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<mods:title id="B7AD295394D0163B57CDE691F270BD28">Extraordinary majoid crabs: the genus Esopus A. Milne-Edwards, 1875 in the new subfamily Esopinae subfam. nov., and erection of Paulitinae subfam. nov. (Crustacea, Decapoda, Brachyura, Majoidea, Inachoididae Dana, 1851)</mods:title>
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Stenorhynchinae
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Stenorhynchinae
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Stenorhynchinae
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Davie
<emphasis id="DCE4EAC2FFB8A61EB55CFC95FCA2FCCF" box="[819,871,863,887]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
2015b: 949
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genus
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.
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<emphasis id="DCE4EAC2FFB8A61EB75DFC31FE13FC66" box="[306,470,964,990]" italics="true" pageId="17" pageNumber="118">Stenorhynchus</emphasis>
Lamarck, 1815
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[
<typeStatus id="312B8872FFB8A61EB4F2FC30FD15FC67" box="[669,720,965,991]" pageId="17" pageNumber="118">type</typeStatus>
species by designation by
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:
<taxonomicName id="29904D53FFB8A61EB2BCFC30FEEBFBBB" authority="Herbst, 1788" authorityName="Herbst" authorityYear="1788" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="DCE4EAC2FFB8A61EB2BCFC30FA59FC66" box="[1235,1436,965,991]" italics="true" pageId="17" pageNumber="118">Cancer seticornis</emphasis>
Herbst, 1788
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].
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. Carapace longer than broad, narrow, not sculpted, smooth, naked or minutely pubescent. Rostrum unpaired, very long, longer than carapace, very slender, with spinous margins. Orbits completely absent; only small postocular spine, distant from unprotected eye, situated along long ‘neck’. Basal article of antenna slender, convex ventrally, not longitudinally sulcate; flagellum concealed beneath rostrum. Orbits not defined. Postorbital spine small. Proepistome not completely reaching short epistomial process. Epistome very large. Pleurites 5–8 exposed but relatively small (particularly pleurite 5), forming narrow, smooth, interrupted collar, never expanded nor salient. Carapace partially resting on weak setting gutter. First pleonal somite encroaching on concave posterior carapace margin; first, second pleonal somites integrated into cephalothorax of males; in adult females pleonal somites 1–4, portion of
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prolongation of carapace, dorsally visible. Thoracic sternal suture 3/4 only weakly visible laterally, not marked by deep depression. Sternum/pterygostome junction complete. Sternal extensions from P1–P4 connecting to sternum. Chelipeds long, narrow, cylindrical. P2–P5 long, spinous (
<bibRefCitation id="8A014B21FFB8A61EB586FA6DFB62FA0B" author="Hendrickx" box="[1001,1191,1432,1459]" firstAuthor="Hendrickx" pageId="17" pageNumber="118" refId="ref22525" refString="Hendrickx, M. E. (1999) Los Cangrejos Braquiuros (Crustacea: Brachyura: Majoidea y Parthenopoidea) del Pacifico Mexicano. Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO) e Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico City, 274 pp., 13 pls." type="book" year="1999">Hendrickx 1999</bibRefCitation>
: fig. 26B:
<taxonomicName id="29904D53FFB8A61EB34CFA6CFA49FA0A" baseAuthorityName="SI Smith" baseAuthorityYear="1871" box="[1315,1420,1432,1459]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="DCE4EAC2FFB8A61EB34CFA6CFA49FA0A" box="[1315,1420,1432,1459]" italics="true" pageId="17" pageNumber="118">S. debilis</emphasis>
</taxonomicName>
). Male pleon with 6 somites (pleotelson); first and seemingly second somites integrated into cephalothorax; female pleon with 5 somites, somites 1–4 and a portion of somite
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the prolongation of the carapace, visible dorsally. Male gonopore at extremity of coxo-sternal condyle (
<bibRefCitation id="8A014B21FFB8A61EB482F9F1FC45F9A7" author="Guinot" box="[749,896,1540,1567]" firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
;
<bibRefCitation id="8A014B21FFB8A61EB5E4F9F1FB9EF9A7" author="Guinot" box="[907,1115,1540,1567]" etAl="et al." firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot
<emphasis id="DCE4EAC2FFB8A61EB58FF9F0FBD4F9A6" box="[992,1041,1540,1566]" italics="true" pageId="17" pageNumber="118">et al</emphasis>
. 2013
</bibRefCitation>
: fig. 31D). G1 with lobe and aperture subdistal, with setae (
<bibRefCitation id="8A014B21FFB8A61EB781F9DDFD48F9FB" author="Brocchi" box="[494,653,1576,1603]" firstAuthor="Brocchi" pageId="17" pageNumber="118" pagination="1 - 131" refId="ref19837" refString="Brocchi, P. (1875) Recherches sur les organes genitaux males des Crustaces Decapodes. Annales de Sciences naturelles, Zoolo- gie, Series 6, 2, 1 - 131." type="journal article" year="1875">Brocchi 1875</bibRefCitation>
: pl. 19, fig. 17;
<bibRefCitation id="8A014B21FFB8A61EB52EF9DDFC13F9FB" author="Monod" box="[833,982,1576,1603]" firstAuthor="Monod" pageId="17" pageNumber="118" pagination="1 - 674" refId="ref24178" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique Noire, 45, 1 - 674." type="journal article" year="1956">Monod 1956</bibRefCitation>
: fig. 839;
<bibRefCitation id="8A014B21FFB8A61EB226F9DDFB08F9FB" author="Garth" box="[1097,1229,1576,1603]" firstAuthor="Garth" pageId="17" pageNumber="118" pagination="1 - 55" refId="ref21540" refString="Garth, J. S. (1958) Brachyura of the Pacific coast of America, Oxyrhyncha. Allan Hancock Pacific Expeditions, 21 (1 &amp; 2), i-xxii + 1 - 499 &amp; 501 - 854, pls. A-Z 4 + 1 - 55." type="book chapter" year="1958">Garth 1958</bibRefCitation>
: pl. B, fig. 7;
<bibRefCitation id="8A014B21FFB8A61EB305F9DDFED2F9DF" author="Williams" firstAuthor="Williams" pageId="17" pageNumber="118" refId="ref25881" refString="Williams, A. B. (1984) Shrimps, Lobsters, and Crabs of the Atlantic Coast of the Eastern United States, Maine to Florida. Smithsonian Institution Press, Washington, D. C., 550 pp." type="book" year="1984">Williams 1984</bibRefCitation>
: fig. 241o).
</paragraph>
</subSubSection>
<subSubSection id="A68A655BFFB8A61DB6A8F985FEFEFEFB" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118" type="discussion">
<paragraph id="EE2F36D0FFB8A61EB6A8F985FDE3F933" blockId="17.[151,1437,1072,2035]" box="[199,550,1648,1675]" pageId="17" pageNumber="118">
<emphasis id="DCE4EAC2FFB8A61EB6A8F985FE11F933" bold="true" box="[199,468,1648,1675]" pageId="17" pageNumber="118">Other included genera</emphasis>
. None.
</paragraph>
<paragraph id="EE2F36D0FFB8A61DB6A8F960FD4BFF44" blockId="17.[151,1437,1072,2035]" lastBlockId="18.[151,1437,153,323]" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118">
<emphasis id="DCE4EAC2FFB8A61EB6A8F960FEF1F917" bold="true" box="[199,308,1685,1711]" pageId="17" pageNumber="118">Remarks</emphasis>
.
<taxonomicName id="29904D53FFB8A61EB72AF961FE2CF916" authorityName="Lamarck" authorityYear="1815" box="[325,489,1684,1710]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB8A61EB72AF961FE2CF916" box="[325,489,1684,1710]" italics="true" pageId="17" pageNumber="118">Stenorhynchus</emphasis>
</taxonomicName>
occupies a quite distinct position among the
<taxonomicName id="29904D53FFB8A61EB591F961FB4AF916" authorityName="Drach &amp; Guinot" authorityYear="1983" box="[1022,1167,1684,1710]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="family">Inachoididae</taxonomicName>
and therefore deserves a special status in the family (
<bibRefCitation id="8A014B21FFB8A61EB780F94DFD6AF96B" author="Guinot" box="[495,687,1720,1747]" firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012: 31</bibRefCitation>
;
<bibRefCitation id="8A014B21FFB8A61EB4D2F94DFC08F96B" author="Guinot" box="[701,973,1720,1747]" etAl="et al." firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot
<emphasis id="DCE4EAC2FFB8A61EB57BF94CFC82F96A" box="[788,839,1720,1746]" italics="true" pageId="17" pageNumber="118">et al</emphasis>
. 2013: 225
</bibRefCitation>
). The exposed pleurites do not show the continuous collar as in other inachoidids, the exposed pleurites appearing more as sclerites as in
<taxonomicName id="29904D53FFB8A61EB29BF929FAA0F94E" authorityName="MacLeay" authorityYear="1838" box="[1268,1381,1756,1782]" class="Malacostraca" family="Inachidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="family">Inachidae</taxonomicName>
. The expansion of the pleurites seems already present in the first crab stage of
<taxonomicName id="29904D53FFB8A61EB5BDF8F4FB9EF8A2" authorityName="Herbst" authorityYear="1788" box="[978,1115,1793,1819]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="DCE4EAC2FFB8A61EB5BDF8F4FB9EF8A2" box="[978,1115,1793,1819]" italics="true" pageId="17" pageNumber="118">S. seticornis</emphasis>
</taxonomicName>
(
<bibRefCitation id="8A014B21FFB8A61EB205F8F4FB2FF8A2" author="Yang" box="[1130,1258,1792,1819]" firstAuthor="Yang" pageId="17" pageNumber="118" pagination="157 - 177" refId="ref26018" refString="Yang, W. T. (1976) Studies on the western Atlantic arrow crab genus Stenorhynchus (Decapod Brachyura, Majidae) I. Larval characters of two species and comparison with other larvae of Inachinae. Crustaceana, 31 (2), 157 - 177. https: // doi. org / 10.1163 / 156854076 X 00198" type="journal article" year="1976">Yang 1976</bibRefCitation>
: fig. 7A). Complete larval development of
<taxonomicName id="29904D53FFB8A61EB7BCF8D0FDB5F886" baseAuthorityName="Brulle" baseAuthorityYear="1837" box="[467,624,1828,1855]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="lanceolatus">
<emphasis id="DCE4EAC2FFB8A61EB7BCF8D0FDB5F886" box="[467,624,1828,1855]" italics="true" pageId="17" pageNumber="118">S. lanceolatus</emphasis>
</taxonomicName>
has shown affinities with other inachoidids as well as its separate status (
<bibRefCitation id="8A014B21FFB8A61EB6F0F8BDFE72F8DB" author="Paula &amp; Cartaxana" box="[159,439,1864,1891]" firstAuthor="Paula" pageId="17" pageNumber="118" pagination="113 - 122" refId="ref24414" refString="Paula, J. &amp; Cartaxana, A. (1991) Complete larval development of the spider crab Stenorhynchus lanceolatus (Brulle, 1837) (Decapoda, Brachyura, Majidae), reared in the laboratory. Crustaceana, 60 (2), 113 - 122. https: // doi. org / 10.1163 / 156854091 X 00335" type="journal article" year="1991">Paula &amp; Cartaxana 1991</bibRefCitation>
), justifying the erection of its own subfamily. According to
<bibRefCitation id="8A014B21FFB8A61EB237F8BCFA59F8DB" author="Marques &amp; Pohle" box="[1112,1436,1864,1891]" firstAuthor="Marques" pageId="17" pageNumber="118" pagination="71 - 82" refId="ref23399" refString="Marques, F. P. L. &amp; Pohle, G. (2003) Searching for larval support for majid subfamilies (Crustacea: Brachyura) with particular reference to Inachoidinae Dana, 1851. Invertebrate Reproduction and Development, 43, 71 - 82. https: // doi. org / 10.1080 / 07924259.2003.9652523" type="journal article" year="2003">Marques &amp; Pohle (2003: 77)</bibRefCitation>
larval evidence revealed that
<taxonomicName id="29904D53FFB8A61EB78AF899FD4CF83E" authorityName="Lamarck" authorityYear="1815" box="[485,649,1900,1926]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB8A61EB78AF899FD4CF83E" box="[485,649,1900,1926]" italics="true" pageId="17" pageNumber="118">Stenorhynchus</emphasis>
</taxonomicName>
nested basal to
<taxonomicName id="29904D53FFB8A61EB52DF899FC13F83E" authorityName="Drach &amp; Guinot" authorityYear="1983" box="[834,982,1900,1926]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="family">Inachoididae</taxonomicName>
.
<taxonomicName id="29904D53FFB8A61EB58DF899FB5BF83E" authorityName="Dana, 1851 (Guinot" authorityYear="2012" box="[994,1182,1900,1926]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="subFamily" subFamily="Stenorhynchinae">Stenorhynchinae</taxonomicName>
is the only inachoidid representative occurring outside the Western Hemisphere, namely
<taxonomicName id="29904D53FFB8A61EB510F864FBD9F812" baseAuthorityName="Brulle" baseAuthorityYear="1837" box="[895,1052,1936,1963]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="lanceolatus">
<emphasis id="DCE4EAC2FFB8A61EB510F864FBD9F812" box="[895,1052,1936,1963]" italics="true" pageId="17" pageNumber="118">S. lanceolatus</emphasis>
</taxonomicName>
, the ‘eastern Atlantic arrow crab’, that is found in the eastern Atlantic (Madeira, Canary Is.,
<collectingCountry id="96877640FFB8A61EB57EF840FC54F876" box="[785,913,1972,1999]" name="Cape Verde" pageId="17" pageNumber="118">Cape Verde</collectingCountry>
Is., and numerous west-African localities from
<collectingCountry id="96877640FFB8A61EB6F8F82CFE80F84A" box="[151,325,2008,2035]" name="Western Sahara" pageId="17" pageNumber="118">Western Sahara</collectingCountry>
to
<collectingCountry id="96877640FFB8A61EB708F82DFE7EF84B" box="[359,443,2008,2035]" name="Angola" pageId="17" pageNumber="118">Angola</collectingCountry>
) (
<bibRefCitation id="8A014B21FFB8A61EB7BDF82CFD4FF84B" author="Capart" box="[466,650,2008,2035]" firstAuthor="Capart" pageId="17" pageNumber="118" pagination="11 - 205" refId="ref19900" refString="Capart, A. (1951) Crustaces Decapodes Brachyures. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de I'Atlantique Sud (1948 - 1949). Resultats Scientifiques, 3 (1), 11 - 205." type="journal article" year="1951">Capart 1951: 81</bibRefCitation>
, fig. 25;
<bibRefCitation id="8A014B21FFB8A61EB482F82DFC7CF84B" author="Monod" box="[749,953,2008,2035]" firstAuthor="Monod" pageId="17" pageNumber="118" pagination="1 - 674" refId="ref24178" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique Noire, 45, 1 - 674." type="journal article" year="1956">Monod 1956: 567</bibRefCitation>
, figs. 838, 839;
<bibRefCitation id="8A014B21FFB8A61DB203F82DFF01FF0B" author="Manning &amp; Holthuis" firstAuthor="Manning" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118" pagination="1 - 379" refId="ref23202" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, i-xii + 1 - 379. https: // doi. org / 10.5479 / si. 00810282.306" type="journal article" year="1981">Manning &amp; Holthuis 1981: 304</bibRefCitation>
, fig. 78b). Species of
<taxonomicName id="29904D53FFBBA61DB7D6FF6CFD98FF0B" authorityName="Lamarck" authorityYear="1815" box="[441,605,153,179]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFBBA61DB7D6FF6CFD98FF0B" box="[441,605,153,179]" italics="true" pageId="18" pageNumber="119">Stenorhynchus</emphasis>
</taxonomicName>
are for most not decorator crabs and display a peculiar behaviour (
<bibRefCitation id="8A014B21FFBBA61DB322FF6CFEA1FF6F" author="Guinot &amp; Wicksten" firstAuthor="Guinot" pageId="18" pageNumber="119" pagination="583 - 638" refId="ref22202" refString="Guinot, D. &amp; Wicksten, M. K. (2015) Chapter 71 - 11. Camouflage: carrying behaviour, decoration behaviour, and other modalities of concealment. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. R. &amp; von Vaupel Klein, J. C. (Eds.), Decapoda: Brachyura, Treatise on Zoology-Anatomy, Taxonomy, Biology. Crustacea. Vol. 9. Part C- 1, Brill, Leiden and Boston, pp. 583 - 638. https: // doi. org / 10.1163 / 9789004190832 _ 013" type="book chapter" year="2015">Guinot &amp; Wicksten 2015</bibRefCitation>
: fig. 71.11.6G). Molecular sequences obtained from
<specimenCount id="F896FD59FFBBA61DB5ADFF48FBE9FF6F" box="[962,1068,189,215]" pageId="18" pageNumber="119" type="larva">42 larvae</specimenCount>
of
<taxonomicName id="29904D53FFBBA61DB239FF48FAB4FF6F" baseAuthorityName="Herbst" baseAuthorityYear="1788" box="[1110,1393,189,215]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="DCE4EAC2FFBBA61DB239FF48FAB4FF6F" box="[1110,1393,189,215]" italics="true" pageId="18" pageNumber="119">Stenorhynchus seticornis</emphasis>
</taxonomicName>
are deposited in GenBank (
<bibRefCitation id="8A014B21FFBBA61DB7F0FF14FD47FF44" author="Brandao" box="[415,642,225,252]" etAl="et al." firstAuthor="Brandao" pageId="18" pageNumber="119" pagination="288 - 302" refId="ref19729" refString="Brandao, M. C., Freire, A. F. &amp; Burton, R. S. (2016) Estimating diversity of crabs (Decapoda: Brachyura) in a no-take marine protected area of the SW Atlantic coast through DNA barcoding of larvae. Systematics and Biodiversity, 14, 288 - 302. https: // doi. org / 10.1080 / 14772000.2016.1140245" type="journal article" year="2016">
Brandão
<emphasis id="DCE4EAC2FFBBA61DB469FF17FDF2FF43" box="[518,567,225,251]" italics="true" pageId="18" pageNumber="119">et al</emphasis>
. 2016
</bibRefCitation>
).
</paragraph>
<paragraph id="EE2F36D0FFBBA61DB6A8FEF0FEFEFEFB" blockId="18.[151,1437,153,323]" pageId="18" pageNumber="119">
The family
<taxonomicName id="29904D53FFBBA61DB726FEF0FE1FFEA7" authorityName="Drach &amp; Guinot" authorityYear="1983" box="[329,474,261,287]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="119" phylum="Arthropoda" rank="family">Inachoididae</taxonomicName>
did not invade areas outside the Americas and did not radiate in the Old World, except
<taxonomicName id="29904D53FFBBA61DB6F8FEDCFEF1FEFB" baseAuthorityName="Brulle" baseAuthorityYear="1837" box="[151,308,297,323]" class="Malacostraca" family="Inachoididae" genus="Stenorhynchus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="lanceolatus">
<emphasis id="DCE4EAC2FFBBA61DB6F8FEDCFEF1FEFB" box="[151,308,297,323]" italics="true" pageId="18" pageNumber="119">S. lanceolatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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