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<document id="4968B8581CC544BC5EB7B6ACD0CD5348" ID-CLB-Dataset="50968" ID-DOI="10.5281/zenodo.273289" ID-GBIF-Dataset="4397fa5a-9228-4234-ac2f-4f20d3548456" ID-ISSN="1175­5326" ID-Zenodo-Dep="273289" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459746611236" checkinUser="plazi" docAuthor="Binda, Maria Grazia, Pilato, Giovanni &amp; Lisi, Oscar" docDate="2005" docId="6656879B9668FF97FEEFA93F181BFB6E" docLanguage="en" docName="zt01075p068.pdf" docOrigin="Zootaxa 1075" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrobiotus sicheli Binda, Pilato &amp; Lisi, 2005, sp. nov." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="9A6FFFE39662FF9BFFE7AB0B1C39FFA0" masterDocTitle="Remarks on Macrobiotus furciger Murray, 1906 and description of three new species of the furciger group (Eutardigrada, Macrobiotidae)" masterLastPageNumber="68" masterPageNumber="55" pageNumber="65" updateTime="1698213948434" updateUser="ExternalLinkService">
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<mods:title id="3549399037A20F95C1C882CC2E9C6153">Remarks on Macrobiotus furciger Murray, 1906 and description of three new species of the furciger group (Eutardigrada, Macrobiotidae)</mods:title>
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<mods:namePart id="7F458917CCA39EB9E54EFD46A1F7A943">Binda, Maria Grazia</mods:namePart>
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<mods:namePart id="BA116DB796401D01E950165AC47B4EA3">Pilato, Giovanni</mods:namePart>
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<mods:namePart id="9CC96AAF4F3D98C6D85358700D587944">Lisi, Oscar</mods:namePart>
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<mods:title id="2D5A9A2F96DEC9C11181F8279138DF1F">Zootaxa</mods:title>
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<mods:part id="93E5BE3CCC3A4F47E9ECF3F2B9BC59D5">
<mods:date id="1C4C4762858E405220CC28C38F44FCD5">2005</mods:date>
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<treatment id="6656879B9668FF97FEEFA93F181BFB6E" ID-DOI="http://doi.org/10.5281/zenodo.5669893" ID-GBIF-Taxon="119336538" ID-Zenodo-Dep="5669893" LSID="urn:lsid:plazi:treatment:6656879B9668FF97FEEFA93F181BFB6E" httpUri="http://treatment.plazi.org/id/6656879B9668FF97FEEFA93F181BFB6E" lastPageId="12" lastPageNumber="67" pageId="10" pageNumber="65">
<subSubSection id="A6E565069668FF91FEEFA93F1E85FDED" box="[264,700,563,590]" pageId="10" pageNumber="65" type="nomenclature">
<paragraph id="EE40368D9668FF91FEEFA93F1E85FDED" blockId="10.[264,700,563,590]" box="[264,700,563,590]" pageId="10" pageNumber="65">
<heading id="B50881E19668FF91FEEFA93F1E85FDED" bold="true" box="[264,700,563,590]" fontSize="11" level="1" pageId="10" pageNumber="65" reason="1">
<emphasis id="DC8BEA9F9668FF91FEEFA93F1E85FDED" bold="true" box="[264,700,563,590]" pageId="10" pageNumber="65">
<taxonomicName id="29FF4D0E9668FF91FEEFA93F1DCEFDEE" ID-CoL="3X2WB" box="[264,503,564,590]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="65" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F9668FF91FEEFA93F1DCEFDEE" bold="true" box="[264,503,564,590]" italics="true" pageId="10" pageNumber="65">Macrobiotus sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E49668FF91FE19A9381E62FDED" box="[510,603,563,589]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="76C42A089668FF91FD8CA9381E8AFDED" box="[619,691,563,589]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A6E565069668FF97FEEFA974181BFB6E" lastPageId="12" lastPageNumber="67" pageId="10" pageNumber="65" type="materials_examined">
<paragraph id="EE40368D9668FF91FEEFA974185DFD39" blockId="10.[264,1324,639,1905]" box="[264,1124,639,665]" pageId="10" pageNumber="65">
<typeStatus id="3144882F9668FF91FEEFA9741D7BFD39" box="[264,322,639,665]" pageId="10" pageNumber="65">Type</typeStatus>
locality.
<collectingCountry id="96E8761D9668FF91FE49A9741E71FD39" box="[430,584,639,665]" name="South Africa" pageId="10" pageNumber="65">South Africa</collectingCountry>
, Tzitzikama (an undetermined moss sample).
</paragraph>
<paragraph id="EE40368D9668FF91FEDFA9AC18CBFD61" blockId="10.[264,1324,639,1905]" box="[312,1266,679,705]" pageId="10" pageNumber="65">
Material examined.
<collectingCountry id="96E8761D9668FF91FDC4A9AC1E84FD61" box="[547,701,679,705]" name="South Africa" pageId="10" pageNumber="65">South Africa</collectingCountry>
, Tzitzikama:
<typeStatus id="3144882F9668FF91FCBBA9AC1FFFFD61" box="[860,966,679,705]" pageId="10" pageNumber="65" type="holotype">holotype</typeStatus>
, 29
<typeStatus id="3144882F9668FF91FC11A9AC185FFD61" box="[1014,1126,679,705]" pageId="10" pageNumber="65" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F8F9FD049668FF91FB78A9AC18D7FD61" box="[1183,1262,679,705]" pageId="10" pageNumber="65" type="egg">7 eggs</specimenCount>
.
</paragraph>
<paragraph id="EE40368D9668FF91FEDFA9C41F2AFCB1" blockId="10.[264,1324,639,1905]" pageId="10" pageNumber="65">
<typeStatus id="3144882F9668FF91FEDFA9C41D4BFD49" box="[312,370,719,745]" pageId="10" pageNumber="65">Type</typeStatus>
repository: Collection of Binda and Pilato: Department of Animal Biology “Marcello La Greca”, University of Catania.
</paragraph>
<paragraph id="EE40368D9668FF91FEDFA8141D87FBA1" blockId="10.[264,1324,639,1905]" pageId="10" pageNumber="65">
<bibRefCitation id="8A6E4B7C9668FF91FEDFA8141DECFC99" author="Binda" box="[312,469,799,825]" pageId="10" pageNumber="67" refString="Binda, M. G. (1984) Notizie sui Tardigradi dell'Africa meridionale con descrizione di una nuova specie di Apodibius (Eutardigrada). Animalia, 11, 5 - 15." type="journal article" year="1984">Binda (1984)</bibRefCitation>
attributed to
<taxonomicName id="29FF4D0E9668FF91FD9EA8141EC6FC98" box="[633,767,799,824]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="65" phylum="Tardigrada" rank="species" species="furciger">
<emphasis id="DC8BEA9F9668FF91FD9EA8141EC6FC98" box="[633,767,799,824]" italics="true" pageId="10" pageNumber="65">M. furciger</emphasis>
</taxonomicName>
specimens and eggs from
<collectingCountry id="96E8761D9668FF91FBA5A81418E6FC99" box="[1090,1247,799,825]" name="South Africa" pageId="10" pageNumber="65">South Africa</collectingCountry>
; some differences with respect to the description of this species were attributed to the individual variability. We compared the material from
<collectingCountry id="96E8761D9668FF91FCFDA8641F8DFC29" box="[794,948,879,905]" name="South Africa" pageId="10" pageNumber="65">South Africa</collectingCountry>
with specimens of
<taxonomicName id="29FF4D0E9668FF91FB44A8641912FC28" box="[1187,1323,879,904]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="65" phylum="Tardigrada" rank="species" species="furciger">
<emphasis id="DC8BEA9F9668FF91FB44A8641912FC28" box="[1187,1323,879,904]" italics="true" pageId="10" pageNumber="65">M. furciger</emphasis>
</taxonomicName>
from the
<collectingCountry id="96E8761D9668FF91FE9CA89C1E11FC11" box="[379,552,919,945]" name="South Georgia and the South Sandwich Islands" pageId="10" pageNumber="65">South Georgia</collectingCountry>
described above, and noted some differences that induce us to attribute the South African specimens to a new species here described and named
<taxonomicName id="29FF4D0E9668FF91FAECA8B41D6DFBA0" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="65" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F9668FF91FAECA8B41D6DFBA0" italics="true" pageId="10" pageNumber="65">M. sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E49668FF91FE84A8EC1D87FBA1" box="[355,446,999,1025]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="EE40368D9668FF91FEDFAF041F34FA19" blockId="10.[264,1324,639,1905]" pageId="10" pageNumber="65">
Description of the
<typeStatus id="3144882F9668FF91FDF8AF041EB0FB89" box="[543,649,1039,1065]" pageId="10" pageNumber="65" type="holotype">holotype</typeStatus>
. Body length
<quantity id="29079B689668FF91FCDEAF041FA2FB89" box="[825,923,1039,1065]" metricMagnitude="2" metricUnit="m" metricValue="3.29" pageId="10" pageNumber="65" unit="m" value="329.0">329 m</quantity>
; colourless, eyes present, cuticle with small dots difficult to see (
<figureCitation id="76C42A089668FF91FD9EAF3C1E86FBF1" box="[633,703,1079,1105]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
C). Buccopharyngeal apparatus of
<taxonomicName id="29FF4D0E9668FF91FBBBAF3C18C8FBF0" box="[1116,1265,1079,1104]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="65" phylum="Tardigrada" rank="genus">
<emphasis id="DC8BEA9F9668FF91FBBBAF3C18C8FBF0" box="[1116,1265,1079,1104]" italics="true" pageId="10" pageNumber="65">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="3144882F9668FF91FB1FAF3C1912FBF1" box="[1272,1323,1079,1105]" pageId="10" pageNumber="65">type</typeStatus>
(
<figureCitation id="76C42A089668FF91FEF6AF541D64FBD9" box="[273,349,1119,1145]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
A); mouth terminal with ten peribuccal lamellae; buccal cavity wall with an anterior band of fine teeth, a posterior crown of triangular teeth and three dorsal and three ventral transverse ridges. Also some supplementary teeth present. Buccal tube
<quantity id="29079B689668FF91FB22AFA41912FB69" box="[1221,1323,1199,1225]" metricMagnitude="1" metricUnit="m" metricValue="4.54" pageId="10" pageNumber="65" unit="m" value="45.4">45.4 m</quantity>
long and
<quantity id="29079B689668FF91FE93AFDC1DF1FB51" box="[372,456,1239,1265]" metricMagnitude="0" metricUnit="m" metricValue="8.3" pageId="10" pageNumber="65" unit="m" value="8.3">8.3 m</quantity>
wide (
<emphasis id="DC8BEA9F9668FF91FDFEAFDC1E09FB50" box="[537,560,1239,1264]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 18.3). Stylet supports inserted on the buccal tube at 80 % of its length (
<emphasis id="DC8BEA9F9668FF91FE84AFF41D43FAB8" box="[355,378,1279,1304]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 80). The pharyngeal bulb (
<quantity id="29079B689668FF91FD37AFF41F0DFAB9" box="[720,820,1279,1305]" metricMagnitude="1" metricUnit="m" metricValue="4.63" pageId="10" pageNumber="65" unit="m" value="46.3">46.3 m</quantity>
<date id="9A41104D9668FF91FCDBAFF41F56FAB9" box="[828,879,1279,1305]" pageId="10" pageNumber="65" value="1932-10">
x
<quantity id="29079B689668FF91FCB5AFF41F56FAB9" box="[850,879,1279,1305]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="65" unit="m" value="32.0">32</quantity>
</date>
m) has pharyngeal apophyses, three rod­shaped macroplacoids and microplacoid. First macroplacoid
<quantity id="29079B689668FF91FBEFAE2C1866FAE1" box="[1032,1119,1319,1345]" metricMagnitude="0" metricUnit="m" metricValue="7.4" pageId="10" pageNumber="65" unit="m" value="7.4">7.4 m</quantity>
long (
<emphasis id="DC8BEA9F9668FF91FB49AE2C18FCFAE0" box="[1198,1221,1319,1344]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 16.3), second
<quantity id="29079B689668FF91FE81AE441DF8FAC9" box="[358,449,1359,1385]" metricMagnitude="0" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="65" unit="m" value="6.1">6.1 m</quantity>
(
<emphasis id="DC8BEA9F9668FF91FE30AE441DD7FAC8" box="[471,494,1359,1384]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 13.4), third, with a preterminal narrowing,
<quantity id="29079B689668FF91FBD4AE4418B4FAC9" box="[1075,1165,1359,1385]" metricMagnitude="0" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="65" unit="m" value="7.3">7.3 m</quantity>
(
<emphasis id="DC8BEA9F9668FF91FB43AE441882FAC8" box="[1188,1211,1359,1384]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 16.1), microplacoid
<quantity id="29079B689668FF91FE57AE7C1E33FA31" box="[432,522,1399,1425]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="10" pageNumber="65" unit="m" value="5.2">5.2 m</quantity>
long (
<emphasis id="DC8BEA9F9668FF91FD85AE7C1E40FA30" box="[610,633,1399,1424]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 11.4); entire placoid row
<quantity id="29079B689668FF91FC04AE7C1875FA31" box="[995,1100,1399,1425]" metricMagnitude="1" metricUnit="m" metricValue="2.77" pageId="10" pageNumber="65" unit="m" value="27.7">27.7 m</quantity>
long (
<emphasis id="DC8BEA9F9668FF91FB43AE7C1882FA30" box="[1188,1211,1399,1424]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 61.0), macroplacoid row
<quantity id="29079B689668FF91FE04AE941E7FFA19" box="[483,582,1439,1465]" metricMagnitude="1" metricUnit="m" metricValue="2.18" pageId="10" pageNumber="65" unit="m" value="21.8">21.8 m</quantity>
long (
<emphasis id="DC8BEA9F9668FF91FD74AE941E93FA18" box="[659,682,1439,1464]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 48.0).
</paragraph>
<paragraph id="EE40368D9668FF91FEDFAECC1FBDF921" blockId="10.[264,1324,639,1905]" pageId="10" pageNumber="65">
The claws, of the
<emphasis id="DC8BEA9F9668FF91FDF1AECC1EBDFA40" box="[534,644,1479,1504]" italics="true" pageId="10" pageNumber="65">hufelandi</emphasis>
<typeStatus id="3144882F9668FF91FD68AECC1EFDFA41" box="[655,708,1479,1505]" pageId="10" pageNumber="65">type</typeStatus>
, are short with accessory points on main branches (
<figureCitation id="76C42A089668FF91FEF6AEE41D6EF9A9" box="[273,343,1519,1545]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
B). External claws of third pair of legs
<quantity id="29079B689668FF91FCD6AEE41FBEF9A9" box="[817,903,1519,1545]" metricMagnitude="0" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="65" unit="m" value="9.5">9.5 m</quantity>
long (
<emphasis id="DC8BEA9F9668FF91FC32AEE41FD5F9A8" box="[981,1004,1519,1544]" italics="true" pageId="10" pageNumber="65">pt</emphasis>
= 20.9). The unfavourable orientation does not allow measurement of other claws; in
<tableCitation id="A37D03369668FF91FC5FAD1C1836F991" box="[952,1039,1559,1585]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="10" pageNumber="65" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
the length of claws of a
<typeStatus id="3144882F9668FF91FEEFAD341D54F9F9" box="[264,365,1599,1625]" pageId="10" pageNumber="65" type="paratype">paratype</typeStatus>
<quantity id="29079B689668FF91FE91AD341DEDF9F9" box="[374,468,1599,1625]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="65" unit="m" value="230.0">230 m</quantity>
long is indicated. Smooth lunules are present; a cuticular thickening is present near the lunules on the first three pairs of legs.
</paragraph>
<paragraph id="EE40368D9668FF90FEDFAD841823FE0E" blockId="10.[264,1324,639,1905]" lastBlockId="11.[264,1323,284,430]" lastPageId="11" lastPageNumber="66" pageId="10" pageNumber="65">
The eggs (
<figureCitation id="76C42A089668FF91FE56AD841DCEF909" box="[433,503,1679,1705]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
D, E), freely laid, are spherical with trunco­conical processes with the extremity divided (the division may be dichotomic or multiple); the diameter is
<quantity id="29079B689668FF91FB29ADBC1D09F959" metricMagnitude="1" metricUnit="m" metricValue="7.575" metricValueMax="8.55" metricValueMin="6.6" pageId="10" pageNumber="65" unit="m" value="75.75" valueMax="85.5" valueMin="66.0">6685.5 m</quantity>
excluding the processes,
<quantity id="29079B689668FF91FD90ADD41ECAF959" box="[631,755,1759,1785]" metricMagnitude="1" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="10" pageNumber="65" unit="m" value="88.5" valueMax="95.0" valueMin="82.0">8295 m</quantity>
including them. The processes,
<quantity id="29079B689668FF91FB69ADD418C0F959" box="[1166,1273,1759,1785]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.842" metricValueMin="3.81" pageId="10" pageNumber="65" unit="in" value="19.0" valueMax="23.0" valueMin="15.0">1523 in</quantity>
the circumference, have a large basal portion and a narrower terminal portion divided twice or thrice (more rarely once) at the apex; the branches are therefore short, and the terminal branches very short (
<figureCitation id="76C42A089668FF91FDEBAC5C1E6EF8D1" box="[524,599,1879,1905]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="10" pageNumber="65">Fig. 5</figureCitation>
D). Process surface smooth without refracting areas. The processes in the largest measured egg are
<quantity id="29079B689669FF90FD11AA171F42FE96" box="[758,891,284,310]" metricMagnitude="0" metricUnit="m" metricValue="7.15" metricValueMax="7.8" metricValueMin="6.5" pageId="11" pageNumber="66" unit="m" value="7.15" valueMax="7.8" valueMin="6.5">6.57.8 m</quantity>
high, the basal diameter in the same egg is
<quantity id="29079B689669FF90FEB0AA4F1DD7FEFE" box="[343,494,324,350]" metricMagnitude="1" metricUnit="m" metricValue="1.005" metricValueMax="1.22" metricValueMin="0.79" pageId="11" pageNumber="66" unit="m" value="10.05" valueMax="12.2" valueMin="7.9">7.912.2 m</quantity>
.. The base of each process has a crown of projections forming a very evident crown of dots. The egg shell is dotted; some dots may be contiguous to one another and form a trace of an irregular reticular design (
<figureCitation id="76C42A089669FF90FC4DAA9F1FC9FE0E" box="[938,1008,404,430]" captionStart="FIGURE 5" captionStartId="11.[264,368,1591,1615]" captionTargetBox="[264,1316,473,1553]" captionTargetId="figure@11.[264,1323,470,1564]" captionTargetPageId="11" captionText="FIGURE 5. Macrobiotus sicheli sp. nov.: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="11" pageNumber="66">Fig. 5</figureCitation>
E).
</paragraph>
<caption id="BA8066059669FF90FEEFAD3C1FF2F933" httpUri="https://zenodo.org/record/273294/files/figure.png" pageId="11" pageNumber="66" targetBox="[264,1316,473,1553]" targetPageId="11">
<paragraph id="EE40368D9669FF90FEEFAD3C1FF2F933" blockId="11.[264,1323,1591,1683]" pageId="11" pageNumber="66">
<emphasis id="DC8BEA9F9669FF90FEEFAD3C1DB3F9EF" bold="true" box="[264,394,1591,1615]" pageId="11" pageNumber="66">FIGURE 5.</emphasis>
<taxonomicName id="29FF4D0E9669FF90FE75AD331E55F9EF" box="[402,620,1592,1615]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="66" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F9669FF90FE75AD331E55F9EF" box="[402,620,1592,1615]" italics="true" pageId="11" pageNumber="66">Macrobiotus sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E49669FF90FD94AD3C1EF3F9EE" box="[627,714,1591,1614]" pageId="11" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
: A, buccopharyngeal apparatus of the holotype; B, claws of the third pair of legs; C, cuticular dots, and claws of the fourth pair of legs of the holotype; D, egg; E, detail of the egg shell ornamentation. (Scale bar = 10 m)
</paragraph>
</caption>
<paragraph id="EE40368D9669FF90FEDFADCC1ED9F8A9" blockId="11.[264,1323,1735,1881]" pageId="11" pageNumber="66">
Etymology. The name
<taxonomicName id="29FF4D0E9669FF90FDA1ADC31EAAF941" box="[582,659,1736,1761]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="66" phylum="Tardigrada" rank="species" species="sicheli">
<emphasis id="DC8BEA9F9669FF90FDA1ADC31EAAF941" box="[582,659,1736,1761]" italics="true" pageId="11" pageNumber="66">sicheli</emphasis>
</taxonomicName>
is in honour of Prof. Giovanni Sichel, cellular biologist of University of Catania and our friend.
</paragraph>
<paragraph id="EE40368D9669FF97FEDFAC1C1858FE5E" blockId="11.[264,1323,1735,1881]" lastBlockId="12.[264,1323,284,1230]" lastPageId="12" lastPageNumber="67" pageId="11" pageNumber="66">
Remarks.
<taxonomicName id="29FF4D0E9669FF90FE51AC131E9FF891" box="[438,678,1816,1841]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="66" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F9669FF90FE51AC131E9FF891" box="[438,678,1816,1841]" italics="true" pageId="11" pageNumber="66">Macrobiotus sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E49669FF90FD53AC1C1F2FF891" box="[692,790,1815,1841]" pageId="11" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="29FF4D0E9669FF90FC21AC13186BF891" box="[966,1106,1816,1841]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="66" phylum="Tardigrada" rank="species" species="furciger">
<emphasis id="DC8BEA9F9669FF90FC21AC13186BF891" box="[966,1106,1816,1841]" italics="true" pageId="11" pageNumber="66">M. furciger</emphasis>
</taxonomicName>
in the following features: cuticle finely dotted; wider buccal tube (
<tableCitation id="A37D03369669FF90FCB3AC341F94F8F9" box="[852,941,1855,1881]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="11" pageNumber="66" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
); stylet supports inserted on the buccal tube in a more caudal position (
<emphasis id="DC8BEA9F966EFF97FD3DAA171EC9FE95" box="[730,752,284,309]" italics="true" pageId="12" pageNumber="67">pt</emphasis>
= 79.380.0 in
<taxonomicName id="29FF4D0E966EFF97FC49AA17181DFE95" box="[942,1060,284,309]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FC49AA17181DFE95" box="[942,1060,284,309]" italics="true" pageId="12" pageNumber="67">M. sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FBCBAA1718B2FE96" box="[1068,1163,284,310]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
,
<quantity id="29079B68966EFF97FB7FAA171912FE96" box="[1176,1323,284,310]" metricMagnitude="0" metricUnit="m" metricValue="1.94691" metricValueMax="1.9583399999999997" metricValueMin="1.93548" pageId="12" pageNumber="67" unit="in" value="76.65" valueMax="77.1" valueMin="76.2">76.277.1 in</quantity>
<taxonomicName id="29FF4D0E966EFF97FEEFAA4F1DB5FEFD" box="[264,396,324,349]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="furciger">
<emphasis id="DC8BEA9F966EFF97FEEFAA4F1DB5FEFD" box="[264,396,324,349]" italics="true" pageId="12" pageNumber="67">M. furciger</emphasis>
</taxonomicName>
); third macroplacoid about long as the first one (
<tableCitation id="A37D0336966EFF97FC2CAA4F181AFEFE" box="[971,1059,324,350]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
); claws shorter (
<tableCitation id="A37D0336966EFF97FB0FAA4F1D2FFE26" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
); accessory points less developed; and in some characters of the egg (smaller dimensions; processes shorter, without refracting areas, divided only in the terminal portion and therefore with branches shorter; basal crown of projections more evident; egg shell with more evident dots but without a true, regular, reticular design).
</paragraph>
<paragraph id="EE40368D966EFF97FEDFA9071E17FCC6" blockId="12.[264,1323,284,1230]" pageId="12" pageNumber="67">
The new species differs from
<taxonomicName id="29FF4D0E966EFF97FD40A9071F12FD85" box="[679,811,524,550]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="aradasi" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FD40A9071E86FD85" box="[679,703,524,549]" italics="true" pageId="12" pageNumber="67">M</emphasis>
.
<emphasis id="DC8BEA9F966EFF97FD36A9071F12FD85" box="[721,811,524,549]" italics="true" pageId="12" pageNumber="67">aradasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FCD1A9071FADFD86" box="[822,916,524,550]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
in the following features: cuticle finely dotted; buccal armature with anterior band of teeth present; posterior crown of teeth more developed; wider buccal tube (
<tableCitation id="A37D0336966EFF97FD55A9571F30FDD6" box="[690,777,604,630]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
); stylet supports inserted on the buccal tube in more caudal position (
<emphasis id="DC8BEA9F966EFF97FDFBA98F1E0AFD3D" box="[540,563,644,669]" italics="true" pageId="12" pageNumber="67">pt</emphasis>
= 79.380.0 in
<taxonomicName id="29FF4D0E966EFF97FD17A98F1F5FFD3D" box="[752,870,644,670]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FD17A98F1F31FD3D" box="[752,776,644,669]" italics="true" pageId="12" pageNumber="67">M</emphasis>
.
<emphasis id="DC8BEA9F966EFF97FCFEA98F1F5FFD3D" box="[793,870,644,669]" italics="true" pageId="12" pageNumber="67">sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FC89A98F1FF4FD3E" box="[878,973,644,670]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
, 72.0
<quantity id="29079B68966EFF97FBFDA98F184CFD3E" box="[1050,1141,644,670]" metricMagnitude="0" metricUnit="m" metricValue="1.8491199999999999" pageId="12" pageNumber="67" unit="in" value="72.8">72.8 in</quantity>
<taxonomicName id="29FF4D0E966EFF97FB99A98F1939FD3D" box="[1150,1280,644,670]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="aradasi" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FB99A98F18AFFD3D" box="[1150,1174,644,669]" italics="true" pageId="12" pageNumber="67">M</emphasis>
.
<emphasis id="DC8BEA9F966EFF97FB40A98F1939FD3D" box="[1191,1280,644,669]" italics="true" pageId="12" pageNumber="67">aradasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FAEEA98F1D07FD66" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
); microplacoid slightly longer (
<tableCitation id="A37D0336966EFF97FD4CA9A71F38FD66" box="[683,769,684,710]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
), claws shorter (
<tableCitation id="A37D0336966EFF97FC23A9A71823FD66" box="[964,1050,684,710]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
) with accessory points less developed; and in some characters of the eggs (egg processes divided only in the terminal portion and therefore with branches clearly shorter; basal projections forming a more evident basal crown of dots; egg shell with more evident dots but without a true, regular, reticular design).
</paragraph>
<paragraph id="EE40368D966EFF97FEDFA87F181BFB6E" blockId="12.[264,1323,284,1230]" pageId="12" pageNumber="67">
<taxonomicName id="29FF4D0E966EFF97FEDFA87F1E11FC2D" box="[312,552,884,909]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FEDFA87F1E11FC2D" box="[312,552,884,909]" italics="true" pageId="12" pageNumber="67">Macrobiotus sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FDD2A87F1EAEFC2E" box="[565,663,884,910]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="29FF4D0E966EFF97FCA2A87F1FDCFC2D" box="[837,997,884,909]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="divergens" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FCA2A87F1FDCFC2D" box="[837,997,884,909]" italics="true" pageId="12" pageNumber="67">M. divergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FC14A87F186DFC2E" box="[1011,1108,884,910]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
in the following features: cuticle finely dotted; eyes present; stylet sheaths normally shaped,
<emphasis id="DC8BEA9F966EFF97FB73A8971886FC15" box="[1172,1215,924,949]" italics="true" pageId="12" pageNumber="67">i.e.,</emphasis>
not very thick and with the caudal portion not diverging from the buccal tube walls; stylet supports inserted on the buccal tube in a more caudal position (
<emphasis id="DC8BEA9F966EFF97FC66A8E71FA1FBA5" box="[897,920,1004,1029]" italics="true" pageId="12" pageNumber="67">pt</emphasis>
= 79.380.0 in
<taxonomicName id="29FF4D0E966EFF97FBA8A8E718FBFBA5" box="[1103,1218,1004,1030]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="sicheli" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FBA8A8E7185EFBA5" box="[1103,1127,1004,1029]" italics="true" pageId="12" pageNumber="67">M</emphasis>
.
<emphasis id="DC8BEA9F966EFF97FB92A8E718FBFBA5" box="[1141,1218,1004,1029]" italics="true" pageId="12" pageNumber="67">sicheli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FB2EA8E7191FFBA6" box="[1225,1318,1004,1030]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
,
<quantity id="29079B68966EFF97FEEFAF1F1DA2FB8E" box="[264,411,1044,1070]" metricMagnitude="0" metricUnit="m" metricValue="1.93548" metricValueMax="1.9685" metricValueMin="1.90246" pageId="12" pageNumber="67" unit="in" value="76.2" valueMax="77.5" valueMin="74.9">74.977.5 in</quantity>
<taxonomicName id="29FF4D0E966EFF97FE43AF1F1E07FB8D" box="[420,574,1044,1070]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="67" phylum="Tardigrada" rank="species" species="divergens" status="sp. nov.">
<emphasis id="DC8BEA9F966EFF97FE43AF1F1D85FB8D" box="[420,444,1044,1069]" italics="true" pageId="12" pageNumber="67">M</emphasis>
.
<emphasis id="DC8BEA9F966EFF97FE29AF1F1E07FB8D" box="[462,574,1044,1069]" italics="true" pageId="12" pageNumber="67">divergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7B857E4966EFF97FDAFAF1F1E90FB8E" box="[584,681,1044,1070]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
); microplacoid more developed (
<tableCitation id="A37D0336966EFF97FBD3AF1F18B7FB8E" box="[1076,1166,1044,1070]" captionStart="TABLE 1" captionStartId="3.[264,352,643,667]" captionTargetBox="[264,1294,814,1896]" captionTargetPageId="3" captionText="TABLE 1. Dimensions of some structures of the smallest and the largest measured specimens of Macrobiotus furciger, M. aradasi sp. nov., M. divergens sp. nov. and M. sicheli sp. nov." httpUri="http://table.plazi.org/id/BA8066059661FF98FEEFA9881890FD62" pageId="12" pageNumber="67" tableUuid="BA8066059661FF98FEEFA9881890FD62">Table 1</tableCitation>
); claws with accessory points less developed; and in some characters of the eggs (processes more developed, divided only in the terminal portion and therefore with branches shorter; basal projections more developed and forming a more evident crown of basal dots; egg shell with more evident dots but without a true, regular, reticular design).
</paragraph>
</subSubSection>
</treatment>
</document>