279 lines
40 KiB
XML
279 lines
40 KiB
XML
<document id="DC768F924B83032458ABC6A57DBA7FC0" ID-DOI="10.11646/zootaxa.5514.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849826" ID-ZooBank="D745608E-E51C-401E-A1A1-3573A24C51BD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727441634877" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany, Merchan-Vargas, Geraldine Lucia & Soares, Matheus M. M." docDate="2024" docId="03A71175FFC23C66CB966B37825DFEC8" docLanguage="en" docName="zootaxa.5514.3.3.pdf" docOrigin="Zootaxa 5514 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus grossus Ramos-Pastrana, Merchan-Vargas & Soares, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="267" masterDocId="FF9E690DFFCA3C6CCB01695C832BFF88" masterDocTitle="First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species" masterLastPageNumber="272" masterPageNumber="257" pageNumber="265" updateTime="1727443949234" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DBA4BABF23565A376528929A10B658BA">
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<mods:title id="B3A9F499A557BE5DF6D7422F4094AADC">First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species</mods:title>
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<mods:name id="407A57252D77E451368A3E6E929A6623" type="personal">
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<mods:namePart id="52692DF315B0E8B4F3865865066E54C7">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="F0E2D11C1807D9F96A19767EC964C3F2">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:namePart id="8D1462FED029C77A21D2E2EB12567CA3">Merchan-Vargas, Geraldine Lucia</mods:namePart>
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<mods:affiliation id="576276D092C1890AD9335F1F38F9FA4B">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:namePart id="F5D58F8981E5A0173C480C3238D20088">Soares, Matheus M. M.</mods:namePart>
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<mods:affiliation id="CC6905C9EFDB27988E4FB89BD9387EAE">Museu de Zoologia da Universidade de São Paulo, Laboratório de Diptera, Av. Nazaré, 481, 04263 - 000 - Ipiranga, São Paulo, São Paulo, Brazil</mods:affiliation>
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<mods:title id="39A2D06DD4EDD3B231CF967877577A05">Zootaxa</mods:title>
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<mods:part id="19639F0A75CF8712F614914A24B02D88">
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<mods:date id="F56C07F5E94B9A5359ACA321C0BE68EA">2024</mods:date>
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<mods:number id="B33E9CFCB062CB90402DD628863445F6">2024-09-27</mods:number>
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<mods:number id="CD5331BDF22C14813474E75D7172A1C0">5514</mods:number>
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<mods:classification id="01395C4B0226D241716EF7781CF3485C">journal article</mods:classification>
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<treatment id="03A71175FFC23C66CB966B37825DFEC8" ID-DOI="http://doi.org/10.5281/zenodo.13849926" ID-Zenodo-Dep="13849926" LSID="urn:lsid:plazi:treatment:03A71175FFC23C66CB966B37825DFEC8" httpUri="http://treatment.plazi.org/id/03A71175FFC23C66CB966B37825DFEC8" lastPageId="10" lastPageNumber="267" pageId="8" pageNumber="265">
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<subSubSection id="C314F3E8FFC23C64CB966B3782E2FD0D" box="[151,457,619,645]" pageId="8" pageNumber="265" type="nomenclature">
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<paragraph id="8BB1A063FFC23C64CB966B3782E2FD0D" blockId="8.[151,457,619,681]" box="[151,457,619,645]" pageId="8" pageNumber="265">
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<heading id="D0F9170FFFC23C64CB966B3782E2FD0D" bold="true" box="[151,457,619,645]" fontSize="11" level="1" pageId="8" pageNumber="265" reason="1">
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<emphasis id="B97A7C71FFC23C64CB966B3782E2FD0D" bold="true" box="[151,457,619,645]" pageId="8" pageNumber="265">
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<taxonomicName id="4C0EDBE0FFC23C64CB966B378248FD0D" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[151,355,619,645]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="grossus" status="sp. nov.">
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<emphasis id="B97A7C71FFC23C64CB966B378248FD0D" bold="true" box="[151,355,619,645]" italics="true" pageId="8" pageNumber="265">Systenus grossus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A249C10AFFC23C64CA6A6B3782E2FD0D" box="[363,457,619,645]" pageId="8" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C314F3E8FFC23C64CB966BD38220FD21" box="[151,267,655,681]" pageId="8" pageNumber="265" type="description">
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<paragraph id="8BB1A063FFC23C64CB966BD38220FD21" blockId="8.[151,457,619,681]" box="[151,267,655,681]" pageId="8" pageNumber="265">
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(
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<figureCitation id="1335BCE6FFC23C64CB9E6BD383C3FD21" box="[159,232,655,681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Figs 4</figureCitation>
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,
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<figureCitation id="1335BCE6FFC23C64CBF56BD38228FD21" box="[244,259,655,681]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[301,1286,1429,1917]" captionTargetId="figure-56@13.[299,1288,1427,1919]" captionTargetPageId="13" captionText="FIGURE 7. Geographical records of Systenus Loew in Colombia. S. beatae Naglis, 2000; S. bickeli sp. nov.; S. colombiensis sp. nov.; S. flaviatus Naglis, 2000; S. fuscus sp. nov.; S. grossus sp. nov.; S. maculipennis Bickel, 2015; S. naranjensis Bickel, 2015 and S. parkeri Bickel, 2015." figureDoi="http://doi.org/10.5281/zenodo.13849845" httpUri="https://zenodo.org/record/13849845/files/figure.png" pageId="8" pageNumber="265">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C314F3E8FFC23C64CB966B8A816FFC2D" pageId="8" pageNumber="265" type="diagnosis">
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<paragraph id="8BB1A063FFC23C64CB966B8A816FFC2D" blockId="8.[151,1437,726,1905]" pageId="8" pageNumber="265">
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<emphasis id="B97A7C71FFC23C64CB966B8A8239FD78" bold="true" box="[151,274,726,752]" pageId="8" pageNumber="265">Diagnosis.</emphasis>
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Antenna mostly dark brown, except ventral edge of scape and pedicel and a narrow triangular basoventral area of postpedicel orangish yellow (
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<figureCitation id="1335BCE6FFC23C64C93D6BA781BDFC9D" box="[572,662,763,789]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4B</figureCitation>
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). Postpedicel abruptly narrowed at 1/2 apical, 3.5X as long as basal height; arista short, about 1/2 as long as postpedicel (
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<figureCitation id="1335BCE6FFC23C64C9F26A438067FCB1" box="[755,844,799,825]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4B</figureCitation>
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). Metepimeron brown (
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<figureCitation id="1335BCE6FFC23C64CF596A438798FCB1" box="[1112,1203,799,825]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A</figureCitation>
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). Wing hyaline (
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<figureCitation id="1335BCE6FFC23C64CE6E6A438395FCD5" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4D</figureCitation>
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). Legs yellow, except basal 2/3 of coxa II, It
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<subScript id="178AA226FFC23C64C9BD6A0981EFFCEB" attach="left" box="[700,708,853,867]" fontSize="6" pageId="8" pageNumber="265">5</subScript>
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, IIt
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<subScript id="178AA226FFC23C64C9EF6A09802CFCEB" attach="left" box="[750,775,853,867]" fontSize="6" pageId="8" pageNumber="265">3–5</subScript>
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and IIIt
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<subScript id="178AA226FFC23C64C8656A098056FCEB" attach="left" box="[868,893,853,867]" fontSize="6" pageId="8" pageNumber="265">3–5</subScript>
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brown (
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<figureCitation id="1335BCE6FFC23C64C8DC6A1F8711FCD5" box="[989,1082,835,861]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A</figureCitation>
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). Surstylus slightly longer than epandrial capsule, very thickened medially, abruptly narrowed and curved ventrally at apex, 1 digitiform process at basal 1/3 of inner surface (
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<figureCitation id="1335BCE6FFC23C64CAC36AD7811CFC2D" box="[450,567,907,933]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4F, G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C314F3E8FFC23C65CBC66AF28289F8EC" lastPageId="9" lastPageNumber="266" pageId="8" pageNumber="265" type="description">
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<paragraph id="8BB1A063FFC23C64CBC66AF28743F8F9" blockId="8.[151,1437,726,1905]" pageId="8" pageNumber="265">
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<emphasis id="B97A7C71FFC23C64CBC66AF282B5FC41" bold="true" box="[199,414,942,969]" pageId="8" pageNumber="265">Description. Male</emphasis>
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(
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<figureCitation id="1335BCE6FFC23C64CAAF6AF382DFFC41" box="[430,500,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4</figureCitation>
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). Body length:
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<quantity id="4CF60D86FFC23C64C9A56AF381D5FC41" box="[676,766,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="265" unit="mm" value="2.6">2.6 mm</quantity>
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; wing
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<quantity id="4CF60D86FFC23C64C84A6AF3808EFC41" box="[843,933,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="265" unit="mm" value="1.9">1.9 mm</quantity>
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.
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<emphasis id="B97A7C71FFC23C64C8B06AF380DAFC41" bold="true" box="[945,1009,943,969]" pageId="8" pageNumber="265">Head</emphasis>
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(
|
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<figureCitation id="1335BCE6FFC23C64CF006AF38757FC41" box="[1025,1148,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A, B</figureCitation>
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). Similar to
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<taxonomicName id="4C0EDBE0FFC23C64CE0A6AF38658FC41" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[1291,1395,943,969]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
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<emphasis id="B97A7C71FFC23C64CE0A6AF38632FC41" box="[1291,1305,943,969]" italics="true" pageId="8" pageNumber="265">S</emphasis>
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||
.
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<emphasis id="B97A7C71FFC23C64CE296AF38658FC41" box="[1320,1395,943,969]" italics="true" pageId="8" pageNumber="265">bickeli</emphasis>
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</taxonomicName>
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||
<emphasis id="B97A7C71FFC23C64CE7A6AF383E3FC65" bold="true" pageId="8" pageNumber="265">
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<taxonomicNameLabel id="A249C10AFFC23C64CE7A6AF383E3FC65" pageId="8" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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except as noted: Palpus pale brown with short, black setae at apical 1/2. Proboscis pale brown. Antenna mostly dark brown, except ventral edge of scape and pedicel and narrow triangular basoventral area of postpedicel orangish yellow; postpedicel abruptly narrowed at 1/2 apical, 3.5X as long as basal height; arista short, about 1/2 as long as postpedicel. One row of postocular setae, uppermost 5 setae black, remaining setae pale brown.
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<emphasis id="B97A7C71FFC23C64CFD46D638605FBD1" bold="true" box="[1237,1326,1087,1113]" pageId="8" pageNumber="265">Thorax</emphasis>
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(
|
||
<figureCitation id="1335BCE6FFC23C64CE3F6D638386FBF5" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A, C</figureCitation>
|
||
). Mostly metallic light green with weak bluish reflections. Metepimeron brown.
|
||
<emphasis id="B97A7C71FFC23C64CF3E6D3F875CFBF5" bold="true" box="[1087,1143,1123,1149]" pageId="8" pageNumber="265">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC23C64CF866D3F87C8FBF5" box="[1159,1251,1123,1149]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A</figureCitation>
|
||
). Yellow, except basal 2/3 of coxa II, It
|
||
<subScript id="178AA226FFC23C64CA8A6DC582B8FB2F" attach="right" box="[395,403,1177,1191]" fontSize="6" pageId="8" pageNumber="265">5</subScript>
|
||
, IIt
|
||
<subScript id="178AA226FFC23C64CABB6DC582F8FB2F" attach="left" box="[442,467,1177,1191]" fontSize="6" pageId="8" pageNumber="265">3–5</subScript>
|
||
and IIIt
|
||
<subScript id="178AA226FFC23C64C92D6DC5816FFB2F" attach="left" box="[556,580,1177,1191]" fontSize="6" pageId="8" pageNumber="265">3–5</subScript>
|
||
brown.
|
||
<emphasis id="B97A7C71FFC23C64C99E6DDB81CFFB29" bold="true" box="[671,740,1159,1185]" pageId="8" pageNumber="265">Leg I.</emphasis>
|
||
Podomere ratios: 55, 50, 22/12/9/7/6. Anterior surface of coxa I covered by short vestiture of yellow setae, 3 long yellow setae at apical edge. Femur I clothed by short vestiture of short, brown setae, except ventral surface with yellow setae basal, ventral surface of basal 1/2 mostly bare. Tibia I clothed by short vestiture of yellow setae.
|
||
<emphasis id="B97A7C71FFC23C64C96F6DAF8195FA85" bold="true" box="[622,702,1267,1293]" pageId="8" pageNumber="265">Leg II.</emphasis>
|
||
Podomere ratios: 60, 63, 33/22/16/12/ 8. Anterior surface of coxa II covered by short vestiture of yellow setae, longer at apical 1/2. Femur II clothed by short vestiture of brown setae, except ventral surface mostly bare, apical 1/2 with yellow setae. Tibia II with 2 pairs of antero- and posterodorsal setae at basal 1/4 and at middle, pair of ventral setae at apical 2/4, 1 pair of posterodorsal setae, 1 at basal 1/5 and 1 at apical 1/5 and crown of setae at apex: 1 ventral, 1 anterodorsal, 1 posterodorsal and 1 dorsal.
|
||
<emphasis id="B97A7C71FFC23C64CFC36CDF860FFA15" bold="true" box="[1218,1316,1411,1437]" pageId="8" pageNumber="265">Leg. III.</emphasis>
|
||
Podomere ratios: 60, 78, 14/30/20/14/8. Lateral surface of coxa III with 1 strong, yellow seta near middle. Femur III with dorsal surface clothed by short vestiture of brown setae, ventral and anterior surface clothed by short vestiture of yellow setae, ventral and posterior surface mostly bare. Tibia III with row of short, strong setae dorsally, starting at basal 1/4 and ending at apical 3/4.
|
||
<emphasis id="B97A7C71FFC23C64C93D6F4E8156F9A4" bold="true" box="[572,637,1554,1580]" pageId="8" pageNumber="265">Wing</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC23C64C98C6F4F81C2F9A5" box="[653,745,1555,1581]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4D</figureCitation>
|
||
). Hyaline, veins pale brown. R
|
||
<subScript id="178AA226FFC23C64CF476F798774F9BB" attach="left" box="[1094,1119,1573,1587]" fontSize="6" pageId="8" pageNumber="265">2+3</subScript>
|
||
nearly straight, R
|
||
<subScript id="178AA226FFC23C64CE2B6F79866FF9BB" attach="left" box="[1322,1348,1573,1587]" fontSize="6" pageId="8" pageNumber="265">4+5</subScript>
|
||
and M
|
||
<subScript id="178AA226FFC23C64CE956F7986B7F9BB" attach="left" box="[1428,1436,1573,1587]" fontSize="6" pageId="8" pageNumber="265">1</subScript>
|
||
slightly bowed at apical 6.5/8 and ending parallel at apex.
|
||
<emphasis id="B97A7C71FFC23C64C8166F6A80A7F9D8" bold="true" box="[791,908,1590,1616]" pageId="8" pageNumber="265">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC23C64C8986F6B873BF9D9" box="[921,1040,1591,1617]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4A, E</figureCitation>
|
||
). Mostly metallic green, covered by short vestiture of black setae, tergite 1 with long setae at posterior margin, tergite 2–5 with brown medial and lateral bands, tergite 6–7 entirely brown, tergite 7 brown, long and setose, sternite 8 brown.
|
||
<emphasis id="B97A7C71FFC23C64CF686F22862EF910" bold="true" box="[1129,1285,1662,1688]" pageId="8" pageNumber="265">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC23C64CE176F2386BBF911" box="[1302,1424,1663,1689]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4F, G</figureCitation>
|
||
). Surstylus slightly longer than epandrial capsule, very thickened medially, abruptly narrowed and curved ventrally at apex, with 1 digitiform process at basal 1/3 of inner surface (
|
||
<figureCitation id="1335BCE6FFC23C64C8686F9B80E8F969" box="[873,963,1735,1761]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="8" pageNumber="265">Fig. 4G</figureCitation>
|
||
), 3 long setae at basal 1/2 of ventral edge and 1 ribbon-like seta at apex preceded by long seta before apical bend. Epandrial lobe as wide as 1/3 of surstylus at widest point, only with apex strongly curved ventrally, with 1 short digitiform basodorsal process and 2 long setae at ventral edge, 1 basally and 1 apically. Cercus yellow, with rounded swollen base, gradually narrowed at apex, as long as epandrial capsule, about as long as surstylus and covered by short yellow setae.
|
||
</paragraph>
|
||
<caption id="DF71F0EBFFC33C65CB966FFC8045F888" ID-DOI="http://doi.org/10.5281/zenodo.13849838" ID-Zenodo-Dep="13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="9" pageNumber="266" startId="9.[151,250,1696,1720]" targetBox="[162,1426,185,1668]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BB1A063FFC33C65CB966FFC8045F888" blockId="9.[151,1436,1696,1792]" pageId="9" pageNumber="266">
|
||
<emphasis id="B97A7C71FFC33C65CB966FFC823FF930" bold="true" box="[151,276,1696,1720]" pageId="9" pageNumber="266">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C0EDBE0FFC33C65CA1D6FFD82ECF930" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[284,455,1697,1720]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="266" phylum="Arthropoda" rank="species" species="grossus" status="sp. nov.">
|
||
<emphasis id="B97A7C71FFC33C65CA1D6FFD82ECF930" box="[284,455,1697,1720]" italics="true" pageId="9" pageNumber="266">Systenus grossus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A7C71FFC33C65CACE6FFC810AF930" bold="true" box="[463,545,1696,1720]" pageId="9" pageNumber="266">
|
||
<taxonomicNameLabel id="A249C10AFFC33C65CACE6FFC810AF930" box="[463,545,1696,1720]" pageId="9" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (before dissection).
|
||
<emphasis id="B97A7C71FFC33C65C8976FFC8084F930" bold="true" box="[918,943,1696,1720]" pageId="9" pageNumber="266">A.</emphasis>
|
||
Habitus, lateral view;
|
||
<emphasis id="B97A7C71FFC33C65CF9B6FFC879AF930" bold="true" box="[1178,1201,1696,1720]" pageId="9" pageNumber="266">B.</emphasis>
|
||
Head, lateral view;
|
||
<emphasis id="B97A7C71FFC33C65CE856FFC86B7F930" bold="true" box="[1412,1436,1696,1720]" pageId="9" pageNumber="266">C.</emphasis>
|
||
Thorax, dorsal view;
|
||
<emphasis id="B97A7C71FFC33C65CA716F9882A3F954" bold="true" box="[368,392,1732,1756]" pageId="9" pageNumber="266">D.</emphasis>
|
||
Left wing;
|
||
<emphasis id="B97A7C71FFC33C65CAFE6F98813DF954" bold="true" box="[511,534,1732,1756]" pageId="9" pageNumber="266">E.</emphasis>
|
||
Abdomen, dorsal view;
|
||
<emphasis id="B97A7C71FFC33C65C80E6F988009F954" bold="true" box="[783,802,1732,1756]" pageId="9" pageNumber="266">F.</emphasis>
|
||
Hypopygium, left lateral view;
|
||
<emphasis id="B97A7C71FFC33C65CF666F9887AAF954" bold="true" box="[1127,1153,1732,1756]" pageId="9" pageNumber="266">G.</emphasis>
|
||
Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB1A063FFC33C65CBC66E178289F8EC" blockId="9.[151,1437,1866,2036]" box="[199,418,1866,1893]" pageId="9" pageNumber="266">
|
||
<emphasis id="B97A7C71FFC33C65CBC66E17820EF8ED" bold="true" box="[199,293,1867,1893]" pageId="9" pageNumber="266">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFC33C65CBC66E3382E9F87C" pageId="9" pageNumber="266" type="materials_examined">
|
||
<paragraph id="8BB1A063FFC33C65CBC66E3382E9F87C" blockId="9.[151,1437,1866,2036]" pageId="9" pageNumber="266">
|
||
<emphasis id="B97A7C71FFC33C65CBC66E33825CF800" bold="true" box="[199,375,1902,1929]" pageId="9" pageNumber="266">Type material.</emphasis>
|
||
<materialsCitation id="3B66AA3EFFC33C65CA816E328037F858" collectingDate="2017-03-01" collectingDateMax="2017-03-15" collectingDateMin="2017-03-01" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana Leg." country="Colombia" county="Florencia" elevation="643" latitude="1.7066667" location="San Francisco" longLatPrecision="21" longitude="-75.61" municipality="Vereda" pageId="9" pageNumber="266" specimenCount="1" specimenCount-male="1" stateProvince="Caqueta" typeStatus="holotype">
|
||
<emphasis id="B97A7C71FFC33C65CA816E328134F800" bold="true" box="[384,543,1902,1928]" pageId="9" pageNumber="266">
|
||
<typeStatus id="54B51EC1FFC33C65CA816E328134F800" box="[384,543,1902,1928]" pageId="9" pageNumber="266" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D086BEAFFC33C65C9296E338114F801" box="[552,575,1903,1929]" count="1" pageId="9" pageNumber="266" type="male">♂</specimenCount>
|
||
, labelled: “
|
||
<collectingCountry id="F319E0F3FFC33C65C9C16E338070F801" box="[704,859,1903,1929]" name="Colombia" pageId="9" pageNumber="266">COLOMBIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CA6E81FFC33C65C8696E3380ECF801" box="[872,967,1903,1929]" country="Colombia" name="Caqueta" pageId="9" pageNumber="266">Caquetá</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D0D8EFFFC33C65C8D56E32876AF800" box="[980,1089,1902,1928]" pageId="9" pageNumber="266">Florencia</collectingCounty>
|
||
, Vda. [
|
||
<collectingMunicipality id="6BD53A19FFC33C65CF936E3287CDF800" box="[1170,1254,1902,1928]" pageId="9" pageNumber="266">Vereda</collectingMunicipality>
|
||
]
|
||
<location id="8ED1F6B8FFC33C65CFF46E3386B3F800" LSID="urn:lsid:plazi:treatment:03A71175FFC23C66CB966B37825DFEC8:8ED1F6B8FFC33C65CFF46E3386B3F800" box="[1269,1432,1902,1929]" country="Colombia" county="Florencia" latitude="1.7066667" longLatPrecision="21" longitude="-75.61" municipality="Vereda" name="San Francisco" pageId="9" pageNumber="266" stateProvince="Caqueta">San Francisco</location>
|
||
, Fca. [
|
||
<location id="8ED1F6B8FFC33C65CBD46ECE8232F824" LSID="urn:lsid:plazi:treatment:03A71175FFC23C66CB966B37825DFEC8:8ED1F6B8FFC33C65CBD46ECE8232F824" box="[213,281,1938,1964]" country="Colombia" county="Florencia" latitude="1.7066667" longLatPrecision="21" longitude="-75.61" municipality="Vereda" name="Finca" pageId="9" pageNumber="266" stateProvince="Caqueta">Finca</location>
|
||
]
|
||
<location id="8ED1F6B8FFC33C65CA276ECE82B2F825" LSID="urn:lsid:plazi:treatment:03A71175FFC23C66CB966B37825DFEC8:8ED1F6B8FFC33C65CA276ECE82B2F825" box="[294,409,1938,1965]" country="Colombia" county="Florencia" latitude="1.7066667" longLatPrecision="21" longitude="-75.61" municipality="Vereda" name="El Recreo" pageId="9" pageNumber="266" stateProvince="Caqueta">El Recreo</location>
|
||
,
|
||
<geoCoordinate id="EE3AC6A4FFC33C65CAA46ECE8106F825" box="[421,557,1938,1965]" degrees="01" direction="north" minutes="42" orientation="latitude" pageId="9" pageNumber="266" precision="15" seconds="24" value="1.7066667">01º42’24’’N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE3AC6A4FFC33C65C9366ECE81E3F825" box="[567,712,1938,1965]" degrees="75" direction="west" minutes="36" orientation="longitude" pageId="9" pageNumber="266" precision="15" seconds="36" value="-75.61">75º36’36’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CF60D86FFC33C65C9D56ECE8035F825" box="[724,798,1938,1965]" metricMagnitude="2" metricUnit="m" metricValue="6.43" pageId="9" pageNumber="266" unit="m" value="643.0">
|
||
<elevation id="00234750FFC33C65C9D56ECE8035F825" box="[724,798,1938,1965]" metricMagnitude="2" metricUnit="m" metricValue="6.43" pageId="9" pageNumber="266" unit="m" value="643.0">643 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB086A3FFC33C65C82B6ECE80F9F825" box="[810,978,1938,1965]" pageId="9" pageNumber="266" value="2017-03-01" valueMax="2017-03-15" valueMin="2017-03-01">
|
||
<collectingDate id="EFF47F4BFFC33C65C82B6ECE80F9F825" box="[810,978,1938,1965]" pageId="9" pageNumber="266" value="2017-03-01" valueMax="2017-03-15" valueMin="2017-03-01">01–15.III.2017</collectingDate>
|
||
</date>
|
||
/
|
||
<location id="8ED1F6B8FFC33C65C8E96ECF8637F825" LSID="urn:lsid:plazi:treatment:03A71175FFC23C66CB966B37825DFEC8:8ED1F6B8FFC33C65C8E96ECF8637F825" box="[1000,1308,1938,1965]" country="Colombia" county="Florencia" latitude="1.7066667" longLatPrecision="21" longitude="-75.61" municipality="Vereda" name="Trampa Malaise en Bosque" pageId="9" pageNumber="266" stateProvince="Caqueta">Trampa Malaise en Bosque</location>
|
||
(Dosel),
|
||
<collectorName id="26FBC5B5FFC33C65CE846ECF82A3F859" pageId="9" pageNumber="266">Y. Ramos-Pastrana Leg.</collectorName>
|
||
” (
|
||
<collectionCode id="ED1F38A6FFC33C65CAA16EEB82C5F859" box="[416,494,1975,2001]" pageId="9" pageNumber="266">LEUA</collectionCode>
|
||
) (photographed specimen)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54B51EC1FFC33C65C8246EEA80A6F858" box="[805,909,1974,2000]" pageId="9" pageNumber="266" type="holotype">Holotype</typeStatus>
|
||
condition: good, abdomen dissected and stored in microvial with glycerin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFC03C66CBC669CA800DFF38" box="[199,806,150,176]" pageId="10" pageNumber="267" type="distribution">
|
||
<paragraph id="8BB1A063FFC03C66CBC669CA800DFF38" blockId="10.[151,1436,150,321]" box="[199,806,150,176]" pageId="10" pageNumber="267">
|
||
<emphasis id="B97A7C71FFC03C66CBC669CA824AFF38" bold="true" box="[199,353,150,176]" pageId="10" pageNumber="267">Distribution.</emphasis>
|
||
<collectingCountry id="F319E0F3FFC03C66CA6669CA82FEFF38" box="[359,469,150,176]" name="Colombia" pageId="10" pageNumber="267">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CA6E81FFC03C66CAE469CA816CFF38" box="[485,583,150,176]" country="Colombia" name="Caqueta" pageId="10" pageNumber="267">Caquetá</collectingRegion>
|
||
: Florencia) (
|
||
<figureCitation id="1335BCE6FFC03C66C9D269CA8033FF38" box="[723,792,150,176]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[301,1286,1429,1917]" captionTargetId="figure-56@13.[299,1288,1427,1919]" captionTargetPageId="13" captionText="FIGURE 7. Geographical records of Systenus Loew in Colombia. S. beatae Naglis, 2000; S. bickeli sp. nov.; S. colombiensis sp. nov.; S. flaviatus Naglis, 2000; S. fuscus sp. nov.; S. grossus sp. nov.; S. maculipennis Bickel, 2015; S. naranjensis Bickel, 2015 and S. parkeri Bickel, 2015." figureDoi="http://doi.org/10.5281/zenodo.13849845" httpUri="https://zenodo.org/record/13849845/files/figure.png" pageId="10" pageNumber="267">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFC03C66CBC669E687DFFF70" pageId="10" pageNumber="267" type="discussion">
|
||
<paragraph id="8BB1A063FFC03C66CBC669E687DFFF70" blockId="10.[151,1436,150,321]" pageId="10" pageNumber="267">
|
||
<emphasis id="B97A7C71FFC03C66CBC669E68210FF5C" bold="true" box="[199,315,186,212]" pageId="10" pageNumber="267">Remarks.</emphasis>
|
||
The new species is easily distinguished from congeners by the remarkably wide surstylus (
|
||
<figureCitation id="1335BCE6FFC03C66CE4169E686BEFF5C" box="[1344,1429,186,212]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="10" pageNumber="267">Fig. 4F</figureCitation>
|
||
) (visible without dissection) and by the digitiform process at basal 1/3 of the inner surface (
|
||
<figureCitation id="1335BCE6FFC03C66CF8C698287C3FF70" box="[1165,1256,222,248]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="10" pageNumber="267">Fig. 4G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFC03C66CBC6685E825DFEC8" pageId="10" pageNumber="267" type="etymology">
|
||
<paragraph id="8BB1A063FFC03C66CBC6685E825DFEC8" blockId="10.[151,1436,150,321]" pageId="10" pageNumber="267">
|
||
<emphasis id="B97A7C71FFC03C66CBC6685E8265FE94" bold="true" box="[199,334,258,284]" pageId="10" pageNumber="267">Etymology.</emphasis>
|
||
From Latin
|
||
<taxonomicName id="4C0EDBE0FFC03C66CADD685E8111FE94" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[476,570,258,284]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="grossus">
|
||
‘
|
||
<emphasis id="B97A7C71FFC03C66CAE4685F8111FE94" box="[485,570,259,284]" italics="true" pageId="10" pageNumber="267">grossus</emphasis>
|
||
</taxonomicName>
|
||
’ (= thickness), referring to the shape of the surstylus. Species epithet is to be treated as adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |