect.);
<treatmentCitation id="0A8E647DFFBC453CFC8414FCFB82FE68" author="Monne, M. A. &amp; Hovore, F. T." box="[873,1069,408,432]" page="482" pageId="7" pageNumber="584" year="2006">
<bibRefCitation id="EFBE3F9DFFBC453CFC8414FCFB82FE68" author="Monne, M. A. &amp; Hovore, F. T." box="[873,1069,408,432]" pageId="7" pageNumber="584" refId="ref12527" refString="Monne, M. A. &amp; Hovore, F. T. 2006. Checklist of the Cerambycidae, or longhorned wood-boring beetles, of the Western Hemisphere. BioQuip, Rancho Dominguez, 394 p." type="book" year="2006">Monné, 2006: 482</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33511E7FFBC4532FD4514BDFDF5FBD7" lastPageId="9" lastPageNumber="586" pageId="7" pageNumber="584" type="diagnosis">
<paragraph id="8B90426CFFBC453CFD4514BDFC98FD88" blockId="7.[680,1229,472,592]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD4514BDFC49FE28" box="[680,998,472,496]" italics="true" pageId="7" pageNumber="584">
Diagnosis:
<taxonomicName id="4C2F39EFFFBC453CFCF214BDFC49FE28" authorityName="Bates" authorityYear="1870" box="[799,998,472,496]" class="Insecta" family="Cerambycidae" genus="Isthmiade" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="ichneumoniformis">I. ichneumoniformis</taxonomicName>
</emphasis>
is readily separated from other Bolivian species by its relatively large size, uniform black head, pale, elongate elytra and orange-red abdomen.
</paragraph>
<paragraph id="8B90426CFFBC453CFD45171CFCC8FCC8" blockId="7.[680,1229,632,784]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD45171CFCDAFD48" box="[680,885,632,656]" italics="true" pageId="7" pageNumber="584">Original description:</emphasis>
Black, shining, elytra testaceousgold, abdomen red (except apex); legs testaceous red, hind leg femora broadly ringed dusky, tibia and tarsus dusky; wings yellow, apical band black. Length 5-7 lines male, female.
</paragraph>
<paragraph id="8B90426CFFBC453CFD45165CFB1AFA48" blockId="7.[680,1229,824,1424]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD45165CFC9EFC88" box="[680,817,824,848]" italics="true" pageId="7" pageNumber="584">Redescription.</emphasis>
Male g
<emphasis id="B95B9E7EFFBC453CFC96165CFC52FC88" box="[891,1021,824,848]" italics="true" pageId="7" pageNumber="584">eneral colour:</emphasis>
Head, pro- and mesosternum, scutellum, urosternites
<collectionCode id="ED3EDAA9FFBC453CFBF7163CFB9AFCB7" box="[1050,1077,856,879]" pageId="7" pageNumber="584">IV</collectionCode>
and
<collectionCode id="ED3EDAA9FFBC453CFB9D163CFB2DFCB7" box="[1136,1154,856,879]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="584" type="Museum">V</collectionCode>
, black. Pronotum usually translucent orange-red across middle, broadly black at front margin and less so at hind margin, but pigmentation of hind margin often protrudes on to disc, or extends along midline to front margin, or, less commonly, black colour reduced to narrow band at front margin (among the material examined pronotum never entirely black, nor entirely orange-red); extreme sides of pronotum black. Elytra translucent orange-yellow (almost vitreous) except area around scutellum, epipleuron from behind humeri to apex, and extreme apices rufous. Broad fascia across apical third of wings translucent yellow to orange-yellow. Metasternum and metepisternum orange-red, urosternites I-III usually so, but most of abdomen may be black. Legs translucent orange-yellow except coxae, apical half of metatibia, and all of metatarsus black; apical two-thirds of metafemoral club dusky.
</paragraph>
<paragraph id="8B90426CFFBC453CFD4510DCFB64F928" blockId="7.[680,1229,1464,1776]" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD4510DCFCC8FA08" box="[680,871,1464,1488]" italics="true" pageId="7" pageNumber="584">General pubescence:</emphasis>
Head almost glabrous, genae with short, recumbent pubescence. Submentum and gula glabrous with group of 2-3 long setae at each side. Sides of pronotum towards base with round patch of dense, long, recumbent pubescence. Prothorax anterior to prosternal process and apical half of metasternum densely hirsute, mesepimeron and base of metasternum clothed with glistening, recumbent, golden pubescence; metepisternum and urosternites I-III almost glabrous,
<collectionCode id="ED3EDAA9FFBC453CFD4513BCFD6CF937" box="[680,707,1752,1775]" pageId="7" pageNumber="584">IV</collectionCode>
and
<collectionCode id="ED3EDAA9FFBC453CFD1913BCFCA9F937" box="[756,774,1752,1775]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="584" type="Museum">V</collectionCode>
with rows of long, erect hairs, denser at sides.
</paragraph>
<paragraph id="8B90426CFFBC4533FD45127CFD0DFD17" blockId="7.[680,1228,1816,1872]" lastBlockId="8.[132,681,184,720]" lastPageId="8" lastPageNumber="585" pageId="7" pageNumber="584">
<emphasis id="B95B9E7EFFBC453CFD45127CFC20F8E8" box="[680,911,1816,1840]" italics="true" pageId="7" pageNumber="584">Surface ornamentation:</emphasis>
Labrum with row of six large punctures interspersed with smaller ones, clypeus impunctate, frons with mix of large and small, confluent punctures in depressed area, otherwise impunctate; vertex between antennal tubercles with single row of confluent punctures, these extending around inner and hind margins of superior lobes in two irregular rows (which may be invested by short, narrow carinas), leaving vertex largely impunctate; submentum with large, rectangular patch of anastomosed punctures, gula impunctate; prothorax anterior to prosternal process with irregular, contiguous punctures, interstices micro-punctate; sides of pronotum under pubescence coarsely and closely punctured (representing the sexual puncturation), the interstices reticulate; elytra with uniform, sparse, small punctures at midline; centre of metasternum with moderately dense, uniform, small punctures; metepisternum and urosternites almost impunctate, some sparse punctures at sides of
<collectionCode id="ED3EDAA9FFB34533FDD317DCFDF6FD17" box="[574,601,696,719]" pageId="8" pageNumber="585">IV</collectionCode>
and
<collectionCode id="ED3EDAA9FFB34533FD6617DCFD32FD17" box="[651,669,696,719]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="585" type="Museum">V</collectionCode>
.
</paragraph>
<paragraph id="8B90426CFFB34533FF69179CFDDBFB28" blockId="8.[132,681,760,1264]" pageId="8" pageNumber="585">
<emphasis id="B95B9E7EFFB34533FF69179CFF4BFCD7" box="[132,228,760,783]" italics="true" pageId="8" pageNumber="585">Structure:</emphasis>
Head. Rostrum distinctly short; frons separated from clypeus by strong, transverse declivity; lengths of gena/inferior lobe 0.7/1.0; inferior lobes subcontiguous, interocular distance/width of lobe 0.2/
<quantity id="4CD7EF89FFB34533FF08161CFE98FC57" box="[229,311,888,912]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="585" unit="mm" value="1.1">1.1 mm</quantity>
; frontal suture represented by deep V-shaped depression extending to hind margin of superior lobes; superior lobes relatively proximate, interocular distance/width of lobe 0.50/
<quantity id="4CD7EF89FFB34533FDAE16BCFD0AFC37" box="[579,677,984,1008]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="585" unit="mm" value="0.25">0.25 mm</quantity>
. Area of submentum depressed with evanescent arced carinas, not separated from gula by distinct declivity. Antennae slender, somewhat short, reaching base of urosternite
<collectionCode id="ED3EDAA9FFB34533FEF6113CFE97FBB7" box="[283,312,1112,1135]" pageId="8" pageNumber="585">III</collectionCode>
; scape (length 1.0 mm) more than three times longer than pedicel, two-thirds length of antennomere
<collectionCode id="ED3EDAA9FFB34533FEF811FCFE9FFB77" box="[277,304,1176,1199]" pageId="8" pageNumber="585">III</collectionCode>
(
<quantity id="4CD7EF89FFB34533FEAC11FCFE39FB68" box="[321,406,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="585" unit="mm" value="1.6">1.6 mm</quantity>
);
<collectionCode id="ED3EDAA9FFB34533FE4111FCFE68FB77" box="[428,455,1176,1199]" pageId="8" pageNumber="585">IV</collectionCode>
(
<quantity id="4CD7EF89FFB34533FE3511FCFD81FB68" box="[472,558,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="585" unit="mm" value="0.9">0.9 mm</quantity>
),
<collectionCode id="ED3EDAA9FFB34533FDAE11FCFDFAFB77" box="[579,597,1176,1199]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="585" type="Museum">V</collectionCode>
and
<collectionCode id="ED3EDAA9FFB34533FD6011FCFD07FB77" box="[653,680,1176,1199]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="585">VI</collectionCode>
equal (
<quantity id="4CD7EF89FFB34533FF2311DCFE8BFB08" box="[206,292,1208,1232]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="585" unit="mm" value="1.5">1.5 mm</quantity>
), remaining antennomeres gradually shortening to X (
<quantity id="4CD7EF89FFB34533FED811BCFE25FB28" box="[309,394,1240,1264]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="585" unit="mm" value="1.1">1.1 mm</quantity>
), X1 longer (
<quantity id="4CD7EF89FFB34533FDFC11BCFDC7FB28" box="[529,616,1240,1264]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="585" unit="mm" value="1.2">1.2 mm</quantity>
).
</paragraph>
<paragraph id="8B90426CFFB34533FF69107CFDA8F988" blockId="8.[132,681,1304,1872]" pageId="8" pageNumber="585">
Thorax. Prothorax distinctly longer (
<quantity id="4CD7EF89FFB34533FDFF107CFDC9FAE8" box="[530,614,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="585" unit="mm" value="2.4">2.4 mm</quantity>
) than width of front and hind margins (both
<quantity id="4CD7EF89FFB34533FDAA105CFD33FA88" box="[583,668,1336,1360]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="585" unit="mm" value="2.1">2.1 mm</quantity>
), deeply constricted at base and apex, the sides strongly rounded and wide (
<quantity id="4CD7EF89FFB34533FEBF101CFE07FA48" box="[338,424,1400,1424]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="585" unit="mm" value="2.4">2.4 mm</quantity>
); disc of pronotum with five distinct calli, those to either side of centre subcircular, central one elongate; base of prosternal process slightly arched, elongate and very narrow (ca. 12 times narrower than width of coxal cavity), apex triangular, large and declivous across apex, the margins narrow and slightly raised, but not explanate.
</paragraph>
<paragraph id="8B90426CFFB34533FF50133CFC71FEE8" blockId="8.[132,681,1304,1872]" lastBlockId="8.[718,1267,184,816]" pageId="8" pageNumber="585">
Mesothorax abruptly declivous before mesosternal process, the latter moderately broad at base at base (1/4 width of coxal cavity), apex weakly cordiform with well elevated sides. Scutellum large, elongate, subtriangular, apex broadly rounded, not bifid; elytral surface adjacent to scutellum elevated and dusky in colour. Elytra elongate (2.6 times width of humeri), reaching middle of urosternite
<collectionCode id="ED3EDAA9FFB34533FE36125CFE40F897" box="[475,495,1848,1871]" pageId="8" pageNumber="585">II</collectionCode>
, strongly acuminate from humerus to apex; dehiscent for apical two-thirds; humero-apical costa weak for middle third, only slightly salient to apex; apices with smooth, blunt, truncate lobe at tip.
</paragraph>
<paragraph id="8B90426CFFB34533FCEA145CFBCBFE48" blockId="8.[718,1267,184,816]" pageId="8" pageNumber="585">Metasternum tumid and convex (planar with pro- and mesocoxae), longitudinal suture not deeply impressed, occupying apical four-fifths.</paragraph>
<paragraph id="8B90426CFFB34533FCEA14FCFBE4FCE8" blockId="8.[718,1267,184,816]" pageId="8" pageNumber="585">
Legs robust; profemur short (3.0 mm), peduncle almost absent, clave large, widest at middle; mesocoxae flattened, mesofemur longer (4.0 mm), peduncle moderately long (
<quantity id="4CD7EF89FFB34533FC02149CFBEAFDC8" box="[1007,1093,504,528]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="585" unit="mm" value="1.2">1.2 mm</quantity>
), clave large, attenuate to apex; metafemur subcylindrical without distinct clave, long (
<quantity id="4CD7EF89FFB34533FC5F175CFBA7FD88" box="[946,1032,568,592]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="585" unit="mm" value="7.5">7.5 mm</quantity>
), reaching middle of urosternite
<collectionCode id="ED3EDAA9FFB34533FCA5173CFCCCFDB7" box="[840,867,600,623]" pageId="8" pageNumber="585">IV</collectionCode>
. Tarsi robust; protarsomeres I and
<collectionCode id="ED3EDAA9FFB34533FB0D173CFB5DFDB7" box="[1248,1266,600,623]" pageId="8" pageNumber="585">II</collectionCode>
subequal, both shorter and much narrower than
<collectionCode id="ED3EDAA9FFB34533FB3A171CFB5DFD57" box="[1239,1266,632,655]" pageId="8" pageNumber="585">III</collectionCode>
(0.45 x
<quantity id="4CD7EF89FFB34533FCCF17FCFC2BFD68" box="[802,900,664,688]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="585" unit="mm" value="0.5">0.50 mm</quantity>
), mesotarsomere I and
<collectionCode id="ED3EDAA9FFB34533FB9617FCFB39FD77" box="[1147,1174,664,687]" pageId="8" pageNumber="585">III</collectionCode>
of equal length (
<quantity id="4CD7EF89FFB34533FCF117DCFCDDFD08" box="[796,882,696,720]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="585" unit="mm" value="0.5">0.5 mm</quantity>
), I subcylindrical, slightly longer than
<collectionCode id="ED3EDAA9FFB34533FD2317BCFD4DFD37" box="[718,738,728,751]" pageId="8" pageNumber="585">II</collectionCode>
, both I and
<collectionCode id="ED3EDAA9FFB34533FC8B17BCFCD7FD37" box="[870,888,728,751]" pageId="8" pageNumber="585">II</collectionCode>
much narrower than
<collectionCode id="ED3EDAA9FFB34533FBB117BCFBD6FD37" box="[1116,1145,728,751]" pageId="8" pageNumber="585">III</collectionCode>
; metatarsomeres I and
<collectionCode id="ED3EDAA9FFB34533FCBC179CFCCCFCD7" box="[849,867,760,783]" pageId="8" pageNumber="585">II</collectionCode>
cylindrical, I longer (
<quantity id="4CD7EF89FFB34533FBAB179CFB35FCC8" box="[1094,1178,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="585" unit="mm" value="1.5">1.5 mm</quantity>
) than
<collectionCode id="ED3EDAA9FFB34533FB0D179CFB5DFCD7" box="[1248,1266,760,783]" pageId="8" pageNumber="585">II</collectionCode>
(
<quantity id="4CD7EF89FFB34533FD3B167CFC85FCE8" box="[726,810,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="585" unit="mm" value="0.7">0.7 mm</quantity>
) and
<collectionCode id="ED3EDAA9FFB34533FC88167CFC2FFCF7" box="[869,896,792,815]" pageId="8" pageNumber="585">III</collectionCode>
(
<quantity id="4CD7EF89FFB34533FC63167CFC4DFCE8" box="[910,994,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="585" unit="mm" value="0.5">0.5 mm</quantity>
) together.
</paragraph>
<paragraph id="8B90426CFFB34533FD23163CFB88FB88" blockId="8.[718,1267,856,1104]" pageId="8" pageNumber="585">
Abdomen. Moderately robust and elongate, subparallel, urosternite I longest (
<quantity id="4CD7EF89FFB34533FBF9161CFBC5FC48" box="[1044,1130,888,912]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="585" unit="mm" value="2.9">2.9 mm</quantity>
), trapezoidal and constricted at middle;
<collectionCode id="ED3EDAA9FFB34533FC0116FCFC51FC77" box="[1004,1022,920,943]" pageId="8" pageNumber="585">II</collectionCode>
(
<quantity id="4CD7EF89FFB34533FBE216FCFBCAFC68" box="[1039,1125,920,944]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="585" unit="mm" value="2.2">2.2 mm</quantity>
), rectangular, subparallel and weakly constricted at basal third;
<collectionCode id="ED3EDAA9FFB34533FB3A16DCFB5DFC17" box="[1239,1266,952,975]" pageId="8" pageNumber="585">III</collectionCode>
(
<quantity id="4CD7EF89FFB34533FD3B16BCFC85FC28" box="[726,810,984,1008]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="585" unit="mm" value="2.1">2.1 mm</quantity>
) quadrate;
<collectionCode id="ED3EDAA9FFB34533FC5D16BCFC64FC37" box="[944,971,984,1007]" pageId="8" pageNumber="585">IV</collectionCode>
subquadrate (
<quantity id="4CD7EF89FFB34533FB8216BCFB6AFC28" box="[1135,1221,984,1008]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="585" unit="mm" value="1.6">1.6 mm</quantity>
);
<collectionCode id="ED3EDAA9FFB34533FB0D16BCFB5DFC37" box="[1248,1266,984,1007]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="585" type="Museum">V</collectionCode>
(
<quantity id="4CD7EF89FFB34533FD3A169CFC85FBC8" box="[727,810,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="585" unit="mm" value="1.1">1.1 mm</quantity>
) trapezoidal, slightly depressed from apex to middle, apical margin slightly sinuate; abdominal process long and narrow, whitish.
</paragraph>
<paragraph id="8B90426CFFB34533FD23111CFB4DF968" blockId="8.[718,1267,1144,1712]" pageId="8" pageNumber="585">
<emphasis id="B95B9E7EFFB34533FD23111CFCB6FB48" box="[718,793,1144,1168]" italics="true" pageId="8" pageNumber="585">Female:</emphasis>
Rostrum hardly longer than male, length of genae equal to length of inferior lobe (
<quantity id="4CD7EF89FFB34533FB8E11FCFB17FB68" box="[1123,1208,1176,1200]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="585" unit="mm" value="0.7">0.7 mm</quantity>
), distance between inferior lobes (
<quantity id="4CD7EF89FFB34533FC1311DCFBFCFB08" box="[1022,1107,1208,1232]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="585" unit="mm" value="0.7">0.7 mm</quantity>
), width of lobe (
<quantity id="4CD7EF89FFB34533FD3B11BCFC84FB28" box="[726,811,1240,1264]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="585" unit="mm" value="0.8">0.8 mm</quantity>
); frons multi carinate from antennal tubercles to apical declivity, each side (between outer margin and frontal suture) with three very narrow carinas and three slightly broader sulci, this arrangement sometimes subdivided by further short carinas in an asymmetrical manner. Patch of pubescence at sides of pronotum in males much reduced in extent and density, and size of punctures minute. Metasternum more convex behind and more salient than mesocoxa (even though these are more salient than in male), the latter not flattened. Abdomen robust, fusiform; urosternite
<collectionCode id="ED3EDAA9FFB34533FB9A137CFB3DF9F7" box="[1143,1170,1560,1583]" pageId="8" pageNumber="585">IV</collectionCode>
elongate (0.1.6 mm) and trapezoidal;
<collectionCode id="ED3EDAA9FFB34533FBE5135CFBB5F997" box="[1032,1050,1592,1615]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="585" type="Museum">V</collectionCode>
elongate (2.0 mm), conical (apex rounded, not angular), narrow (basal margin
<quantity id="4CD7EF89FFB34533FCF3131CFCDEF957" box="[798,881,1656,1680]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="585" unit="mm" value="1.2">1.2 mm</quantity>
, apical margin
<quantity id="4CD7EF89FFB34533FBE0131CFBF0F948" box="[1037,1119,1656,1680]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="585" unit="mm" value="0.7">0.7 mm</quantity>
) and regularly convex; abdominal process same colour as abdomen.
</paragraph>
<paragraph id="8B90426CFFB34532FD2313BCFE6CFD08" blockId="8.[718,1268,1752,1872]" lastBlockId="9.[94,643,184,720]" lastPageId="9" lastPageNumber="586" pageId="8" pageNumber="585">
Measurements (mm):
<specimenCount id="9D2989E5FFB34533FC5D13BCFBA1F928" box="[944,1038,1752,1776]" count="24" pageId="8" pageNumber="585" type="male">24 males</specimenCount>
/
<specimenCount id="9D2989E5FFB34533FBF813BCFBDCF928" box="[1045,1139,1752,1776]" count="4" pageId="8" pageNumber="585" type="female">4 females</specimenCount>
respectively: total length 15.3-18.5/17.4-18.3; length of pronotum 2.1-2.6/2.3-2.4;widthofpronotum2.0-2.6/2.4;lengthof elytra 6.4-7.9/7.2-7.4; width of humeri 2.1-2.8/2.7-2.9.
<emphasis id="B95B9E7EFFB24532FFB315DCFDA1FF08" box="[94,526,184,208]" italics="true" pageId="9" pageNumber="586">
Material examined (in the
<collectionCode id="ED3EDAA9FFB24532FE8B15DCFE0EFF17" box="[358,417,184,207]" pageId="9" pageNumber="586">RCSZ</collectionCode>
collection):
</emphasis>
<collectingCountry id="F33802FCFFB24532FDFB15DCFDD1FF08" box="[534,638,184,208]" name="Bolivia" pageId="9" pageNumber="586">BOLIVIA</collectingCountry>
,
<collectingRegion id="49EB8C8EFFB24532FFB215BCFF7BFF28" box="[95,212,216,240]" country="Bolivia" name="Santa Cruz" pageId="9" pageNumber="586">Santa Cruz</collectingRegion>
, 17°29’96”S/
<geoCoordinate id="EE1B24ABFFB24532FEB015BCFE49FF28" box="[349,486,216,240]" degrees="63" direction="west" minutes="39" orientation="longitude" pageId="9" pageNumber="586" precision="15" seconds="13" value="-63.653614">63° 39’13”W</geoCoordinate>
,
<quantity id="4CD7EF89FFB24532FE1B15BCFD95FF37" box="[502,570,216,240]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="586" unit="m" value="420.0">420 m</quantity>
, Hotel Flora &amp; Fauna,
<quantity id="4CD7EF89FFB24532FF14159CFE82FEC8" box="[249,301,248,272]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="586" unit="km" value="5.0">5 km</quantity>
SSE Buena Vista,
<collectionCode id="ED3EDAA9FFB24532FE0C159CFE5AFED7" box="[481,501,248,271]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="586">R</collectionCode>
. Clarke/S. Zamalloa col. The following on/flying to flowers of “Bejuco hoja lanuda”:
<specimenCount id="9D2989E5FFB24532FF05145CFEEFFE88" box="[232,320,312,336]" count="11" pageId="9" pageNumber="586" type="male">11 males</specimenCount>
<date id="FF9164ACFFB24532FEAB145CFE16FE88" box="[326,441,312,336]" pageId="9" pageNumber="586" value="2005-04-21">
21.
<collectionCode id="ED3EDAA9FFB24532FE8F145CFED0FE88" box="[354,383,312,336]" pageId="9" pageNumber="586">IV</collectionCode>
.2005
</date>
,
<specimenCount id="9D2989E5FFB24532FE29145CFDA9FE88" box="[452,518,312,336]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FDE0145CFDC4FE88" box="[525,619,312,336]" pageId="9" pageNumber="586" value="2005-05-06">
6.
<collectionCode id="ED3EDAA9FFB24532FDF0145CFD9EFE88" box="[541,561,312,336]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="586" type="Museum">V</collectionCode>
.2005
</date>
,
<specimenCount id="9D2989E5FFB24532FD98145CFF22FEA8" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FF7C143CFF54FEA8" box="[145,251,344,368]" pageId="9" pageNumber="586" value="2005-05-10">
10.
<collectionCode id="ED3EDAA9FFB24532FF40143CFF6EFEA8" box="[173,193,344,368]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="586" type="Museum">V</collectionCode>
.2005
</date>
,
<specimenCount id="9D2989E5FFB24532FEEE143CFEEDFEA8" box="[259,322,344,368]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FEAB143CFE1FFEA8" box="[326,432,344,368]" pageId="9" pageNumber="586" value="2005-05-11">
11.
<collectionCode id="ED3EDAA9FFB24532FE8D143CFEDBFEA8" box="[352,372,344,368]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="586" type="Museum">V</collectionCode>
.2005
</date>
,
<specimenCount id="9D2989E5FFB24532FE5A143CFDBEFEA8" box="[439,529,344,368]" count="2" pageId="9" pageNumber="586" type="female">2 females</specimenCount>
<date id="FF9164ACFFB24532FDF9143CFD2FFEA8" box="[532,640,344,368]" pageId="9" pageNumber="586" value="2005-05-12">
12.
<collectionCode id="ED3EDAA9FFB24532FDDD143CFDEBFEA8" box="[560,580,344,368]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="586" type="Museum">V</collectionCode>
.2005
</date>
. The following on/flying to flowers of “Barbasquillo” vine:
<specimenCount id="9D2989E5FFB24532FF7A14FCFF72FE68" box="[151,221,408,432]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FF0A14FCFECEFE68" box="[231,353,408,432]" pageId="9" pageNumber="586" value="2005-08-03">3.VIII.2005</date>
,
<specimenCount id="9D2989E5FFB24532FE8214FCFE12FE68" box="[367,445,408,432]" count="2" pageId="9" pageNumber="586" type="male">2 males</specimenCount>
<date id="FF9164ACFFB24532FE2B14FCFDE1FE68" box="[454,590,408,432]" pageId="9" pageNumber="586" value="2005-08-30">30.VIII.2005</date>
. The following on/flying to flowers of “Sapamoisi”:
<specimenCount id="9D2989E5FFB24532FDD814DCFD2CFE08" box="[565,643,440,464]" count="3" pageId="9" pageNumber="586" type="male">3 males</specimenCount>
<date id="FF9164ACFFB24532FFB214BCFF73FE28" box="[95,220,472,496]" pageId="9" pageNumber="586" value="2005-12-21">21.XII.2005</date>
,
<specimenCount id="9D2989E5FFB24532FF0A14BCFE86FE28" box="[231,297,472,496]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FEDD14BCFE02FE28" box="[304,429,472,496]" pageId="9" pageNumber="586" value="2005-12-24">24.XII.2005</date>
,
<specimenCount id="9D2989E5FFB24532FE5514BCFE55FE28" box="[440,506,472,496]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FDEC14BCFDD0FE28" box="[513,639,472,496]" pageId="9" pageNumber="586" value="2005-12-27">27.XII.2005</date>
. The following on/flying to flowers of “Tutumillo espinoso”:
<specimenCount id="9D2989E5FFB24532FF4B177CFF42FDE8" box="[166,237,536,560]" count="1" pageId="9" pageNumber="586" type="male">1 male</specimenCount>
<date id="FF9164ACFFB24532FF1A177CFECBFDE8" box="[247,356,536,560]" pageId="9" pageNumber="586" value="2006-10-20">20.X.2006</date>
. The following on/flying to flowers of “Sama blanca”:
<specimenCount id="9D2989E5FFB24532FEB1175CFE00FD88" box="[348,431,568,592]" count="1" pageId="9" pageNumber="586" type="female">1 female</specimenCount>
<date id="FF9164ACFFB24532FE59175CFD84FD88" box="[436,555,568,592]" pageId="9" pageNumber="586" value="2006-11-11">11.XI.2006</date>
. The following from different locality and host flower:
<collectingCountry id="F33802FCFFB24532FDC2173CFFD5FD48" name="Bolivia" pageId="9" pageNumber="586">BOLIVIA</collectingCountry>
,
<collectingRegion id="49EB8C8EFFB24532FF68171CFF58FD48" box="[133,247,632,656]" country="Bolivia" name="Santa Cruz" pageId="9" pageNumber="586">Santa Cruz</collectingRegion>
, road to San Javier,
<quantity id="4CD7EF89FFB24532FE2F171CFDACFD48" box="[450,515,632,656]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="586" unit="km" value="12.0">12 km</quantity>
ENE Buena Vista,
<collectionCode id="ED3EDAA9FFB24532FF7117FCFF1FFD77" box="[156,176,664,687]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="586">R</collectionCode>
. Clarke/S. Zamalloa col., on/flying to flowers of “Esquiziton”:
<specimenCount id="9D2989E5FFB24532FF0A17DCFE94FD08" box="[231,315,696,720]" count="1" pageId="9" pageNumber="586" type="female">1 female</specimenCount>
<date id="FF9164ACFFB24532FEAD17DCFE10FD08" box="[320,447,696,720]" pageId="9" pageNumber="586" value="2005-12-14">14.XII.2005</date>
.
</paragraph>
<paragraph id="8B90426CFFB24532FFB3179DFDF5FBD7" blockId="9.[94,643,760,1040]" pageId="9" pageNumber="586">
<emphasis id="B95B9E7EFFB24532FFB3179DFF66FCC8" box="[94,201,761,784]" italics="true" pageId="9" pageNumber="586">Discussion:</emphasis>
Maybe
<taxonomicName id="4C2F39EFFFB24532FEFA179DFE74FCC8" authorityName="Bates" authorityYear="1870" box="[279,475,760,784]" class="Insecta" family="Cerambycidae" genus="Isthmiade" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="586" phylum="Arthropoda" rank="species" species="ichneumoniformis">
<emphasis id="B95B9E7EFFB24532FEFA179DFE74FCC8" box="[279,475,760,784]" italics="true" pageId="9" pageNumber="586">I. ichneumoniformis</emphasis>
</taxonomicName>
is more than one species, or a superspecies, but the Bolivian specimens look very like the illustration of the
<typeStatus id="5494FCCEFFB24532FE32165CFDECFC88" box="[479,579,824,848]" pageId="9" pageNumber="586" type="lectotype">Lectotype</typeStatus>
given by
<bibRefCitation id="EFBE3F9DFFB24532FF93163CFDA1FCA8" author="Tavakilian, G. L. &amp; PeNahererra-Leiva, A. Y." box="[126,526,856,880]" pageId="9" pageNumber="586" pagination="27 - 50" refId="ref12599" refString="Tavakilian, G. L. &amp; PeNahererra-Leiva, A. Y. 2005. Nouvelles especes et nouveaux genres de Rhinotragini (Coleoptera, Cerambycidae) IV. Coleopteres, 11 (5): 27 - 50." type="journal article" year="2005">Tavakilian &amp; Peñahererra-Leiva (2005)</bibRefCitation>
except this specimen has entirely black pronotum and dark fascia on pro- and mesofemora. Since Bates’ description, four lines of Latin, is inadequate for comparative purposes, only evaluation of specimens from all parts of its geographical distribution will determine its status.
</paragraph>
</subSubSection>
</treatment>
</document>
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