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<mods:title id="C9B44A65B8440A5896D3EDB47257C167">Further addition to the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae), with new genera and species records, and the description of two new species</mods:title>
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<mods:namePart id="C641E7B8472583CCDDC40B0C30866855">Abu El-Ghiet, Usama M.</mods:namePart>
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<heading id="D0AB818BFF94641EFF52F970FBD2F900" bold="true" box="[151,1150,1716,1743]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
<taxonomicName id="4C5C4D64FF94641EFF52F970FBD2F900" ID-CoL="6K9VM" ID-ENA="2495025" authority="Pulawski in de Beaumont, Bytinski-Salz &amp; Pulawski, 1973" authorityName="Pulawski in de Beaumont, Bytinski-Salz &amp; Pulawski" authorityYear="1973" box="[151,1150,1716,1743]" class="Insecta" family="Crabronidae" genus="Gastrosericus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="sanctus">
<emphasis id="B928EAF5FF94641EFF52F970FBD2F900" bold="true" box="[151,1150,1716,1743]" pageId="9" pageNumber="160">
<emphasis id="B928EAF5FF94641EFF52F970FE0CF900" bold="true" box="[151,416,1717,1743]" italics="true" pageId="9" pageNumber="160">Gastrosericus sanctus</emphasis>
Pulawski
<emphasis id="B928EAF5FF94641EFDE4F970FD95F900" bold="true" box="[545,569,1717,1743]" italics="true" pageId="9" pageNumber="160">in</emphasis>
<bibRefCitation id="EFCD4B16FF94641EFD84F970FBD2F900" author="de Beaumont, J. &amp; Bytinski-Salz, H. &amp; Pulawski, W." box="[577,1150,1716,1743]" pageId="9" pageNumber="160" pagination="1 - 26" refId="ref8371" refString="de Beaumont, J., Bytinski-Salz, H. &amp; Pulawski, W. (1973) The Sphecidae (Hym.) of Eretz Israel III. Subfamilies: Astatinae, Larrinae, Trypoxyloninae, Pemphredoninae, Crabronidae, Oxybelinae. Israel Journal of Entomology, 8, 1 - 26." type="journal article" year="1973">de Beaumont, Bytinski-Salz &amp; Pulawski, 1973</bibRefCitation>
</emphasis>
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<subSubSection id="C346656CFF94641EFF52F91CFEA3F93C" box="[151,271,1753,1779]" pageId="9" pageNumber="160" type="description">
<paragraph id="8BE336E7FF94641EFF52F91CFEA3F93C" blockId="9.[151,1150,1716,1779]" box="[151,271,1753,1779]" pageId="9" pageNumber="160">
<figureCitation id="13672A62FF94641EFF52F91CFEA3F93C" box="[151,271,1753,1779]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="9" pageNumber="160">Fig. 7AG</figureCitation>
</paragraph>
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<emphasis id="B928EAF5FF94641EFF52F8DBFEDBF8FA" box="[151,375,1822,1845]" italics="true" pageId="9" pageNumber="160">Gastrosericus sanctus</emphasis>
Pulawski
<emphasis id="B928EAF5FF94641EFE25F8DBFE58F8FA" box="[480,500,1822,1845]" italics="true" pageId="9" pageNumber="160">in</emphasis>
<bibRefCitation id="EFCD4B16FF94641EFE3FF8D8FC50F8FA" author="de Beaumont, J. &amp; Bytinski-Salz, H. &amp; Pulawski, W." box="[506,1020,1821,1845]" pageId="9" pageNumber="160" pagination="1 - 26" refId="ref8371" refString="de Beaumont, J., Bytinski-Salz, H. &amp; Pulawski, W. (1973) The Sphecidae (Hym.) of Eretz Israel III. Subfamilies: Astatinae, Larrinae, Trypoxyloninae, Pemphredoninae, Crabronidae, Oxybelinae. Israel Journal of Entomology, 8, 1 - 26." type="journal article" year="1973">de Beaumont, Bytinski-Salz &amp; Pulawski, 1973: 16</bibRefCitation>
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,
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, ♁.
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<emphasis id="B928EAF5FF94641EFF52F8A6FEBEF8B2" bold="true" box="[151,274,1891,1917]" pageId="9" pageNumber="160">Diagnosis.</emphasis>
Male: body length: 7.0 mm. Body colour: see
<figureCitation id="13672A62FF94641EFE49F842FE61F86D" box="[396,461,1927,1954]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="9" pageNumber="160">Fig. 7</figureCitation>
(AF).
</paragraph>
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<paragraph id="8BE336E7FF97641DFF5DF852FAC0F81B" blockId="10.[151,1436,1943,2009]" pageId="10" pageNumber="161">
<emphasis id="B928EAF5FF97641DFF5DF852FEB4F87F" bold="true" box="[152,280,1943,1968]" pageId="10" pageNumber="161">FIGURE 7.</emphasis>
<taxonomicName id="4C5C4D64FF97641DFEDBF85DFE51F860" authorityName="Pulawski in de Beaumont, Bytinski-Salz &amp; Pulawski" authorityYear="1973" box="[286,509,1944,1967]" class="Insecta" family="Crabronidae" genus="Gastrosericus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="sanctus">
<emphasis id="B928EAF5FF97641DFEDBF85DFE51F860" box="[286,509,1944,1967]" italics="true" pageId="10" pageNumber="161">Gastrosericus sanctus</emphasis>
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(♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S
<subScript id="17D834A2FF97641DFB80F80EFBE0F816" attach="left" box="[1093,1100,1995,2009]" fontSize="6" pageId="10" pageNumber="161">3</subScript>
and S
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); F, last gastral terga.
</paragraph>
</caption>
<paragraph id="8BE336E7FF96641CFF02FF52FBF6FEAA" blockId="11.[151,1436,151,645]" pageId="11" pageNumber="162">
The male of this species shares with that of
<taxonomicName id="4C5C4D64FF96641CFD6EFF52FC8FFF7F" authorityName="E. Saunders" authorityYear="1910" box="[683,803,151,177]" class="Insecta" family="Crabronidae" genus="Gastrosericus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B928EAF5FF96641CFD6EFF52FC8FFF7F" box="[683,803,151,177]" italics="true" pageId="11" pageNumber="162">G. moricei</emphasis>
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the appressed genal (
<figureCitation id="13672A62FF96641CFBD4FF52FBC4FF7E" box="[1041,1128,151,177]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7D</figureCitation>
) and propodeal pubescence (
<figureCitation id="13672A62FF96641CFF5AFF7EFF54FF1A" box="[159,248,187,213]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7C</figureCitation>
); sternal fimbriae of S
<subScript id="17D834A2FF96641CFE37FF08FE53FF14" attach="left" box="[498,511,205,219]" fontSize="6" pageId="11" pageNumber="162">3,</subScript>
S
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noticeably long, slightly extending beyond posterior margin of sterna (
<figureCitation id="13672A62FF96641CFAFDFF7EFA23FF1A" box="[1336,1423,187,213]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7E</figureCitation>
); gastral terga without yellow bands (
<figureCitation id="13672A62FF96641CFDE2FF1AFD2CFF36" box="[551,640,223,249]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7A</figureCitation>
). It differs from that of
<taxonomicName id="4C5C4D64FF96641CFC42FF1AFBACFF37" authorityName="E. Saunders" authorityYear="1910" box="[903,1024,223,249]" class="Insecta" family="Crabronidae" genus="Gastrosericus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B928EAF5FF96641CFC42FF1AFBACFF37" box="[903,1024,223,249]" italics="true" pageId="11" pageNumber="162">G. moricei</emphasis>
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in the following: clypeal lobe broad, with arcuate to slightly sinuate free margin (
<figureCitation id="13672A62FF96641CFD49FEC6FD4EFED2" box="[652,738,259,285]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7B</figureCitation>
) and well-defined corner (obtusely pointed in
<taxonomicName id="4C5C4D64FF96641CFB34FEC6FAC6FED3" authorityName="E. Saunders" authorityYear="1910" box="[1265,1386,259,285]" class="Insecta" family="Crabronidae" genus="Gastrosericus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B928EAF5FF96641CFB34FEC6FAC6FED3" box="[1265,1386,259,285]" italics="true" pageId="11" pageNumber="162">G. moricei</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFCD4B16FF96641CFF52FEE2FEC5FE8E" author="Pulawski, W. J." box="[151,361,294,321]" pageId="11" pageNumber="162" pagination="1 - 173" refId="ref9639" refString="Pulawski, W. J. (1995) The wasp genus Gastrosericus Spinola, 1839 (Hymenoptera: Sphecidae). Memoirs of the California Academy of Sciences, 18, i - vi + 1 - 173." type="journal article" year="1995">Pulawski 1995: 94</bibRefCitation>
, fig. 78f, h); S
<subScript id="17D834A2FF96641CFDCFFEFCFDBEFE88" attach="left" box="[522,530,313,327]" fontSize="6" pageId="11" pageNumber="162">3</subScript>
and S
<subScript id="17D834A2FF96641CFD90FEFCFDF1FE88" attach="left" box="[597,605,313,327]" fontSize="6" pageId="11" pageNumber="162">4</subScript>
depressed medially (
<figureCitation id="13672A62FF96641CFC8DFEE2FC32FE8E" box="[840,926,295,321]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[184,1404,181,1920]" captionTargetId="figure-20@10.[184,1404,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Gastrosericus sanctus (♁): A, dorsal habitus; B, frontal view of head; C, dorsal view of head and thorax; D, lateral view of head and thorax; E, ventral view of gaster (showing large fimbriate impressions on S3 and S4); F, last gastral terga." figureDoi="http://doi.org/10.5281/zenodo.8182337" httpUri="https://zenodo.org/record/8182337/files/figure.png" pageId="11" pageNumber="162">Fig. 7E</figureCitation>
), not fimbriate laterally (fimbriae of S
<subScript id="17D834A2FF96641CFA8CFEFCFAFDFE88" attach="left" box="[1353,1361,313,327]" fontSize="6" pageId="11" pageNumber="162">3</subScript>
and S
<subScript id="17D834A2FF96641CFA51FEFCFA30FE88" attach="left" box="[1428,1436,313,327]" fontSize="6" pageId="11" pageNumber="162">4</subScript>
present side to side, sterna not depressed medially, see
<bibRefCitation id="EFCD4B16FF96641CFD3BFE8EFC7CFEAA" author="Pulawski, W. J." box="[766,976,330,357]" pageId="11" pageNumber="162" pagination="1 - 173" refId="ref9639" refString="Pulawski, W. J. (1995) The wasp genus Gastrosericus Spinola, 1839 (Hymenoptera: Sphecidae). Memoirs of the California Academy of Sciences, 18, i - vi + 1 - 173." type="journal article" year="1995">Pulawski 1995: 95</bibRefCitation>
, fig. 79e, f).
</paragraph>
</subSubSection>
<subSubSection id="C346656CFF96641CFF02FEABFC42FE62" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8BE336E7FF96641CFF02FEABFAF6FE46" blockId="11.[151,1436,151,645]" box="[199,1370,366,395]" pageId="11" pageNumber="162">
<emphasis id="B928EAF5FF96641CFF02FEABFE1FFE47" bold="true" box="[199,435,366,392]" pageId="11" pageNumber="162">Material examined:</emphasis>
2 ♁, Farasan, Abdulmajeed farm [16
<superScript id="7C299BAFFF96641CFC91FEABFCF2FEB3" attach="left" box="[852,862,366,380]" fontSize="6" pageId="11" pageNumber="162">Q</superScript>
42.07970N; 42
<superScript id="7C299BAFFF96641CFBD2FEABFB8DFEB3" attach="left" box="[1047,1057,366,380]" fontSize="6" pageId="11" pageNumber="162">Q</superScript>
08.5781E],
<date id="FFE21027FF96641CFB74FEAAFAFAFE46" box="[1201,1366,367,393]" pageId="11" pageNumber="162" value="2022-09-23" valueMax="2022-09-30" valueMin="2022-09-23">2330.ix.2022</date>
.
</paragraph>
<paragraph id="8BE336E7FF96641CFF02FE57FC42FE62" blockId="11.[151,1436,151,645]" box="[199,1006,402,429]" pageId="11" pageNumber="162">
<emphasis id="B928EAF5FF96641CFF02FE57FDE1FE62" bold="true" box="[199,589,402,429]" pageId="11" pageNumber="162">Previous Saudi Arabian records:</emphasis>
Abu Arish,
<collectingRegion id="4998F805FF96641CFD13FE56FC8BFE62" box="[726,807,403,429]" country="Saudi Arabia" name="Ar Riyad" pageId="11" pageNumber="162">Riyadh</collectingRegion>
(
<bibRefCitation id="EFCD4B16FF96641CFCF3FE56FC4DFE62" author="Pulawski, W. J." box="[822,993,402,429]" pageId="11" pageNumber="162" pagination="1 - 173" refId="ref9639" refString="Pulawski, W. J. (1995) The wasp genus Gastrosericus Spinola, 1839 (Hymenoptera: Sphecidae). Memoirs of the California Academy of Sciences, 18, i - vi + 1 - 173." type="journal article" year="1995">Pulawski 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346656CFF96641CFF02FE73FEBBFDD6" pageId="11" pageNumber="162" type="distribution">
<paragraph id="8BE336E7FF96641CFF02FE73FEBBFDD6" blockId="11.[151,1436,151,645]" pageId="11" pageNumber="162">
<emphasis id="B928EAF5FF96641CFF02FE73FECFFE1F" bold="true" box="[199,355,438,464]" pageId="11" pageNumber="162">Distribution:</emphasis>
<collectingCountry id="F34B7677FF96641CFEABFE72FDA7FE1E" box="[366,523,439,465]" name="Burkina Faso" pageId="11" pageNumber="162">Burkina Faso</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFDDFFE72FDCAFE1E" box="[538,614,439,465]" name="Ghana" pageId="11" pageNumber="162">Ghana</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFDB0FE72FD1AFE1E" box="[629,694,439,465]" name="Israel" pageId="11" pageNumber="162">Israel</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFD00FE72FCFCFE1E" box="[709,848,439,465]" name="Ivory Coast" pageId="11" pageNumber="162">Ivory Coast</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFC9AFE72FC07FE1E" box="[863,939,439,465]" name="Kenya" pageId="11" pageNumber="162">Kenya</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFC7FFE72FC5EFE1E" box="[954,1010,439,465]" name="Serbia" pageId="11" pageNumber="162">Mali</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFBC4FE72FBD3FE1E" box="[1025,1151,439,465]" name="Mauritania" pageId="11" pageNumber="162">Mauritania</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFB4BFE72FB5DFE1E" box="[1166,1265,439,465]" name="Namibia" pageId="11" pageNumber="162">Namibia</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFAC5FE72FAEEFE1E" box="[1280,1346,439,465]" name="Niger" pageId="11" pageNumber="162">Niger</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFA94FE72FA34FE1E" box="[1361,1432,439,465]" name="Oman" pageId="11" pageNumber="162">Oman</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFF52FE1EFF55FE3A" box="[151,249,475,501]" name="Pakistan" pageId="11" pageNumber="162">Pakistan</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFECBFE1EFE01FE3A" box="[270,429,475,501]" name="Saudi Arabia" pageId="11" pageNumber="162">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFE06FE1EFDB2FE3A" box="[451,542,475,501]" name="Senegal" pageId="11" pageNumber="162">Senegal</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFDF1FE1EFD3FFE3A" box="[564,659,475,501]" name="Somalia" pageId="11" pageNumber="162">Somalia</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFD6DFE1EFCEFFE3A" box="[680,835,475,501]" name="South Africa" pageId="11" pageNumber="162">South Africa</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFC9CFE1EFC6DFE3A" box="[857,961,475,501]" name="Tanzania" pageId="11" pageNumber="162">Tanzania</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFC10FE1EFBBDFE3A" box="[981,1041,475,501]" name="Togo" pageId="11" pageNumber="162">Togo</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFBE2FE1EFA9EFE3A" box="[1063,1330,475,501]" name="United Arab Emirates" pageId="11" pageNumber="162">United Arab Emirates</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFA82FE1EFA34FE3A" box="[1351,1432,475,501]" name="Yemen" pageId="11" pageNumber="162">Yemen</collectingCountry>
,
<collectingCountry id="F34B7677FF96641CFF52FE3AFEBEFDD6" box="[151,274,511,537]" name="Zimbabwe" pageId="11" pageNumber="162">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C346656CFF96641CFF02FDE7FE12FD4A" pageId="11" pageNumber="162" type="discussion">
<paragraph id="8BE336E7FF96641CFF02FDE7FE12FD4A" blockId="11.[151,1436,151,645]" pageId="11" pageNumber="162">
<emphasis id="B928EAF5FF96641CFF02FDE7FEE3FDF3" bold="true" box="[199,335,546,572]" pageId="11" pageNumber="162">Comments.</emphasis>
Characters of the collected specimens of this species agree with Pulawskis key (1995: 23, couplet 24). It also agrees with Schmid-Egger &amp; van Hartens key (2022: 547, couplet 9), but gastral terga of male black, reddish brown posteriorly.
</paragraph>
</subSubSection>
</treatment>
</document>