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<document id="04017AF0A9A14CE2034708AE219AF661" ID="10.11646/zootaxa.4247.2.8SLASH10643" ID-CLB-Dataset="36191" ID-DOI="10.11646/zootaxa.4247.2.8" ID-GBIF-Dataset="26522fb1-7785-4e6e-88f3-05d0f577530b" ID-ISSN="1175-5326" ID-Zenodo-Dep="438249" ID-ZooBank="9E8C6460-ABF0-4491-A2F5-2CA6E2ADF666" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490602122214" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="03F587FDFFD2FFBBFF52FC69FC8F969E" docLanguage="en" docName="zootaxa.4247.2.8SLASH10643.pdf" docOrigin="Zootaxa 4247 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bellendenker Jałoszyński, 2017, gen. nov." docType="treatment" docVersion="7" lastPageNumber="171" masterDocId="FFCCFF85FFD3FFBCFFC5FF83FFED9716" masterDocTitle="Bellendenker gen. nov., a new Australian genus of Glandulariini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="173" masterPageNumber="165" pageNumber="166" updateTime="1698438423630" updateUser="ExternalLinkService">
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<mods:title id="830785CD8D2A460E4A25C8D6D3F882E8">Bellendenker gen. nov., a new Australian genus of Glandulariini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="E99634CFD1809FAE3687A0DD5258A132" type="personal">
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<mods:namePart id="14CE692C062C7560C2B0803DCDBBBF60">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="9B43DFBF7E8F3B463F629A325D52BCFE">Zootaxa</mods:title>
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<mods:date id="1EDFD0C3584DF7110C8D7B626399FB27">2017</mods:date>
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<mods:number id="12E5D28FE4B945D6D794ABC3958BE3C0">4247</mods:number>
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<mods:identifier id="1478541CDBC5C06C1C1147BDFCD5F3DE" type="DOI">10.11646/zootaxa.4247.2.8</mods:identifier>
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<treatment id="03F587FDFFD2FFBBFF52FC69FC8F969E" ID-DOI="http://doi.org/10.5281/zenodo.6019051" ID-GBIF-Taxon="127852541" ID-Zenodo-Dep="6019051" LSID="urn:lsid:plazi:treatment:03F587FDFFD2FFBBFF52FC69FC8F969E" httpUri="http://treatment.plazi.org/id/03F587FDFFD2FFBBFF52FC69FC8F969E" lastPageId="7" lastPageNumber="171" pageId="1" pageNumber="166">
<subSubSection id="C3466560FFD2FFBDFF52FC69FE409312" box="[151,429,1002,1028]" pageId="1" pageNumber="166" type="nomenclature">
<paragraph id="8BE336EBFFD2FFBDFF52FC69FE409312" blockId="1.[151,429,1002,1028]" box="[151,429,1002,1028]" pageId="1" pageNumber="166">
<heading id="D0AB8187FFD2FFBDFF52FC69FE409312" bold="true" box="[151,429,1002,1028]" fontSize="11" level="1" pageId="1" pageNumber="166" reason="1">
<emphasis id="B928EAF9FFD2FFBDFF52FC69FE409312" bold="true" box="[151,429,1002,1028]" pageId="1" pageNumber="166">
<taxonomicName id="4C5C4D68FFD2FFBDFF52FC69FED59312" ID-CoL="8XJZM" box="[151,312,1002,1028]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD2FFBDFF52FC69FED59312" bold="true" box="[151,312,1002,1028]" italics="true" pageId="1" pageNumber="166">Bellendenker</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5782FFD2FFBDFEFAFC69FE409312" box="[319,429,1002,1028]" pageId="1" pageNumber="166" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3466560FFD2FFBDFF52FBB2FCB3935C" box="[151,862,1073,1098]" pageId="1" pageNumber="166" type="materials_examined">
<paragraph id="8BE336EBFFD2FFBDFF52FBB2FCB3935C" blockId="1.[151,1437,1073,1998]" box="[151,862,1073,1098]" pageId="1" pageNumber="166">
<emphasis id="B928EAF9FFD2FFBDFF52FBB2FEDF935C" bold="true" box="[151,306,1073,1098]" pageId="1" pageNumber="166">Type species.</emphasis>
<taxonomicName id="4C5C4D68FFD2FFBDFEFCFBB2FDC3935F" box="[313,558,1073,1097]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD2FFBDFEFCFBB2FDC3935F" box="[313,558,1073,1097]" italics="true" pageId="1" pageNumber="166">Bellendenker palpalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5782FFD2FFBDFDF0FBB2FD63935F" box="[565,654,1073,1098]" pageId="1" pageNumber="166" rank="species">
<emphasis id="B928EAF9FFD2FFBDFDF0FBB2FD6A935C" bold="true" box="[565,647,1073,1098]" pageId="1" pageNumber="166">sp. nov</emphasis>
.
</taxonomicNameLabel>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="C3466560FFD2FFBDFF02FBD6FEDB9170" pageId="1" pageNumber="166" type="diagnosis">
<paragraph id="8BE336EBFFD2FFBDFF02FBD6FEDB9170" blockId="1.[151,1437,1073,1998]" pageId="1" pageNumber="166">
<emphasis id="B928EAF9FFD2FFBDFF02FBD6FEAC9378" bold="true" box="[199,321,1109,1134]" pageId="1" pageNumber="166">Diagnosis.</emphasis>
Glandulariine genus with uniquely modified labial palps: palpomeres I and II extremely elongate so that labial palps are longer than labium; mandibles falciform, each with distinct mesal tooth located near basal third; eyes located posteriorly, distant from mandibular bases; clypeus much longer than labrum; frontoclypeal groove absent; labrum densely and asymmetrically setose; occipital constriction slightly narrower than frons between eyes; antennae gradually thickened distally; head and prothorax with thick bristles; pronotum broadest indistinctly in front of middle, with three pairs of antebasal pits; prothoracic hypomera lacking hypomeral ridges, each with subrectangular adcoxal lobe, with its anterior margin perpendicular to and connected with notosternal suture; basisternal portion of prosternum slightly shorter than procoxal cavities; prosternal process narrow, carinate, weakly elevated; mesoventrite with distinct pair of impressed procoxal rests separated at middle but not demarcated posteriorly; mesoventral intercoxal process broad, parallel-sided, moderately strongly elevated; metaventral intermetacoxal process subtriangular, narrowly separating metacoxae and with distinct median notch; each elytron with two rudimentary basal foveae covered by pronotal base in intact specimens, lacking basal impressions and humeral calli; hind wings absent; mesocoxa with two tufts of setae on dorsal surface; median lobe of aedeagus and endophallus symmetrical, parameres free, slender, lacking apical setae; spermatheca globular, asymmetrical.
</paragraph>
</subSubSection>
<subSubSection id="C3466560FFD2FFBAFF02F9F2FC1D9213" lastPageId="6" lastPageNumber="171" pageId="1" pageNumber="166" type="description">
<paragraph id="8BE336EBFFD2FFBDFF02F9F2FE6691C7" blockId="1.[151,1437,1073,1998]" pageId="1" pageNumber="166">
<emphasis id="B928EAF9FFD2FFBDFF02F9F2FEB4919C" bold="true" box="[199,345,1649,1674]" pageId="1" pageNumber="166">Description.</emphasis>
Body (
<figureCitation id="13672A6EFFD2FFBDFE73F9F2FE11919C" box="[438,508,1649,1674]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,931,954]" captionTargetBox="[163,1436,193,905]" captionTargetId="figure@2.[151,1436,193,910]" captionTargetPageId="2" captionText="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
) around
<quantity id="4CA49B0EFFD2FFBDFDABF9F1FD21919F" box="[622,716,1650,1674]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="166" unit="mm" value="1.7">1.7 mm</quantity>
, strongly convex, elongate but moderately slender, distinctly constricted between head and pronotum and between pronotum and elytra, with long appendages, pigmentation brown; cuticle setose.
</paragraph>
<paragraph id="8BE336EBFFD2FFBDFF02F95EFC5690D8" blockId="1.[151,1437,1073,1998]" pageId="1" pageNumber="166">
Head (
<figureCitation id="13672A6EFFD2FFBDFED3F95EFE8F91E0" box="[278,354,1757,1782]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,931,954]" captionTargetBox="[163,1436,193,905]" captionTargetId="figure@2.[151,1436,193,910]" captionTargetPageId="2" captionText="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="1" pageNumber="166">Figs 1</figureCitation>
,
<figureCitation id="13672A6EFFD2FFBDFEB0F95EFE7291E0" box="[373,415,1757,1782]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="1" pageNumber="166">35</figureCitation>
) divided by occipital constriction into exposed anterior part and posterior part largely retracted into pronotum; anterior part elongate, distinctly flattened, narrower than pronotum, broadest at eyes; frons (
<figureCitation id="13672A6EFFD2FFBDFF5AF8A6FF089028" box="[159,229,1829,1854]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="1" pageNumber="166">Fig. 3</figureCitation>
;
<emphasis id="B928EAF9FFD2FFBDFF36F8A6FEEB902B" box="[243,262,1829,1853]" italics="true" pageId="1" pageNumber="166">fr</emphasis>
) and vertex (
<figureCitation id="13672A6EFFD2FFBDFE5AF8A6FE089028" box="[415,485,1829,1854]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="1" pageNumber="166">Fig. 3</figureCitation>
;
<emphasis id="B928EAF9FFD2FFBDFE31F8A5FDE5902B" box="[500,520,1830,1853]" italics="true" pageId="1" pageNumber="166">vt</emphasis>
) confluent, together about as long as broad, weakly convex, posterior margin of vertex demarcated from 'neck' by broad impression; tempora (
<figureCitation id="13672A6EFFD2FFBDFCA2F8CAFC429074" box="[871,943,1865,1890]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="1" pageNumber="166">Fig. 3</figureCitation>
;
<emphasis id="B928EAF9FFD2FFBDFC7AF8C9FC369077" box="[959,987,1866,1889]" italics="true" pageId="1" pageNumber="166">tm</emphasis>
) about as long as eyes in dorsal view, strongly convergent posteriorly and nearly straight, with thick bristles; supraantennal tubercles distinct but relatively weakly elevated; clypeus (
<figureCitation id="13672A6EFFD2FFBDFD8BF812FD7790BC" box="[590,666,1937,1962]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="1" pageNumber="166">Fig. 3</figureCitation>
;
<emphasis id="B928EAF9FFD2FFBDFD6BF812FD2F90BF" box="[686,706,1937,1961]" italics="true" pageId="1" pageNumber="166">cl</emphasis>
) strikingly long, posteriorly at middle confluent with frons; compound eyes large and strongly projecting laterally, coarsely faceted.
</paragraph>
<caption id="DF236663FFD1FFBEFF52FC20FB4C94AC" box="[151,1185,931,954]" httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="2" pageNumber="167" targetBox="[163,1436,193,905]" targetPageId="2">
<paragraph id="8BE336EBFFD1FFBEFF52FC20FB4C94AC" blockId="2.[151,1185,931,954]" box="[151,1185,931,954]" pageId="2" pageNumber="167">
<emphasis id="B928EAF9FFD1FFBEFF52FC20FED494AF" bold="true" box="[151,313,931,954]" pageId="2" pageNumber="167">FIGURES 12.</emphasis>
<taxonomicName id="4C5C4D68FFD1FFBEFE85FC27FDF394AC" box="[320,542,932,954]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD1FFBEFE85FC27FDF394AC" box="[320,542,932,954]" italics="true" pageId="2" pageNumber="167">Bellendenker palpalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5782FFD1FFBEFDE1FC27FD9994AF" box="[548,628,932,953]" pageId="2" pageNumber="167" rank="species">
<emphasis id="B928EAF9FFD1FFBEFDE1FC27FD9994AF" bold="true" box="[548,628,932,953]" pageId="2" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male; dorsal habitus (1) and distribution (2).
</paragraph>
</caption>
<paragraph id="8BE336EBFFD1FFBEFF02FC62FD2092BC" blockId="2.[151,1437,993,2026]" pageId="2" pageNumber="167">
Labrum (
<figureCitation id="13672A6EFFD1FFBEFEEBFC62FE7F94EC" box="[302,402,993,1018]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 3, 5</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFE5BFC62FE5294EF" box="[414,447,993,1017]" italics="true" pageId="2" pageNumber="167">lbr</emphasis>
) strongly transverse and short, densely and asymmetrically setose. Mandibles (
<figureCitation id="13672A6EFFD1FFBEFAF7FC62FA7594EC" box="[1330,1432,993,1018]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 36</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF52FB86FF54930B" box="[151,185,1029,1053]" italics="true" pageId="2" pageNumber="167">md</emphasis>
) symmetrical, falciform, each with broad mandibular base (
<figureCitation id="13672A6EFFD1FFBEFC9DFB86FC709308" box="[856,925,1029,1054]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 6</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFC6CFB86FC26930B" box="[937,971,1029,1053]" italics="true" pageId="2" pageNumber="167">mb</emphasis>
) rapidly narrowed at small, subtriangular mesal tooth located near proximal 1/3 of mandible, distal portion of mandible strikingly long and slender, pointed and curved, setose prostheca present but not visible in intact specimens. Maxillae and maxillary palps do not deviate from ground plan of
<taxonomicName id="4C5C4D68FFD1FFBEFE39FBF2FD78939C" box="[508,661,1137,1162]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
. Labium (
<figureCitation id="13672A6EFFD1FFBEFCD1FBF2FC6D939C" box="[788,896,1137,1162]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 46</figureCitation>
) with large submentum (
<figureCitation id="13672A6EFFD1FFBEFB71FBF2FACC939C" box="[1204,1313,1137,1162]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 46</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFAF1FBF1FA8C939F" box="[1332,1377,1138,1161]" italics="true" pageId="2" pageNumber="167">smn</emphasis>
) not demarcated laterally by sutures, but with faint ridges extending posteromesally from cardines; subtrapezoidal mentum (
<figureCitation id="13672A6EFFD1FFBEFECDFB3AFE9993C4" box="[264,372,1209,1234]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 46</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFE43FB39FE4593C7" box="[390,424,1210,1233]" italics="true" pageId="2" pageNumber="167">mn</emphasis>
); and very long, largely membranous prementum (
<figureCitation id="13672A6EFFD1FFBEFBD5FB3AFB9193C4" box="[1040,1148,1209,1234]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Figs 46</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFB4BFB39FB5393C7" box="[1166,1214,1210,1233]" italics="true" pageId="2" pageNumber="167">pmn</emphasis>
) bearing narrowly separated at bases labial palps composed of short, annular palpomere I (
<figureCitation id="13672A6EFFD1FFBEFC7DFB5EFC1193E0" box="[952,1020,1245,1270]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 6</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFBCDFB5EFBC693E3" box="[1032,1067,1245,1269]" italics="true" pageId="2" pageNumber="167">lp1</emphasis>
), very long palpomere II (
<figureCitation id="13672A6EFFD1FFBEFA97FB5EFA7B93E0" box="[1362,1430,1245,1270]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 6</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF52FA82FF56920F" box="[151,187,1281,1305]" italics="true" pageId="2" pageNumber="167">lp2</emphasis>
) narrowing distally and about five times as long as broad at base, and extremely long and slender palpomere III (
<figureCitation id="13672A6EFFD1FFBEFF5AFAA6FF099228" box="[159,228,1317,1342]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 6</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF34FAA6FEF9922B" box="[241,276,1317,1341]" italics="true" pageId="2" pageNumber="167">lp3</emphasis>
) narrowing distally, weakly curved and about ten times as long as broad at base, labial palps are longer than labium; area between labial palps with a pair of long setae, but not forming defined ligula. Hypostomal ridges (
<figureCitation id="13672A6EFFD1FFBEFF5AFAEEFF079290" box="[159,234,1389,1414]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 4</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF3EFAEEFEF99293" box="[251,276,1389,1413]" italics="true" pageId="2" pageNumber="167">hr</emphasis>
) complete, strongly curved posteromesally and with their posterior ends recurved posteriorly, not connected and distant from posterior tentorial pits.
</paragraph>
<paragraph id="8BE336EBFFD1FFBEFF02FA36FD479100" blockId="2.[151,1437,993,2026]" pageId="2" pageNumber="167">
Gular plate (
<figureCitation id="13672A6EFFD1FFBEFE92FA36FE7292D8" box="[343,415,1461,1486]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 4</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFE68FA35FE2492DB" box="[429,457,1462,1485]" italics="true" pageId="2" pageNumber="167">gp</emphasis>
) large and subtrapezoidal, with rapidly narrowed unsculptured anterior portion; gular sutures indistinct; posterior tentorial pits (
<figureCitation id="13672A6EFFD1FFBEFDB4FA5AFD5592E4" box="[625,696,1497,1522]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@3.[151,1436,193,1712]" captionTargetPageId="3" captionText="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="2" pageNumber="167">Fig. 4</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFD00FA59FD0492E7" box="[709,745,1498,1521]" italics="true" pageId="2" pageNumber="167">ptp</emphasis>
) narrow and strongly elongate, located in front of transverse impression ventrally demarcating 'neck' region.
</paragraph>
<paragraph id="8BE336EBFFD1FFBEFF02F9A1FEFE9148" blockId="2.[151,1437,993,2026]" pageId="2" pageNumber="167">
Antennae (
<figureCitation id="13672A6EFFD1FFBEFE80F9A2FE65912C" box="[325,392,1569,1594]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,931,954]" captionTargetBox="[163,1436,193,905]" captionTargetId="figure@2.[151,1436,193,910]" captionTargetPageId="2" captionText="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="2" pageNumber="167">Fig. 1</figureCitation>
) long and slender, gradually but only slightly thickened distally; antennomeres compactly assembled.
</paragraph>
<paragraph id="8BE336EBFFD1FFBEFF02F9EAFD799004" blockId="2.[151,1437,993,2026]" pageId="2" pageNumber="167">
Prothorax (
<figureCitation id="13672A6EFFD1FFBEFE81F9EAFE669194" box="[324,395,1641,1666]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,931,954]" captionTargetBox="[163,1436,193,905]" captionTargetId="figure@2.[151,1436,193,910]" captionTargetPageId="2" captionText="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="2" pageNumber="167">Figs 1</figureCitation>
,
<figureCitation id="13672A6EFFD1FFBEFE5FF9E9FE289194" box="[410,453,1642,1666]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">79</figureCitation>
) in dorsal view elongate and broadest slightly in front of middle, with rounded anterior margin and anterior portions of lateral margins; anterior corners not marked; posterior corners distinct but blunt and nearly right-angled; posterior margin nearly straight. Base of pronotum with three pairs of distinct antebasal pits (
<figureCitation id="13672A6EFFD1FFBEFF5AF956FF0B91FB" box="[159,230,1749,1774]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 7</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF31F956FEF391FB" box="[244,286,1749,1773]" italics="true" pageId="2" pageNumber="167">abp</emphasis>
) not connected by groove, lacking lateral or sublateral carinae or edges. Sides of pronotum covered with thick bristles well visible in dorsal view.
</paragraph>
<paragraph id="8BE336EBFFD1FFBEFF02F89DFB7890FC" blockId="2.[151,1437,993,2026]" pageId="2" pageNumber="167">
Prosternum with moderately long basisternal part (
<figureCitation id="13672A6EFFD1FFBEFCC7F89EFCAB9020" box="[770,838,1821,1846]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 8</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFC91F89EFC999023" box="[852,884,1821,1845]" italics="true" pageId="2" pageNumber="167">bst</emphasis>
) demarcated from procoxal cavities (
<figureCitation id="13672A6EFFD1FFBEFADFF89EFAB29020" box="[1306,1375,1821,1846]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 8</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFAA9F89DFA7F9023" box="[1388,1426,1822,1845]" italics="true" pageId="2" pageNumber="167">pcc</emphasis>
) by arcuate and diffused ridges densely covered with setae; median part of sternum carinate but weakly elevated, forming prosternal process (
<figureCitation id="13672A6EFFD1FFBEFE14F8E6FDF89068" box="[465,533,1893,1918]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 8</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFDE4F8E5FDAA906B" box="[545,583,1894,1917]" italics="true" pageId="2" pageNumber="167">psp</emphasis>
) not separating procoxae; procoxal sockets closed by posterolateral lobes of prosternum; hypomera (
<figureCitation id="13672A6EFFD1FFBEFE69F80AFE1890B4" box="[428,501,1929,1954]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 8</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFDC0F80AFDCD90B7" box="[517,544,1929,1953]" italics="true" pageId="2" pageNumber="167">hy</emphasis>
) elongate, each divided into broad lateral part confluent with pronotum and narrower, elongate subrectangular inner (adcoxal) part with anterior margin perpendicular to notosternal suture (
<figureCitation id="13672A6EFFD1FFBEFF5AF852FF0990FC" box="[159,228,2001,2026]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="2" pageNumber="167">Fig. 8</figureCitation>
;
<emphasis id="B928EAF9FFD1FFBEFF35F851FEF990FF" box="[240,276,2002,2025]" italics="true" pageId="2" pageNumber="167">nss</emphasis>
; largely obscured by dense setae), the latter complete; hypomeral ridges absent.
</paragraph>
<caption id="DF236663FFD0FFBFFF52F947FB6C9021" httpUri="https://zenodo.org/record/438251/files/figure.png" pageId="3" pageNumber="168" targetBox="[151,1436,193,1712]" targetPageId="3">
<paragraph id="8BE336EBFFD0FFBFFF52F947FB6C9021" blockId="3.[151,1436,1732,1847]" pageId="3" pageNumber="168">
<emphasis id="B928EAF9FFD0FFBFFF52F947FED791CC" bold="true" box="[151,314,1732,1755]" pageId="3" pageNumber="168">FIGURES 36.</emphasis>
<taxonomicName id="4C5C4D68FFD0FFBFFE85F946FDF391CD" box="[320,542,1733,1755]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD0FFBFFE85F946FDF391CD" box="[320,542,1733,1755]" italics="true" pageId="3" pageNumber="168">Bellendenker palpalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5782FFD0FFBFFDE1F945FD9991CD" box="[548,628,1734,1755]" pageId="3" pageNumber="168" rank="species">
<emphasis id="B928EAF9FFD0FFBFFDE1F945FD9991CD" bold="true" box="[548,628,1734,1755]" pageId="3" pageNumber="168">sp. nov.</emphasis>
</taxonomicNameLabel>
; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp13, labial palpomere IIII; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex.
</paragraph>
</caption>
<paragraph id="8BE336EBFFD0FFBFFF02F8DDFDB5908D" blockId="3.[151,1436,1886,2019]" pageId="3" pageNumber="168">Mesocutellum subtriangular, in intact specimens not exposed, entirely hidden under posterior margin of pronotum overlapping with elytral base.</paragraph>
<paragraph id="8BE336EBFFD0FFB8FF02F825FC99960B" blockId="3.[151,1436,1886,2019]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168">
Mesoventrite (
<figureCitation id="13672A6EFFD0FFBFFEAAF825FE2B90A9" box="[367,454,1958,1983]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="3" pageNumber="168">Fig. 10</figureCitation>
) with massive and demarcated anterior ridge (
<figureCitation id="13672A6EFFD0FFBFFC33F825FBA290A9" box="[1014,1103,1958,1983]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="3" pageNumber="168">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD0FFBFFBA5F824FB9490A8" box="[1120,1145,1959,1982]" italics="true" pageId="3" pageNumber="168">ar</emphasis>
); mesoventral intercoxal process (
<figureCitation id="13672A6EFFD0FFBFFF3FF849FEA090F5" box="[250,333,1994,2019]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="3" pageNumber="168">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD0FFBFFE9DF848FE7F90F4" box="[344,402,1995,2018]" italics="true" pageId="3" pageNumber="168">msvp</emphasis>
) subrectangular, broad, parallel-sided, moderately elevated; anterior portion of mesoventrite with a pair of shallow setose impressions functioning as procoxal rests (
<figureCitation id="13672A6EFFD7FFB8FC29FF14FBA997A6" box="[1004,1092,151,176]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD7FFB8FB91FF1AFB9497A6" box="[1108,1145,153,176]" italics="true" pageId="4" pageNumber="169">pcr</emphasis>
), their posterior margins indistinct; mesanepisternum with long prepectus (
<figureCitation id="13672A6EFFD7FFB8FD02FF3FFCF697C2" box="[711,795,188,213]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD7FFB8FCE2FF3EFCA697C2" box="[807,843,189,212]" italics="true" pageId="4" pageNumber="169">pre</emphasis>
), sides of mesoventrite with setose ventrolateral and dorsolateral foveae (
<figureCitation id="13672A6EFFD7FFB8FE40FF5CFE2097EE" box="[389,461,223,248]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 9</figureCitation>
;
<emphasis id="B928EAF9FFD7FFB8FE1BFF63FE1797EE" box="[478,506,224,248]" italics="true" pageId="4" pageNumber="169">vlf</emphasis>
,
<emphasis id="B928EAF9FFD7FFB8FDC9FF63FDC497EE" box="[524,553,224,248]" italics="true" pageId="4" pageNumber="169">dlf</emphasis>
); mesocoxal projections (
<figureCitation id="13672A6EFFD7FFB8FC9CFF5CFC5C97EE" box="[857,945,223,248]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD7FFB8FC04FF62FC0297EE" box="[961,1007,225,248]" italics="true" pageId="4" pageNumber="169">mcp</emphasis>
) prominent, with mesocoxal sockets located on their mesoventral surface and exposed in ventral view.
</paragraph>
<paragraph id="8BE336EBFFD7FFB8FF02FEA4FBB9969E" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="169">
Metaventrite (
<figureCitation id="13672A6EFFD7FFB8FEAFFEA4FE2C9656" box="[362,449,295,320]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
;
<date id="FFE2102BFFD7FFB8FE14FEABFE019656" box="[465,492,296,320]" pageId="4" pageNumber="169">
<emphasis id="B928EAF9FFD7FFB8FE14FEABFE019656" box="[465,492,296,320]" italics="true" pageId="4" pageNumber="169">v3</emphasis>
</date>
) short, anteriorly fused with mesoventrite, posteriorly deeply bisinuate and with broadly subtriangular metaventral intercoxal process (
<figureCitation id="13672A6EFFD7FFB8FD37FECFFCA89672" box="[754,837,332,357]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
;
<emphasis id="B928EAF9FFD7FFB8FC94FECEFC6A9672" box="[849,903,333,356]" italics="true" pageId="4" pageNumber="169">mtvp</emphasis>
) bearing distinct inversely drop-shaped median notch. Metanepisterna and metepimera narrow, in intact specimens covered by elytra.
</paragraph>
<caption id="DF236663FFD7FFB8FF52F8C4FD1190CE" httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="4" pageNumber="169" targetBox="[151,1436,445,1842]" targetPageId="4">
<paragraph id="8BE336EBFFD7FFB8FF52F8C4FD1190CE" blockId="4.[151,1436,1863,2008]" pageId="4" pageNumber="169">
<emphasis id="B928EAF9FFD7FFB8FF52F8C4FEA6904A" bold="true" box="[151,331,1863,1886]" pageId="4" pageNumber="169">FIGURES 710.</emphasis>
<taxonomicName id="4C5C4D68FFD7FFB8FE91F8C4FDDB904B" box="[340,566,1863,1885]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD7FFB8FE91F8C4FDDB904B" box="[340,566,1863,1885]" italics="true" pageId="4" pageNumber="169">Bellendenker palpalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5782FFD7FFB8FD85F8CBFD79904B" box="[576,660,1864,1885]" pageId="4" pageNumber="169" rank="species">
<emphasis id="B928EAF9FFD7FFB8FD85F8CBFD79904B" bold="true" box="[576,660,1864,1885]" pageId="4" pageNumber="169">sp. nov.</emphasis>
</taxonomicNameLabel>
; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v3, metaventrite.
</paragraph>
</caption>
<caption id="DF236663FFD6FFB9FF52F8DBFB5F909B" httpUri="https://zenodo.org/record/438253/files/figure.png" pageId="5" pageNumber="170" targetBox="[151,1436,193,1859]" targetPageId="5">
<paragraph id="8BE336EBFFD6FFB9FF52F8DBFB5F909B" blockId="5.[151,1436,1880,1933]" pageId="5" pageNumber="170">
<emphasis id="B928EAF9FFD6FFB9FF52F8DBFEBA907B" bold="true" box="[151,343,1880,1903]" pageId="5" pageNumber="170">FIGURES 1113.</emphasis>
<taxonomicName id="4C5C4D68FFD6FFB9FEA7F8DBFDA89078" box="[354,581,1880,1902]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD6FFB9FEA7F8DBFDA89078" box="[354,581,1880,1902]" italics="true" pageId="5" pageNumber="170">Bellendenker palpalis</emphasis>
</taxonomicName>
<emphasis id="B928EAF9FFD6FFB9FD95F8DAFD489078" bold="true" box="[592,677,1881,1902]" pageId="5" pageNumber="170">
<taxonomicNameLabel id="A21B5782FFD6FFB9FD95F8DAFD489078" box="[592,677,1881,1902]" pageId="5" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; mesocoxa in dorsal view (11); setal tufts on mesocoxa (12); apex of protarsomere V in ventral view (13). Abbreviations: cx, coxa; emp, empodium; fe, femur; tr, trochanter.
</paragraph>
</caption>
<caption id="DF236663FFD5FFBAFF52FCC2FA6A9441" box="[151,1415,833,856]" httpUri="https://zenodo.org/record/438254/files/figure.png" pageId="6" pageNumber="171" targetBox="[163,1430,200,806]" targetPageId="6">
<paragraph id="8BE336EBFFD5FFBAFF52FCC2FA6A9441" blockId="6.[151,1415,833,856]" box="[151,1415,833,856]" pageId="6" pageNumber="171">
<emphasis id="B928EAF9FFD5FFBAFF52FCC2FEBE9440" bold="true" box="[151,339,833,856]" pageId="6" pageNumber="171">FIGURES 1418.</emphasis>
<taxonomicName id="4C5C4D68FFD5FFBAFE9FFCC2FDDA9441" box="[346,567,833,855]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="palpalis" status="sp. nov.">
<emphasis id="B928EAF9FFD5FFBAFE9FFCC2FDDA9441" box="[346,567,833,855]" italics="true" pageId="6" pageNumber="171">Bellendenker palpalis</emphasis>
</taxonomicName>
<emphasis id="B928EAF9FFD5FFBAFDF8FCC1FD609441" bold="true" box="[573,653,834,855]" pageId="6" pageNumber="171">
<taxonomicNameLabel id="A21B5782FFD5FFBAFDF8FCC1FD609441" box="[573,653,834,855]" pageId="6" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; aedeagus in ventral (14, 16), and lateral (15, 17) views; spermatheca (18).
</paragraph>
</caption>
<paragraph id="8BE336EBFFD5FFBAFF02FC07FDA794D6" blockId="6.[151,1437,900,2005]" pageId="6" pageNumber="171">
Elytra (
<figureCitation id="13672A6EFFD5FFBAFED9FC07FE88948B" box="[284,357,900,925]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,931,954]" captionTargetBox="[163,1436,193,905]" captionTargetId="figure@2.[151,1436,193,910]" captionTargetPageId="2" captionText="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." httpUri="https://zenodo.org/record/438250/files/figure.png" pageId="6" pageNumber="171">Figs 1</figureCitation>
,
<figureCitation id="13672A6EFFD5FFBAFEB0FC06FE69948B" box="[373,388,901,925]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="6" pageNumber="171">9</figureCitation>
) oval, strongly convex, lacking humeral calli and basal impressions, each with two asetose rudiments of basal foveae (
<figureCitation id="13672A6EFFD5FFBAFE03FC2BFDE694D6" box="[454,523,936,961]" captionStart="FIGURES 7 10" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[151,1436,445,1842]" captionTargetId="figure@4.[151,1436,445,1842]" captionTargetPageId="4" captionText="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." httpUri="https://zenodo.org/record/438252/files/figure.png" pageId="6" pageNumber="171">Fig. 7</figureCitation>
;
<emphasis id="B928EAF9FFD5FFBAFDDDFC2BFDD794D6" box="[536,570,936,960]" italics="true" pageId="6" pageNumber="171">bef</emphasis>
).
</paragraph>
<paragraph id="8BE336EBFFD5FFBAFF02FC4FFE7294F3" blockId="6.[151,1437,900,2005]" box="[199,415,972,997]" pageId="6" pageNumber="171">Hind wings absent.</paragraph>
<paragraph id="8BE336EBFFD5FFBAFF02FC72FEE2938E" blockId="6.[151,1437,900,2005]" pageId="6" pageNumber="171">
Legs (
<figureCitation id="13672A6EFFD5FFBAFED4FC73FE45931E" box="[273,424,1008,1033]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 6" captionStart-2="FIGURES 7 10" captionStart-3="FIGURES 11 13" captionStartId-0="2.[151,264,931,954]" captionStartId-1="3.[151,264,1732,1755]" captionStartId-2="4.[151,264,1863,1886]" captionStartId-3="5.[151,264,1880,1903]" captionTargetBox-0="[163,1436,193,905]" captionTargetBox-1="[151,1436,193,1712]" captionTargetBox-2="[151,1436,445,1842]" captionTargetBox-3="[151,1436,193,1859]" captionTargetId-0="figure@2.[151,1436,193,910]" captionTargetId-1="figure@3.[151,1436,193,1712]" captionTargetId-2="figure@4.[151,1436,445,1842]" captionTargetId-3="figure@5.[151,1436,193,1859]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1 2. Bellendenker palpalis sp. nov., holotype male; dorsal habitus (1) and distribution (2)." captionText-1="FIGURES 3 6. Bellendenker palpalis sp. nov.; head in dorsal (3), ventral (4) and lateral (5) views; mouthparts in ventral view (6). Abbreviations: cl, clypeus; fr, frons; gp, gular plate; hr, hypostomal ridge; lbr, labrum; lp, labial palp; lp 1 3, labial palpomere I III; mb, mandibular base; md, mandible; mn, mentum; mt, mesal tooth; mxp, maxillary palp; occ, occipital constriction; pmn, prementum; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." captionText-2="FIGURES 7 10. Bellendenker palpalis sp. nov.; prothorax in dorsal (7), ventral (8) and lateral (9) views; pterothorax in ventral view (10). Abbreviations: abp, antebasal pit; ar, anterior ridge; bef, basal elytral fovea; bst, basisternal part of prosternum; dlf, dorsolateral fovea; hy, hypomeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; nss, notosternal suture; pcc, procoxal cavity; pcr, procoxal rest; pre, prepectus; psp, prosternal process; vlf, ventrolateral fovea; v 3, metaventrite." captionText-3="FIGURES 11 13. Bellendenker palpalis sp. nov.; mesocoxa in dorsal view (11); setal tufts on mesocoxa (12); apex of protarsomere V in ventral view (13). Abbreviations: cx, coxa; emp, empodium; fe, femur; tr, trochanter." httpUri-0="https://zenodo.org/record/438250/files/figure.png" httpUri-1="https://zenodo.org/record/438251/files/figure.png" httpUri-2="https://zenodo.org/record/438252/files/figure.png" httpUri-3="https://zenodo.org/record/438253/files/figure.png" pageId="6" pageNumber="171">Figs 1, 113</figureCitation>
) long and robust; pro- and mesocoxae subglobose, metacoxae strongly transverse; all trochanters short and subtriangular; all femora strongly and gradually clavate; tibiae and tarsi moderately slender, terminal tarsomere with distinct empodium (
<figureCitation id="13672A6EFFD5FFBAFD6CFBBBFCEF9346" box="[681,770,1080,1105]" captionStart="FIGURES 11 13" captionStartId="5.[151,264,1880,1903]" captionTargetBox="[151,1436,193,1859]" captionTargetId="figure@5.[151,1436,193,1859]" captionTargetPageId="5" captionText="FIGURES 11 13. Bellendenker palpalis sp. nov.; mesocoxa in dorsal view (11); setal tufts on mesocoxa (12); apex of protarsomere V in ventral view (13). Abbreviations: cx, coxa; emp, empodium; fe, femur; tr, trochanter." httpUri="https://zenodo.org/record/438253/files/figure.png" pageId="6" pageNumber="171">Fig. 13</figureCitation>
; e
<emphasis id="B928EAF9FFD5FFBAFCE4FBBAFCAE9346" box="[801,835,1081,1104]" italics="true" pageId="6" pageNumber="171">mp</emphasis>
) emarginated distally and bearing two long setae. Mesocoxae (
<figureCitation id="13672A6EFFD5FFBAFEE0FBDFFE949363" box="[293,377,1116,1141]" captionStart="FIGURES 11 13" captionStartId="5.[151,264,1880,1903]" captionTargetBox="[151,1436,193,1859]" captionTargetId="figure@5.[151,1436,193,1859]" captionTargetPageId="5" captionText="FIGURES 11 13. Bellendenker palpalis sp. nov.; mesocoxa in dorsal view (11); setal tufts on mesocoxa (12); apex of protarsomere V in ventral view (13). Abbreviations: cx, coxa; emp, empodium; fe, femur; tr, trochanter." httpUri="https://zenodo.org/record/438253/files/figure.png" pageId="6" pageNumber="171">Figs 11</figureCitation>
¯12) with largely asetose dorsal surface (adjacent to mesocoxal cavity) bearing two bunches of long setae.
</paragraph>
<paragraph id="8BE336EBFFD5FFBAFF02FB27FC4C93F6" blockId="6.[151,1437,900,2005]" pageId="6" pageNumber="171">
Aedeagus (
<figureCitation id="13672A6EFFD5FFBAFE89FB27FE3993AB" box="[332,468,1188,1213]" captionStart="FIGURES 14 18" captionStartId="6.[151,264,833,856]" captionTargetBox="[163,1430,200,806]" captionTargetId="figure@6.[151,1436,193,812]" captionTargetPageId="6" captionText="FIGURES 14 18. Bellendenker palpalis sp. nov.; aedeagus in ventral (14, 16), and lateral (15, 17) views; spermatheca (18)." httpUri="https://zenodo.org/record/438254/files/figure.png" pageId="6" pageNumber="171">Figs 1417</figureCitation>
) elongate, thin-walled, with symmetrical median lobe and symmetrical, weakly sclerotized endophallus; parameres free and thick, lacking long setae.
</paragraph>
<paragraph id="8BE336EBFFD5FFBAFF02FB6FFC1D9213" blockId="6.[151,1437,900,2005]" box="[199,1008,1260,1285]" pageId="6" pageNumber="171">
Spermatheca (
<figureCitation id="13672A6EFFD5FFBAFEA2FB6FFE5A9213" box="[359,439,1260,1285]" captionStart="FIGURES 14 18" captionStartId="6.[151,264,833,856]" captionTargetBox="[163,1430,200,806]" captionTargetId="figure@6.[151,1436,193,812]" captionTargetPageId="6" captionText="FIGURES 14 18. Bellendenker palpalis sp. nov.; aedeagus in ventral (14, 16), and lateral (15, 17) views; spermatheca (18)." httpUri="https://zenodo.org/record/438254/files/figure.png" pageId="6" pageNumber="171">Fig. 18</figureCitation>
) globular, asymmetrical, moderately thick-walled.
</paragraph>
</subSubSection>
<subSubSection id="C3466560FFD5FFBAFF02FA93FAAF925B" pageId="6" pageNumber="171" type="etymology">
<paragraph id="8BE336EBFFD5FFBAFF02FA93FAAF925B" blockId="6.[151,1437,900,2005]" pageId="6" pageNumber="171">
<emphasis id="B928EAF9FFD5FFBAFF02FA93FEA3923F" bold="true" box="[199,334,1296,1321]" pageId="6" pageNumber="171">Etymology.</emphasis>
<taxonomicName id="4C5C4D68FFD5FFBAFE93FA93FE07923E" box="[342,490,1296,1320]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFE93FA93FE07923E" box="[342,490,1296,1320]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
is named after the
<typeStatus id="54E78849FFD5FFBAFD0CFA92FD14923F" box="[713,761,1297,1321]" pageId="6" pageNumber="171">type</typeStatus>
locality, Mt. Bellenden Ker in
<collectingRegion id="4998F809FFD5FFBAFBA4FA93FB05923F" box="[1121,1256,1296,1321]" country="Australia" name="Queensland" pageId="6" pageNumber="171">Queensland</collectingRegion>
, which, in turn, was dedicated to an English botanist of the XVIIIXIX c., John Bellenden Ker Gawler. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3466560FFD5FFBBFF02FADBFC8F969E" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171" type="discussion">
<paragraph id="8BE336EBFFD5FFBBFF02FADBFC8F969E" blockId="6.[151,1437,900,2005]" lastBlockId="7.[151,1436,151,392]" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171">
<emphasis id="B928EAF9FFD5FFBAFF02FADBFED69267" bold="true" box="[199,315,1368,1393]" pageId="6" pageNumber="171">Remarks.</emphasis>
<taxonomicName id="4C5C4D68FFD5FFBAFE87FADBFE3B9266" box="[322,470,1368,1392]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFE87FADBFE3B9266" box="[322,470,1368,1392]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
differs from all genera of
<taxonomicName id="4C5C4D68FFD5FFBAFD3BFADBFC7B9267" box="[766,918,1368,1393]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
, except for
<taxonomicName id="4C5C4D68FFD5FFBAFBDFFADBFB2F9266" box="[1050,1218,1368,1392]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFBDFFADBFB2F9266" box="[1050,1218,1368,1392]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
, in peculiar ventral structures of the prothorax: the adcoxal portion of each hypomeron forms an elongate subrectangular lobe with its anterior margin perpendicular in relation to the notosternal suture, and the lobe is not demarcated from the outer portion of hypomeron (i.e., lacking hypomeral ridge). This structure can be found only in
<taxonomicName id="4C5C4D68FFD5FFBAFB16FA47FA8A92CA" box="[1235,1383,1476,1500]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFB16FA47FA8A92CA" box="[1235,1383,1476,1500]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
and Australian
<taxonomicName id="4C5C4D68FFD5FFBAFED6FA6BFE569116" box="[275,443,1512,1536]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFED6FA6BFE569116" box="[275,443,1512,1536]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
; these genera share all prothoracic, most pterothoracic structures, and two tufts of setae on the dorsal surface of mesocoxae. These are almost certainly sister taxa, but they differ in a number of easily observable features: i) the clypeus in
<taxonomicName id="4C5C4D68FFD5FFBAFDFDF9B3FD21915E" box="[568,716,1584,1608]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFDFDF9B3FD21915E" box="[568,716,1584,1608]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
is strongly elongate, much longer than labrum, not declined and not demarcated from frons (in
<taxonomicName id="4C5C4D68FFD5FFBAFDC6F9D7FD46917A" box="[515,683,1620,1644]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFDC6F9D7FD46917A" box="[515,683,1620,1644]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
clypeus much shorter than labrum, declined and demarcated by distinct frontoclypeal groove); ii) the distance between compound eye and mandibular base in
<taxonomicName id="4C5C4D68FFD5FFBAFB2EF9FBFA929186" box="[1259,1407,1656,1680]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFB2EF9FBFA929186" box="[1259,1407,1656,1680]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
is about twice as wide as that between eye and posterior margin of vertex (in
<taxonomicName id="4C5C4D68FFD5FFBAFC1DF91FFB6D91A2" box="[984,1152,1692,1716]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFC1DF91FFB6D91A2" box="[984,1152,1692,1716]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
the distance between eye and mandibular base is about as wide or slightly shorter than that between eye and posterior margin of vertex); iii) the mandibles in
<taxonomicName id="4C5C4D68FFD5FFBAFE9EF967FE0291EA" box="[347,495,1764,1788]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFE9EF967FE0291EA" box="[347,495,1764,1788]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
are extremely slender, with the mesal tooth located in basal 1/3 (in
<taxonomicName id="4C5C4D68FFD5FFBAFB31F967FA7191EA" box="[1268,1436,1764,1788]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFB31F967FA7191EA" box="[1268,1436,1764,1788]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
mandibles short and broad, with mesal tooth located in distal 1/4); iv) the labial palpomeres II and III in
<taxonomicName id="4C5C4D68FFD5FFBAFF52F8AFFEC69052" box="[151,299,1836,1860]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFF52F8AFFEC69052" box="[151,299,1836,1860]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
are strikingly elongate, together much longer than entire labium (in
<taxonomicName id="4C5C4D68FFD5FFBAFBE7F8AFFB279052" box="[1058,1226,1836,1860]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFBE7F8AFFB279052" box="[1058,1226,1836,1860]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
labial palps do not differ from those in all remaining
<taxonomicName id="4C5C4D68FFD5FFBAFDE1F8D3FD51907F" box="[548,700,1872,1897]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
, i.e., they are shorter than labium); v) the hypostomal ridges in
<taxonomicName id="4C5C4D68FFD5FFBAFF52F8F7FEC6909A" box="[151,299,1908,1932]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFF52F8F7FEC6909A" box="[151,299,1908,1932]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
are strongly curved mesally (in
<taxonomicName id="4C5C4D68FFD5FFBAFD53F8F7FCD3909A" box="[662,830,1908,1932]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFD53F8F7FCD3909A" box="[662,830,1908,1932]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
straight, extending posteromesally); vi) the labrum in
<taxonomicName id="4C5C4D68FFD5FFBAFF52F81BFEC690A6" box="[151,299,1944,1968]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFF52F81BFEC690A6" box="[151,299,1944,1968]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
is covered densely with more than 50 asymmetrically distributed setae (in
<taxonomicName id="4C5C4D68FFD5FFBAFBB0F81BFAF090A6" box="[1141,1309,1944,1968]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD5FFBAFBB0F81BFAF090A6" box="[1141,1309,1944,1968]" italics="true" pageId="6" pageNumber="171">Scydmaenozila</emphasis>
</taxonomicName>
only about 15 mostly symmetrically distributed setae); vii) in
<taxonomicName id="4C5C4D68FFD5FFBAFD09F83FFC8D90C2" box="[716,864,1980,2004]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD5FFBAFD09F83FFC8D90C2" box="[716,864,1980,2004]" italics="true" pageId="6" pageNumber="171">Bellendenker</emphasis>
</taxonomicName>
the basal elytral foveae are rudimentary and barely discernible in intact specimens, basal elytral impressions are absent, humeral calli are rounded and indistinct, and wings absent (in
<taxonomicName id="4C5C4D68FFD4FFBBFEA1FF3FFDE197C2" box="[356,524,188,212]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD4FFBBFEA1FF3FFDE197C2" box="[356,524,188,212]" italics="true" pageId="7" pageNumber="172">Scydmaenozila</emphasis>
</taxonomicName>
elytral foveae are deep and large, located in distinct basal impressions, well discernible in intact beetles, humeral calli prominent, and hind wings well developed); viii) in
<taxonomicName id="4C5C4D68FFD4FFBBFB19FF63FA9D97EE" box="[1244,1392,224,248]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD4FFBBFB19FF63FA9D97EE" box="[1244,1392,224,248]" italics="true" pageId="7" pageNumber="172">Bellendenker</emphasis>
</taxonomicName>
the metaventral intercoxal process is deeply notched at middle, the notch is inversely drop-shaped (in
<taxonomicName id="4C5C4D68FFD4FFBBFB31FE87FA71960A" box="[1268,1436,260,284]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD4FFBBFB31FE87FA71960A" box="[1268,1436,260,284]" italics="true" pageId="7" pageNumber="172">Scydmaenozila</emphasis>
</taxonomicName>
posterior margin of metaventral process is indistinctly, broadly emarginate); ix) in
<taxonomicName id="4C5C4D68FFD4FFBBFBF3FEABFB279656" box="[1078,1226,296,320]" class="Insecta" family="Staphylinidae" genus="Bellendenker" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF9FFD4FFBBFBF3FEABFB279656" box="[1078,1226,296,320]" italics="true" pageId="7" pageNumber="172">Bellendenker</emphasis>
</taxonomicName>
the endophallus is simple, composed of weakly sclerotized structures (in
<taxonomicName id="4C5C4D68FFD4FFBBFD33FECFFC739672" box="[758,926,332,356]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF9FFD4FFBBFD33FECFFC739672" box="[758,926,332,356]" italics="true" pageId="7" pageNumber="172">Scydmaenozila</emphasis>
</taxonomicName>
the endophallus is complicated, composed of a large number of at least partly strongly sclerotized structures).
</paragraph>
</subSubSection>
</treatment>
</document>