405 lines
40 KiB
XML
405 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="2621C9F4952125581691806F773AE802" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole longispinosa Forel 1891" docType="treatment" docVersion="4" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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<treatment ID-GBIF-Taxon="161777002" LSID="urn:lsid:plazi:treatment:2621C9F4952125581691806F773AE802" httpUri="http://treatment.plazi.org/id/2621C9F4952125581691806F773AE802" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="Forel, 1891" authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longispinosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longispinosa">Pheidole longispinosa Forel, 1891</taxonomicName>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">Figs 23A-F</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84Y</figureCitation>
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<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longispinosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longispinosa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole longispinosa</emphasis>
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</taxonomicName>
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Forel, 1891: 170, pl. 5, fig. 4 (s.w.q.). Lectotype [designated here]: major worker (top specimen, CASENT0101682): Madagascar, Antananarivo, Ambatomanjaka, Miarinarivo (MHNG) [examined]. Paralectotypes: 1 major worker (CASENT0810542, bottom specimen, the same pin as lectotype) (MHNG) [examined], 3 minor workers (1 pin, CASENT0101598, CASENT0923202, CASENT0923203) (MHNG) [examined], 1 dealate queen (CASENT0101657) (MHNG) [examined]: the same data as lectotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antananarivo</emphasis>
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: •12w., 2s.; 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe; -18.47333, 47.96; alt. 1300 m; 5 Dec 2000; B.L. Fisher et al. leg.; CASENT0406929, CASENT0406930, CASENT0413533, CASENT0413537, CASENT0427722, CASENT0427724, CASENT0427725, CASENT0427733. •1 w.; Ambatolaona;
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<geoCoordinate degrees="18.928" direction="south" orientation="latitude" precision="55" value="-18.928">-18.928</geoCoordinate>
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,
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<geoCoordinate degrees="47.88283" direction="east" orientation="longitude" precision="1" value="47.88283">47.88283</geoCoordinate>
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; alt. 1382 m; 19 Feb 2007; B.L. Fisher et al. leg.; CASENT0120674 (CASC). •4w., 1s, 1q.; Mandraka;
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<geoCoordinate degrees="18.91813" direction="south" orientation="latitude" precision="1" value="-18.91813">-18.91813</geoCoordinate>
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,
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<geoCoordinate degrees="47.91717" direction="east" orientation="longitude" precision="1" value="47.91717">47.91717</geoCoordinate>
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; alt. 1312 m; 20 Mar 2014; B.L. Fisher et al. leg.; CASENT0377078, CASENT0378205, CASENT0378163, CASENT0378568 (CASC). •4w, 1m.; Mandraka Park;
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<geoCoordinate degrees="18.9019" direction="south" orientation="latitude" precision="5" value="-18.9019">-18.9019</geoCoordinate>
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,
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<geoCoordinate degrees="47.90786" direction="east" orientation="longitude" precision="1" value="47.90786">47.90786</geoCoordinate>
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; 1360 m; 11 Mar 2012; B.L. Fisher et al. leg.; CASENT0275803, CASENT0275804, CASENT0275863 (CASC). •11w., 5s., 1m., 1q; Reg. Analamanga, St.
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<normalizedToken originalValue="Forestière">Forestiere</normalizedToken>
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Mandraka;
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<geoCoordinate degrees="18.9183" direction="south" orientation="latitude" precision="5" value="-18.9183">-18.9183</geoCoordinate>
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,
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<geoCoordinate degrees="47.91687" direction="east" orientation="longitude" precision="1" value="47.91687">47.91687</geoCoordinate>
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; alt. 1285 m; 25 Mar 2015; B.L. Fisher et al. leg.; CASENT0390404-CASENT0390406, CASENT0390415, CASENT0390429, CASENT0390496, CASENT0390497, CASENT0390506, CASENT0390533, CASENT0720697, CASENT0720833 (CASC); •3w., 1s.; Tsinjoarivo forest, Ankadivory;
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<geoCoordinate degrees="19.71572" direction="south" orientation="latitude" precision="1" value="-19.71572">-19.71572</geoCoordinate>
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,
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<geoCoordinate degrees="47.82084" direction="east" orientation="longitude" precision="1" value="47.82084">47.82084</geoCoordinate>
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; alt. 1385 m; 23 Aug 2014; B.L. Fisher et al. leg.; CASENT0389650, CASENT0389655, CASENT0389682 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1 w.; 9.2 km WSW Befingotra,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1280 m; 5 Nov 1994; B.L. Fisher leg.; CASENT0198233 (CASC). •3w., 1s., 2q; Betaolana Forest, along Bekona River;
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<geoCoordinate degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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,
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<geoCoordinate degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 4 Mar 2009; B.L. Fisher et al. leg.; CASENT0152375, CASENT0152403, CASENT0152410 (CASC).•3w.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 17 Nov 2003; B.L. Fisher leg.; CASENT0487914 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •5w., 2s.; 2 km W Andrambovato, along river Tatamaly; -21.51167, 47.41; alt. 1075 m; 3 Jun 2005; B.L. Fisher et al. leg.; CASENT0060852, CASENT0060853, CASENT0060932, CASENT0060933, CASENT0061669 (CASC). •1w.; 38 km S Ambalavao,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Andringitra; -22.2, 46.96667; alt. 1680 m; 23 Oct 1993; B.L. Fisher leg.; CASENT0198397 (CASC). •3w., 1q.; 40 km S Ambalavao,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Andringitra;
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<geoCoordinate degrees="22.21667" direction="south" orientation="latitude" precision="1" value="-22.21667">-22.21667</geoCoordinate>
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,
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<geoCoordinate degrees="46.96667" direction="east" orientation="longitude" precision="1" value="46.96667">46.96667</geoCoordinate>
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; alt. 1275 m; 19 Oct 1993; B.L. Fisher leg.; CASENT0198376, CASENT0198377 (CASC). •7w, 1s., 1q.; 8.0 km NE Ivohibe;
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<geoCoordinate degrees="22.42167" direction="south" orientation="latitude" precision="1" value="-22.42167">-22.42167</geoCoordinate>
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,
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<geoCoordinate degrees="46.89833" direction="east" orientation="longitude" precision="1" value="46.89833">46.89833</geoCoordinate>
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; alt. 1200 m; 3 Nov 1997; B.L. Fisher leg.; CASENT0198369, CASENT0198371, CASENT0198803, CASENT0198868 (CASC). •4w., 3s., 1q.; Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango;
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<geoCoordinate degrees="23.84083" direction="south" orientation="latitude" precision="1" value="-23.84083">-23.84083</geoCoordinate>
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,
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<geoCoordinate degrees="46.9575" direction="east" orientation="longitude" precision="5" value="46.9575">46.9575</geoCoordinate>
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; alt. 1250 m; 17 Nov 2006; B.L. Fisher et al. leg.; CASENT0118416, CASENT0119618, CASENT0119592, CASENT0118442 (CASC). •7w., 1s., 1q.; R.S. Ivohibe 8.0 km E Ivohibe;
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<geoCoordinate degrees="22.48333" direction="south" orientation="latitude" precision="1" value="-22.48333">-22.48333</geoCoordinate>
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,
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<geoCoordinate degrees="46.96833" direction="east" orientation="longitude" precision="1" value="46.96833">46.96833</geoCoordinate>
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; alt. 1200 m; 15 Oct 1997; B.L. Fisher leg.; CASENT0196901, CASENT0198003, CASENT0198004, CASENT0198372, CASENT0198805, CASENT0198807, CASENT0198867 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1w.; [Morano-Chrome
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<normalizedToken originalValue="forêt">foret</normalizedToken>
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, 25 km W]; Morarano Chrome; Amparafaravola; -17.75, 47.98333; alt. 1276 m; 15 Jun 1991; A. Pauly leg.; CASENT0198378 (CASC). •1w., 1s., 1q.; 6.9 km NE Ambanizana, Ambohitsitondroina; -15.56667, 50; alt. 1000 m; 9 Dec 1993; B.L. Fisher leg.; CASENT0198379 (CASC). •1w.; Ambanizana, Parc National Masoala;
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<geoCoordinate degrees="15.57167" direction="south" orientation="latitude" precision="1" value="-15.57167">-15.57167</geoCoordinate>
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,
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<geoCoordinate degrees="50.00611" direction="east" orientation="longitude" precision="1" value="50.00611">50.00611</geoCoordinate>
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; alt. 800-897 m; 26 Feb 2003; Andriamalala D. et al. leg.; CASENT0047728 (CASC). •8w.; Ankerana;
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<geoCoordinate degrees="18.40636" direction="south" orientation="latitude" precision="1" value="-18.40636">-18.40636</geoCoordinate>
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,
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<geoCoordinate degrees="48.80254" direction="east" orientation="longitude" precision="1" value="48.80254">48.80254</geoCoordinate>
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; alt. 1108 m; 19 Jan 2012; B.L. Fisher et al. leg.; CASENT0273728, CASENT0274171, CASENT0274172, CASENT0274173, CASENT0274993, CASENT0274994, CASENT0274994 (CASC). •1w., 1s.; Corridor Forestier Analamay-Mantadia, Tsaravoniana;
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<geoCoordinate degrees="18.76124" direction="south" orientation="latitude" precision="1" value="-18.76124">-18.76124</geoCoordinate>
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,
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<geoCoordinate degrees="48.42134" direction="east" orientation="longitude" precision="1" value="48.42134">48.42134</geoCoordinate>
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; alt. 939 m; 2 Dec 2012; B.L. Fisher et al. leg.; CASENT0300341 (CASC). •3w., 2s.; F.C. Didy;
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<geoCoordinate degrees="18.19833" direction="south" orientation="latitude" precision="1" value="-18.19833">-18.19833</geoCoordinate>
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,
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<geoCoordinate degrees="48.57833" direction="east" orientation="longitude" precision="1" value="48.57833">48.57833</geoCoordinate>
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; alt. 960 m; 16 Dec 1998; Ratsirarson H.J. leg.; CASENT0198005, CASENT0198006, CASENT0198373 (CASC). •2w., 1s.; Montagne
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<normalizedToken originalValue="d’Anjanaharibe">d'Anjanaharibe</normalizedToken>
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, 19.5 km 27°NNE Ambinanitelo;
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<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
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,
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<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
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; alt. 1100 m; 12 Mar 2003; B.L. Fisher et al. leg.; CASENT0037455, CASENT0037462, CASENT0048698, CASENT0048727 (CASC). •1w.; P.N. Mantadia;
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<geoCoordinate degrees="18.79167" direction="south" orientation="latitude" precision="1" value="-18.79167">-18.79167</geoCoordinate>
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,
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||
<geoCoordinate degrees="48.42667" direction="east" orientation="longitude" precision="1" value="48.42667">48.42667</geoCoordinate>
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; alt. 895 m; 25 Nov 1998; Ratsirarson H.J. leg.; CASENT0198384 (CASC). •1w., 1s.; Parc National
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<normalizedToken originalValue="d´">d'</normalizedToken>
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Andasibe-Mantadia,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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de Mantadia, 25.7 km 248° Moramanga;
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<geoCoordinate degrees="18.81402" direction="south" orientation="latitude" precision="1" value="-18.81402">-18.81402</geoCoordinate>
|
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,
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<geoCoordinate degrees="48.43028" direction="east" orientation="longitude" precision="1" value="48.43028">48.43028</geoCoordinate>
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; alt. 1040 m; 13 Jul 2006; Raharimalala & Blaimer leg.; CASENT0117442 (CASC). •1w., 1s.; Parc National de Zahamena, Onibe River;
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||
<geoCoordinate degrees="17.75908" direction="south" orientation="latitude" precision="1" value="-17.75908">-17.75908</geoCoordinate>
|
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,
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||
<geoCoordinate degrees="48.85468" direction="east" orientation="longitude" precision="1" value="48.85468">48.85468</geoCoordinate>
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; alt. 780 m; 21 Feb 2009; B.L. Fisher et al. leg.; CASENT0152053 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
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||
: •7w., 1s., 1q.; 13 km NW Enkara,
|
||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||
Andohahela; -24.55, 46.8; alt. 1300 m; 2 Dec 1992; B.L. Fisher leg.; CASENT0198374, CASENT0198375, CASENT0198381, CASENT0198382 (CASC). •31w., 14s., 9q.; Anosy Region, Anosyenne Mts, 29.33 km NW Manantenina;
|
||
<geoCoordinate degrees="24.13993" direction="south" orientation="latitude" precision="1" value="-24.13993">-24.13993</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="47.07418" direction="east" orientation="longitude" precision="1" value="47.07418">47.07418</geoCoordinate>
|
||
; alt. 540 m; 21 Feb 2015; B.L. Fisher et al. leg.; CASENT0704727, CASENT0704727, CASENT0704749, CASENT0704780, CASENT0723465, CASENT0704473, CASENT0704200, CASENT0704201, CASENT0704205, CASENT0704206, CASENT0704217, CASENT0704218, CASENT0704226, CASENT0704227, CASENT0704228, CASENT0704229, CASENT0704242, CASENT0704245, CASENT0704273, CASENT0704353, CASENT0704354, CASENT0704355, CASENT0704393, CASENT0704394, CASENT0704867, CASENT0704874, CASENT0704886, CASENT0704887, CASENT0705875, CASENT0705877, CASENT0721001, CASENT0721005, CASENT0721007, CASENT0721009, CASENT0721013 (CASC). •20w., 2s.; Parc National
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||
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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||
, Col du Sedro, 3.8 km 113°ESE Mahamavo, 37.6 km 341°NNW Tolagnaro;
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||
<geoCoordinate degrees="24.76389" direction="south" orientation="latitude" precision="1" value="-24.76389">-24.76389</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.75167" direction="east" orientation="longitude" precision="1" value="46.75167">46.75167</geoCoordinate>
|
||
; alt. 900 m; 21 Jan 2002; B.L. Fisher et al. leg.; CASENT0078380, CASENT0430695, CASENT0430759, CASENT0451281, CASENT0451303, CASENT0451309, CASENT0456249, CASENT0460156, CASENT0460158, CASENT0483964, CASENT0484129, CASENT0484130 (CASC). •3w.; Parc National de Kirindy Mite, 16.3 km 127°SE Belo sur Mer;
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<geoCoordinate degrees="20.79528" direction="south" orientation="latitude" precision="1" value="-20.79528">-20.79528</geoCoordinate>
|
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,
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||
<geoCoordinate degrees="44.147" direction="east" orientation="longitude" precision="55" value="44.147">44.147</geoCoordinate>
|
||
; alt. 80 m; 6 Dec 2001; B.L. Fisher et al. leg.; CASENT0477205 (CASC). •5w. 2s.,1q.;
|
||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||
Kalambatritra, Ambinanitelo;
|
||
<geoCoordinate degrees="23.4502" direction="south" orientation="latitude" precision="5" value="-23.4502">-23.4502</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.45658" direction="east" orientation="longitude" precision="1" value="46.45658">46.45658</geoCoordinate>
|
||
; alt. 1325 m; 11 Feb 2009; B.L. Fisher et al. leg.; CASENT0148863, CASENT0148866, CASENT0149937-CASENT0149939 (CASC). •18w., 7s., 3m., 11q.;
|
||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||
Kalambatritra, Ampanihy;
|
||
<geoCoordinate degrees="23.463" direction="south" orientation="latitude" precision="55" value="-23.463">-23.463</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.47057" direction="east" orientation="longitude" precision="1" value="46.47057">46.47057</geoCoordinate>
|
||
; alt. 1269 m; 10 Feb 2009; B.L. Fisher et al. leg.; CASENT0148871, CASENT0148872, CASENT0148875, CASENT0148945, CASENT0148953, CASENT0148964, CASENT0148967, CASENT0148972, CASENT0148983, CASENT0148994, CASENT0148995, CASENT0148996, CASENT0149001, CASENT0149008, CASENT0149801, CASENT0149992, CASENT0150511, CASENT0150535, CASENT0151012, CASENT0151013, CASENT0151019, CASENT0151020, CASENT0152223, CASENT0153227, CASENT0235023 (CASC). •6w., 3s., 2q.;
|
||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||
Kalambatritra, Befarara;
|
||
<geoCoordinate degrees="23.4178" direction="south" orientation="latitude" precision="5" value="-23.4178">-23.4178</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.4478" direction="east" orientation="longitude" precision="5" value="46.4478">46.4478</geoCoordinate>
|
||
; alt. 1390 m; 7 Feb 2009; B.L. Fisher et al. leg.; CASENT0149598, CASENT0149599, CASENT0149869, CASENT0149871, CASENT0150723 (CASC). •1w., 1s.;
|
||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||
Kalambatritra, Betanana;
|
||
<geoCoordinate degrees="23.4144" direction="south" orientation="latitude" precision="5" value="-23.4144">-23.4144</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46.459" direction="east" orientation="longitude" precision="55" value="46.459">46.459</geoCoordinate>
|
||
; alt. 1360 m; 8 Feb 2009; B.L. Fisher et al. leg.; CASENT0148654 (CASC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
Large species: HL: 2.62-3.0 (2.77), HW: 2.64-2.92 (2.75), WL: 1.88-2.14 (2.04); propodeal spines very long (PSL: 0.67-0.87 (0.73)); head in full-face view trapezoid, widened posteriorly; sides of the head without pilosity; frons with longitudinal rugae never reaching further than midlength of head; gaster smooth and shiny; body brown to dark brown.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
Large species: HL: 0.99-1.15 (1.1), HW: 0.98-1.06 (1.0), WL: 1.62-1.76 (1.66); propodeal spines very long (PSL: 0.62-0.76 (0.69)); scape, when laid back, surpassing posterior head margin by more than half its length; mesosoma smooth and shiny, only anepisternum, katepisternum, and propodeum sometimes with indistinct and sparse superficial rugulae; promesonotal groove absent; metanotal groove shallow and wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="redescription">
|
||
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 2.62-3.0 (2.77); HW: 2.64-2.92 (2.75); SL: 1.24-1.35 (1.3); EL: 0.24-0.28 (0.26); WL: 1.88-2.14 (2.04); PSL: 0.67-0.87 (0.73); MTL: 1.39-1.55 (1.46); PNW: 0.8-0.97 (0.89); PTW: 0.23-0.31 (0.27); PPW: 0.68-0.82 (0.75); CI: 94.3-106.4 (99.3); SI: 43.9-50.9 (47.4); PSLI: 24.1-29.0 (26.5); PPI: 30.1-42.3 (35.4); PNI: 30.45-33.38 (32.4); MTI: 49.6-57.6 (53.0).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
In full-face view trapezoid, widened posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23B</figureCitation>
|
||
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head without pilosity; frons and vertex with few, long, erect setae. Antennal scrobes absent or very indistinct. Occipital lobes, genae, posterior part of frons smooth and shiny; lateral sides of head and genae with sparse, partially fading superficial rugulae; centre of frons shiny with fine, thin, longitudinal rugae reaching at most to midlength of head; malar area with fine, thin, longitudinal rugae. Centre of clypeus smooth and shiny, lateral sides with fine and sparse rugae; median notch present, shallow, and wide; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching midlength of head; pilosity suberect to erect (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, low, triangular and thick, with rounded apex, closely spaced; outer hypostomal teeth distinct, slightly smaller and thinner than inner hypostomal teeth, lobe-like (Fig.
|
||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84Y</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, short, and relatively flat; pronotum slightly convex; dorsal mesonotum concave; posterior mesonotum relatively steep; promesonotal groove absent; metanotal groove absent; propodeal spines very long, massive basally, with acute apex; humeral area laterally either weakly or not produced (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D</figureCitation>
|
||
). Surface shiny and smooth with weak, sparse, and irregular rugae on pronotum and lateral sides of propodeum. Pilosity very sparse, short, and decumbent; dorsum with few additional, long, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Smooth and shiny; peduncle long and thin, superficially rugulose ventrally; node triangular, with rounded apex, in rear view node dorsoventrally depressed; pilosity sparse, short, and erect (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Smooth and shiny; in dorsal view sides with acute, narrow, moderately short triangular projections; pilosity short to long, sparse, and erect (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
|
||
.
|
||
</emphasis>
|
||
Smooth and shiny; pilosity sparse, erect, and long (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, brown to dark brown (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1" start="Figure 23" startId="F23">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 23.</emphasis>
|
||
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longispinosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longispinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole longispinosa</emphasis>
|
||
</taxonomicName>
|
||
Forel, full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of minor worker (CASENT0451282) and full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of major worker (CASENT0451284).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.99-1.15 (1.1); HW: 0.98-1.06 (1.0); SL: 1.35-1.47 (1.4); EL: 0.14-0.21 (0.18); WL: 1.62-1.76 (1.66); PSL: 0.62-0.76 (0.69); MTL: 1.21-1.29 (1.24); PNW: 0.65-0.7 (0.67); PTW: 0.13-0.17 (0.16); PPW: 0.22-0.27 (0.25); CI: 88.3-99.1 (91.3); SI: 136.7-144.4 (139.7); PSLI: 56.8-67.8 (62.5); PPI: 56.3-66.9 (62.9); PNI: 65.6-68.3 (66.9); MTI: 121.3-128.1 (124.4).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin convex; occipital carina narrow, weakly developed (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23A</figureCitation>
|
||
). Pilosity sparse, long, and erect on frons and median part of occiput. Sculpture smooth and shiny on the whole surface, only antennal sockets with sparse carinae curved outward. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae present. Scape, when laid back, surpassing posterior head margin by more than half of its length; pilosity suberect to erect (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, long, and slightly convex; promesonotal groove absent; metanotal groove indistinct and wide; propodeal spines very long, massive basally, with acute apex (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C</figureCitation>
|
||
). Surface smooth and shiny, only anepisternum, katepisternum, and propodeum sometimes with indistinct and sparse superficial rugulae. Pilosity absent or short, very sparse, suberect (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Smooth and shiny; peduncle long and thin; node bulge-like with rounded apex; pilosity absent (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Moderately long, low, and with slightly convex dorsum; with two long, erect setae at the anterior edge (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
|
||
.
|
||
</emphasis>
|
||
With few long, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, brown to dark brown (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole longispinosa Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0451282) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0451284)." figureDoi="10.3897/zookeys.905.39592.figure23" httpUri="https://binary.pensoft.net/fig/373070" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected at between 80-1680 m in elevation, in montane rainforest, montane shrubland, thicket, tropical dry forest, and urban areas. Nests were located in rotten logs, tree stumps, soil, and rock crevasses.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species is most similar to
|
||
<taxonomicName lsidName="P. mahaboensis" pageId="0" pageNumber="1" rank="species" species="mahaboensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. mahaboensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName lsidName="P. praegrandis" pageId="0" pageNumber="1" rank="species" species="praegrandis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. praegrandis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longispinosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longispinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole longispinosa</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName lsidName="P. mahaboensis" pageId="0" pageNumber="1" rank="species" species="mahaboensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. mahaboensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by absence of metanotal groove, lack of pilosity at the sides of head, and longitudinal rugae never reaching further than midlength of head; from
|
||
<taxonomicName lsidName="P. praegrandis" pageId="0" pageNumber="1" rank="species" species="praegrandis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. praegrandis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by brighter body colouration, reduced head sculpture, and not shagreened first gastral tergite and head.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longispinosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longispinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole longispinosa</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName lsidName="P. mahaboensis" pageId="0" pageNumber="1" rank="species" species="mahaboensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. mahaboensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by absence of promesonotal groove, shallow and wide metanotal groove and reduced sculpture on mesosoma; from
|
||
<taxonomicName lsidName="P. praegrandis" pageId="0" pageNumber="1" rank="species" species="praegrandis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. praegrandis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by mostly smooth and shiny sculpture of head and mesosoma, which is never rugoreticulate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |