381 lines
61 KiB
XML
381 lines
61 KiB
XML
<document id="0B526535806B6A3888714D6C8B3A3340" ID-DOI="10.5281/zenodo.191909" ID-GBIF-Dataset="b5235354-ef06-4a49-b9d1-5498e97c3639" ID-ISSN="1175-5326" ID-Zenodo-Dep="191909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461137864308" checkinUser="plazi" docAuthor="Ng, Peter K. L., Rahayu, Dwi Listyo & Naser, Murtada D." docDate="2009" docId="03F58791D32A0C1FC9CF9D32FD16F8AA" docLanguage="en" docName="zt02312p026.pdf" docOrigin="Zootaxa 2312" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nasima Manning 1991" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FFCCFFE9D3200C12C9589874FFC1FFD4" masterDocTitle="The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat & Jones, 1996 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="11" updateTime="1698598725934" updateUser="plazi">
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<mods:titleInfo id="EB24BA3E5D243C2A930720307D7C10D8">
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<mods:title id="8611C72AA347580FFD84F312061F082D">The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat & Jones, 1996 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="01AEBBC0F85B6D9E3022510218539BF1">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="02E285AAA1066B50ACCDEC378D6FAFED">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="D0D2ECF6063E0C6E0219BBA2598E5A6E">Naser, Murtada D.</mods:namePart>
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<mods:typeOfResource id="3262793FEE41114C1BC83957BB3FFE1D">text</mods:typeOfResource>
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<mods:title id="2CEDD93824A5BCDF865AF50272E1F4B2">Zootaxa</mods:title>
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<mods:part id="82993BE180E72B71489B5B49119453AC">
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<mods:date id="3B48B0E494FB6CAC5FE2554C2AB3ECC5">2009</mods:date>
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<mods:detail id="F0AA2D8280387CE53B3011510C3A7FB4" type="volume">
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<mods:number id="F2566DAE1302771F43B1892705C7A5F8">2312</mods:number>
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<mods:classification id="B251DE5E4DBD8331B114E92FBDE9DF99">journal article</mods:classification>
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<mods:identifier id="C56E3F4B274FDFB2088C24B51638AAE4" type="DOI">10.5281/zenodo.191909</mods:identifier>
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<mods:identifier id="4B1667C7581C95C14AA01DA022B3A4BE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CDA77E3E4757968DB85A94CDA4524A13" type="Zenodo-Dep">191909</mods:identifier>
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<treatment id="03F58791D32A0C1FC9CF9D32FD16F8AA" ID-DOI="http://doi.org/10.5281/zenodo.6226793" ID-GBIF-Taxon="119627620" ID-Zenodo-Dep="6226793" LSID="urn:lsid:plazi:treatment:03F58791D32A0C1FC9CF9D32FD16F8AA" httpUri="http://treatment.plazi.org/id/03F58791D32A0C1FC9CF9D32FD16F8AA" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<subSubSection id="C346650CD32A0C18C9CF9D32FE77FAB4" box="[151,438,1350,1376]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BE33687D32A0C18C9CF9D32FE77FAB4" blockId="10.[151,438,1350,1376]" box="[151,438,1350,1376]" pageId="10" pageNumber="11">
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<heading id="D0AB81EBD32A0C18C9CF9D32FE77FAB4" bold="true" box="[151,438,1350,1376]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<taxonomicName id="4C5C4D04D32A0C18C9CF9D32FE77FAB4" ID-CoL="7PPS4" authority="Manning, 1991" authorityName="Manning" authorityYear="1991" box="[151,438,1350,1376]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B928EA95D32A0C18C9CF9D32FE77FAB4" bold="true" box="[151,438,1350,1376]" pageId="10" pageNumber="11">
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<emphasis id="B928EA95D32A0C18C9CF9D32FF35FAB4" bold="true" box="[151,244,1350,1376]" italics="true" pageId="10" pageNumber="11">Nasima</emphasis>
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<bibRefCitation id="EFCD4B76D32A0C18C9A39D32FE77FAB4" author="Manning" box="[251,438,1350,1376]" pageId="10" pageNumber="11" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning, 1991</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346650CD32A0C18C9CF9DE4FD04FA73" box="[151,709,1424,1447]" pageId="10" pageNumber="11" type="reference_group">
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<paragraph id="8BE33687D32A0C18C9CF9DE4FD04FA73" blockId="10.[151,709,1424,1447]" box="[151,709,1424,1447]" pageId="10" pageNumber="11">
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<treatmentCitationGroup id="AB4C11A9D32A0C18C9CF9DE4FD04FA73" box="[151,709,1424,1447]" pageId="10" pageNumber="11">
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<taxonomicName id="4C5C4D04D32A0C18C9CF9DE4FE0AFA73" ID-CoL="7PPS4" authority="Manning, 1991: 304" authorityName="Manning" authorityPageNumber="304" authorityYear="1991" box="[151,459,1424,1447]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B928EA95D32A0C18C9CF9DE4FF28FA73" box="[151,233,1424,1447]" italics="true" pageId="10" pageNumber="11">Nasima</emphasis>
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<treatmentCitation id="0AFD1096D32A0C18C9B79DE4FE0AFA73" author="Manning" box="[239,459,1424,1447]" page="304" pageId="10" pageNumber="11" year="1991">
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<bibRefCitation id="EFCD4B76D32A0C18C9B79DE4FE54FA73" author="Manning" box="[239,405,1424,1447]" pageId="10" pageNumber="11" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning, 1991</bibRefCitation>
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: 304
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</treatmentCitation>
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</taxonomicName>
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. —Ng
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<emphasis id="B928EA95D32A0C18CB4F9DE4FD8CFA73" box="[535,589,1424,1447]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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2008: 233.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346650CD32A0C18C9CF9DA2FC01FA24" box="[151,960,1494,1520]" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BE33687D32A0C18C9CF9DA2FC01FA24" blockId="10.[151,1437,1494,2023]" box="[151,960,1494,1520]" pageId="10" pageNumber="11">
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||
<emphasis id="B928EA95D32A0C18C9CF9DA2FEF8FA24" bold="true" box="[151,313,1494,1520]" pageId="10" pageNumber="11">
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<typeStatus id="54E78825D32A0C18C9CF9DA2FF15FA24" box="[151,212,1494,1520]" pageId="10" pageNumber="11">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C5C4D04D32A0C18C8189DA3FCD3FA24" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[320,786,1494,1520]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dotilliforme">
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<emphasis id="B928EA95D32A0C18C8189DA3FDAAFA24" box="[320,619,1495,1520]" italics="true" pageId="10" pageNumber="11">Cleistostoma dotilliforme</emphasis>
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<bibRefCitation id="EFCD4B76D32A0C18CB2A9DA2FCD3FA24" author="Alcock" box="[626,786,1494,1520]" pageId="10" pageNumber="11" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3), 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C346650CD32A0C1EC99E9D89FD1EFE98" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BE33687D32A0C1EC99E9D89FD1EFE98" blockId="10.[151,1437,1494,2023]" lastBlockId="12.[151,1437,152,332]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<emphasis id="B928EA95D32A0C18C99E9D89FE87F9C3" bold="true" box="[198,326,1533,1559]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
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Carapace average width to length ratio 1.44; subquadrate, moderately convex; regions of dorsal surface well developed, distinct, gastro-cardiac groove distinct, shallow; surface sparsely hairy, dorsolateral margin distinctly demarcated as raised row of granules; anterolateral margin with low, blunt lobe behind external orbital tooth; epigastric lobes present (
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<figureCitation id="13672A02D32A0C18CA149E05FC66F95F" box="[844,935,1649,1675]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="10" pageNumber="11">Figs. 5</figureCitation>
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,
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||
<figureCitation id="13672A02D32A0C18CAED9E05FC04F95F" box="[949,965,1649,1675]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">6</figureCitation>
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A). Front with broad, shallow groove medially on dorsal surface, with thickened margin; frontomedial margin bilobed (
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<figureCitation id="13672A02D32A0C18CDD29EECFB15F966" box="[1162,1236,1688,1714]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
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A). Supraorbital margin gradually curved, junction between base of front and supraorbital margin gradual, not sharply demarcated (
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<figureCitation id="13672A02D32A0C18C8649E91FE4AF92B" box="[316,395,1765,1791]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
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A); infraorbital margin straight along inner portion, outer portion concave, inner infraorbital tooth triangular, with rounded apex, not very distinctly separated from rest of infraorbital margin. Posterior edge of epistome thick, posterior medial tooth broadly triangular (
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<figureCitation id="13672A02D32A0C18CD519F46FB8EF898" box="[1033,1103,1842,1868]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
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B). Third maxilliped with Yshaped groove on surface; pterygostomian region with closed, shallow Y-shaped sulcus (
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<figureCitation id="13672A02D32A0C18CDEA9F2DFB3BF8A7" box="[1202,1274,1881,1907]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
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C). Chelipeds small, equal, similar in both sexes, tips spatulate, setose (
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<figureCitation id="13672A02D32A0C18CA179FF4FC56F84E" box="[847,919,1920,1946]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
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D, E). Meri of ambulatory legs foliaceous; anterior margins of carpus, propodus with dense seate; P3, P4 ventral margins lined with short sharp spines (
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<figureCitation id="13672A02D32A0C18C9F89FB9FF28F833" box="[160,233,1997,2023]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
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). Male abdomen narrow, somite 1 about as narrow as somite 2, not reaching P5 coxae; somite 5 not constricted, somites 2–5 fused, sutures between somites 3, 4 interrupted very faintly, visible as short lateral grooves, median transverse grooves, somites 4, 5 separated by distinct groove but immobile, somite 6 with limited mobility with somite 5; telson free (
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<figureCitation id="13672A02D32C0C1ECBFB9891FD2DFF2B" box="[675,748,229,255]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
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F); female with subcircular abdomen; somites 1–6, telson free. G1 recurved, lacking distal process, outer edge of recurved portion broadened to form low but distinct flat lobe halfway to tip, apex conical (
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<figureCitation id="13672A02D32C0C1ECB0B9946FD59FE98" box="[595,664,306,332]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
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G, H).
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF23660FD32B0C19C9CF9FC2FBEFF824" httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,185,1952]" targetPageId="11">
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<paragraph id="8BE33687D32B0C19C9CF9FC2FBEFF824" blockId="11.[151,1436,1974,2032]" pageId="11" pageNumber="12">
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<emphasis id="B928EA95D32B0C19C9CF9FC2FEDDF81A" bold="true" box="[151,284,1974,1998]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C5C4D04D32B0C19C87B9FC3FD75F819" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[291,692,1974,1998]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32B0C19C87B9FC3FDC5F81A" box="[291,516,1975,1998]" italics="true" pageId="11" pageNumber="12">Nasima dotilliformis</emphasis>
|
||
(Alcock, 1900)
|
||
</taxonomicName>
|
||
. A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF23660FD32C0C1EC9CF9EC5FF09F8E3" httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="12" pageNumber="13" targetBox="[172,1417,403,1640]" targetPageId="12">
|
||
<paragraph id="8BE33687D32C0C1EC9CF9EC5FF09F8E3" blockId="12.[151,1437,1713,1847]" pageId="12" pageNumber="13">
|
||
<emphasis id="B928EA95D32C0C1EC9CF9EC5FEDFF91D" bold="true" box="[151,286,1713,1737]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C5C4D04D32C0C1EC87D9EC6FD7DF91D" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[293,700,1714,1737]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32C0C1EC87D9EC6FDCBF91D" box="[293,522,1714,1737]" italics="true" pageId="12" pageNumber="13">Nasima dotilliformis</emphasis>
|
||
(Alcock, 1900)
|
||
</taxonomicName>
|
||
, male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G1; H, distal part of left G1. Scales: A–E, G = 1.0 mm; F, H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C346650CD32C0C1FC99E9F18FD16F8AA" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8BE33687D32C0C1FC99E9F18FCA2FE14" blockId="12.[151,1437,1900,2003]" lastBlockId="13.[151,1437,152,1918]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="B928EA95D32C0C1EC99E9F18FE81F852" bold="true" box="[198,320,1900,1926]" pageId="12" pageNumber="13">Remarks.</emphasis>
|
||
<bibRefCitation id="EFCD4B76D32C0C1EC80B9F18FDDEF852" author="Manning" box="[339,543,1900,1926]" pageId="12" pageNumber="13" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning (1991)</bibRefCitation>
|
||
established
|
||
<taxonomicName id="4C5C4D04D32C0C1ECB909F18FCE3F851" box="[712,802,1900,1925]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32C0C1ECB909F18FCE3F851" box="[712,802,1900,1925]" italics="true" pageId="12" pageNumber="13">Nasima</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<taxonomicName id="4C5C4D04D32C0C1ECA329F18FAA3F852" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[874,1378,1900,1926]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dotilliforme">
|
||
<emphasis id="B928EA95D32C0C1ECA329F18FB60F851" box="[874,1185,1900,1925]" italics="true" pageId="12" pageNumber="13">Cleistostoma dotilliforme</emphasis>
|
||
<bibRefCitation id="EFCD4B76D32C0C1ECDED9F18FAA3F852" author="Alcock" box="[1205,1378,1900,1926]" pageId="12" pageNumber="13" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3), 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
. He distinguished it from other genera mainly by the male abdomen having all six somites and telson free, and the chelipeds being of similar size in both sexes. Not all the somites of the male abdomen are actually free, however, although some of the sutures separating them remain visible; somites 2–5 are actually immobile. Somites 2-4 have only a shallow and barely discernible suture separating each other; while the suture between somites 4 and 5 suture is complete, though immobile (
|
||
<figureCitation id="13672A02D32D0C1FCA569891FC95FF2B" box="[782,852,229,255]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||
F). Somites 5 and 6 are also barely mobile, even if the suture is complete and appears deep.
|
||
<bibRefCitation id="EFCD4B76D32D0C1FCB259978FCFFFEF2" author="Manning" box="[637,830,268,294]" pageId="13" pageNumber="14" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning (1991)</bibRefCitation>
|
||
also diagnosed the genus as having the meri of the ambulatory legs without spines, but this is not completely correct since the ventral margins of meri of the P3 and P4 have numerous sharp granules that resemble short spines (
|
||
<figureCitation id="13672A02D32D0C1FCAF1992DFC33FEA7" box="[937,1010,345,371]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
|
||
). The lack of sexual dimorphism in the chelipeds is, however, useful, and several other characters, in addition to those mentioned by
|
||
<bibRefCitation id="EFCD4B76D32D0C1FCC6899F4FF26FE14" author="Manning" pageId="13" pageNumber="14" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning (1991)</bibRefCitation>
|
||
, make
|
||
<taxonomicName id="4C5C4D04D32D0C1FC86299D3FE55FE14" box="[314,404,423,448]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC86299D3FE55FE14" box="[314,404,423,448]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
generically distinct from
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB9B99D3FC9DFE14" box="[707,860,423,448]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCB9B99D3FC9DFE14" box="[707,860,423,448]" italics="true" pageId="13" pageNumber="14">Cleistostoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE33687D32D0C1FC99E99B9FF36FC63" blockId="13.[151,1437,152,1918]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C5C4D04D32D0C1FC99E99B9FE7BFE32" box="[198,442,461,486]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FC99E99B9FE7BFE32" box="[198,442,461,486]" italics="true" pageId="13" pageNumber="14">Nasima dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB0499B9FD31FE32" box="[604,752,461,486]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FCB0499B9FD31FE32" box="[604,752,461,486]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54E78825D32D0C1FCA5A99B9FCF7FE33" box="[770,822,461,487]" pageId="13" pageNumber="14">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA9899B9FB98FE32" box="[960,1113,461,486]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCA9899B9FB98FE32" box="[960,1113,461,486]" italics="true" pageId="13" pageNumber="14">Cleistostoma</emphasis>
|
||
</taxonomicName>
|
||
) in several key characters. The male abdominal somite 1 of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB7F9980FD7FFDD9" box="[551,702,500,525]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FCB7F9980FD7FFDD9" box="[551,702,500,525]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
is transversely elongated, with the ends overreaching the P5 coxae (
|
||
<figureCitation id="13672A02D32D0C1FC9B39A6EFEFEFDE0" box="[235,319,538,564]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
A). In
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8D39A6EFDFFFDE7" box="[395,574,538,563]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FC8D39A6EFDFFFDE7" box="[395,574,538,563]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
, male abdominal somite 1 is only almost as transversely wide as somite 2 and the ends before the P5 coxae (
|
||
<figureCitation id="13672A02D32D0C1FCB719A35FDBEFD8F" box="[553,639,577,603]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
C). The apex of the G
|
||
<quantity id="4CA49B62D32D0C1FCADA9A35FC6AFD8F" box="[898,939,577,603]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="14" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C5C4D04D32D0C1FCAEA9A35FB84FD8E" box="[946,1093,577,602]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FCAEA9A35FB84FD8E" box="[946,1093,577,602]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
is drawn out to form a point, with subapical mushroom-like tubercles on one edge (
|
||
<figureCitation id="13672A02D32D0C1FCA4C9A1CFCA6FD56" box="[788,871,616,642]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
E, F). The apex of the G
|
||
<quantity id="4CA49B62D32D0C1FCDD39A1CFB75FD56" box="[1163,1204,616,642]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="14" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C5C4D04D32D0C1FCDE49A1CFAAEFD55" box="[1212,1391,616,641]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FCDE49A1CFAAEFD55" box="[1212,1391,616,641]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
, on the other hand, is conical and lacks mushroom-like tubercles (
|
||
<figureCitation id="13672A02D32D0C1FCA339AFAFC6EFD7C" box="[875,943,654,680]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||
G, H). In addition, the low lobe present on the recurved portion of the G
|
||
<quantity id="4CA49B62D32D0C1FC8A69AC1FDE8FD1B" box="[510,553,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="14" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB699AC1FD2AFD1A" box="[561,747,693,718]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FCB699AC1FD2AFD1A" box="[561,747,693,718]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13672A02D32D0C1FCBA49AC1FC85FD1B" box="[764,836,693,719]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||
G, H) is absent in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCD799AC1FB79FD1A" box="[1057,1208,693,718]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FCD799AC1FB79FD1A" box="[1057,1208,693,718]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13672A02D32D0C1FCD929AC1FADEFD1B" box="[1226,1311,693,719]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
E, F). The sexual dimorphism of the chelipeds is very evident in adult
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA099AA8FC22FD21" box="[849,995,732,757]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FCA099AA8FC22FD21" box="[849,995,732,757]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
but is not apparent in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCDB19AA8FA5DFD21" box="[1257,1436,732,757]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FCDB19AA8FA5DFD21" box="[1257,1436,732,757]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13672A02D32D0C1FC9F89B76FF28FCC8" box="[160,233,770,796]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
|
||
). The first sternite of the thoracic sternum in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA549B76FC02FCCF" box="[780,963,770,795]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FCA549B76FC02FCCF" box="[780,963,770,795]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
has a rounded anterior edge (
|
||
<figureCitation id="13672A02D32D0C1FCC709B76FABEFCC8" box="[1320,1407,770,796]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
C) while the same structure in
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8A49B5DFD5DFC96" box="[508,668,809,834]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FC8A49B5DFD5DFC96" box="[508,668,809,834]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
is broadly triangular (
|
||
<figureCitation id="13672A02D32D0C1FCAE69B5DFBDDFC97" box="[958,1052,809,835]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones & Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
A). Furthermore, the carapace anterolateral margin in
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8F59B24FD80FCBD" box="[429,577,848,873]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dilatatum">
|
||
<emphasis id="B928EA95D32D0C1FC8F59B24FD80FCBD" box="[429,577,848,873]" italics="true" pageId="13" pageNumber="14">C. dilatatum</emphasis>
|
||
</taxonomicName>
|
||
is entire but in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCBA69B24FC75FCBD" box="[766,948,848,873]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32D0C1FCBA69B24FC75FCBD" box="[766,948,848,873]" italics="true" pageId="13" pageNumber="14">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
, there is a small lobe present just behind the external orbital tooth (
|
||
<figureCitation id="13672A02D32D0C1FC8889B02FDE4FC44" box="[464,549,886,912]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13672A02D32D0C1FCB6B9B02FD82FC44" box="[563,579,886,912]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">6</figureCitation>
|
||
A). These observations support Manning's (1991) action of establishing
|
||
<taxonomicName id="4C5C4D04D32D0C1FC9CF9BE9FF30FC62" box="[151,241,925,950]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC9CF9BE9FF30FC62" box="[151,241,925,950]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE33687D32D0C1FC99E9BB0FD16F8AA" blockId="13.[151,1437,152,1918]" pageId="13" pageNumber="14">
|
||
The relationship of
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8F79BB0FDC8FC09" box="[431,521,964,989]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC8F79BB0FDC8FC09" box="[431,521,964,989]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB159BB0FBF1FC0A" authority="Al-Khayat & Jones, 1996" authorityName="Al-Khayat & Jones" authorityYear="1996" box="[589,1072,964,990]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCB159BB0FD36FC09" box="[589,759,964,989]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
<bibRefCitation id="EFCD4B76D32D0C1FCBA79BB0FBF1FC0A" author="Al-Khayat" box="[767,1072,964,990]" pageId="13" pageNumber="14" refString="Al-Khayat, J. A. & Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat & Jones, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
, however, is problematic. This affinity has never been discussed before, even when
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA419B9EFC06FBD7" box="[793,967,1002,1027]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCA419B9EFC06FBD7" box="[793,967,1002,1027]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
was first described, but this may have been because several of the diagnostic features were misinterpreted by
|
||
<bibRefCitation id="EFCD4B76D32D0C1FCD4E9C65FAA3FBFF" author="Al-Khayat" box="[1046,1378,1041,1067]" pageId="13" pageNumber="14" refString="Al-Khayat, J. A. & Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat & Jones (1996)</bibRefCitation>
|
||
(see discussion for this genus later). It is noteworthy that
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA759C4CFC4AFB85" box="[813,907,1080,1105]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCA759C4CFC4AFB85" box="[813,907,1080,1105]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
shares the diagnostic male abdominal and thoracic sternal generic characters of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB179C2BFBE0FBAC" authority="Jones & Clayton, 1983" authorityName="Jones & Clayton" authorityYear="1983" box="[591,1057,1118,1144]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCB179C2BFD39FBAC" box="[591,760,1119,1144]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCD4B76D32D0C1FCA509C2AFBD8FBAC" author="Jones" box="[776,1049,1118,1144]" pageId="13" pageNumber="14" refString="Jones, D. A. & Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones & Clayton, 1983</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13672A02D32D0C1FCD699C2AFB43FBAC" box="[1073,1154,1118,1144]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">Figs. 6</figureCitation>
|
||
F, 7C, 9B, 10B, C). The G1s of
|
||
<taxonomicName id="4C5C4D04D32D0C1FC9A99CF1FE8CFB4A" box="[241,333,1157,1182]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC9A99CF1FE8CFB4A" box="[241,333,1157,1182]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8D09CF1FDF7FB4A" box="[392,566,1157,1182]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC8D09CF1FDF7FB4A" box="[392,566,1157,1182]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
are also very similar (
|
||
<figureCitation id="13672A02D32D0C1FCA1F9CF1FC5BFB4B" box="[839,922,1157,1183]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">Figs. 6</figureCitation>
|
||
G, H, 11K, L), and there are no significant differences, certainly not at the genus level. The ambulatory legs of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCAE69CD8FBD9FB11" box="[958,1048,1196,1221]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCAE69CD8FBD9FB11" box="[958,1048,1196,1221]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D04D32D0C1FCD0B9CD8FB3CFB11" box="[1107,1277,1196,1221]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCD0B9CD8FB3CFB11" box="[1107,1277,1196,1221]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
are also very similar, sharing the armature along the ventral margins of the P3 and P4 meri (spines relatively weaker in
|
||
<taxonomicName id="4C5C4D04D32D0C1FC9CF9C8DFF30FAC6" box="[151,241,1273,1298]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC9CF9C8DFF30FAC6" box="[151,241,1273,1298]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
) with prominent tufts of setae on the propodus (
|
||
<figureCitation id="13672A02D32D0C1FCA769C8DFC40FAC7" box="[814,897,1273,1299]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13672A02D32D0C1FCAC89C8DFC5EFAC7" box="[912,927,1273,1299]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones & Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="13" pageNumber="14">7</figureCitation>
|
||
A, 8). The carapaces of the two species are also similar, the only major differences in their relative proportions, the carapace of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCDC49D54FB39FAED" box="[1180,1272,1312,1337]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCDC49D54FB39FAED" box="[1180,1272,1312,1337]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
being almost always transversely rectangular and relatively broader (average width to length ratio 1.43, versus
|
||
<quantity id="4CA49B62D32D0C1FCC1F9D32FA5DFAB4" box="[1351,1436,1350,1376]" metricMagnitude="-2" metricUnit="m" metricValue="3.3528" pageId="13" pageNumber="14" unit="in" value="1.32">1.32 in</quantity>
|
||
<taxonomicName id="4C5C4D04D32D0C1FC9CF9D19FE85FA52" box="[151,324,1389,1414]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC9CF9D19FE85FA52" box="[151,324,1389,1414]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
). The carapace regions of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCBD19D19FD25FA52" box="[649,740,1389,1414]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCBD19D19FD25FA52" box="[649,740,1389,1414]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
are also relatively more developed and pronounced, with scattered groups of granules (
|
||
<figureCitation id="13672A02D32D0C1FC8AA9DE0FDF8FA7A" box="[498,569,1428,1454]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCBE19DE0FCA3FA79" box="[697,866,1428,1453]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCBE19DE0FCA3FA79" box="[697,866,1428,1453]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
, the regions are lower and less distinct, with the granules much smaller (
|
||
<figureCitation id="13672A02D32D0C1FC8F79DCFFE3EFA01" box="[431,511,1467,1493]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones & Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="13" pageNumber="14">Figs. 7</figureCitation>
|
||
A, 8). The anterolateral margins also differ; those of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCD3D9DCFFB7EFA00" box="[1125,1215,1467,1492]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCD3D9DCFFB7EFA00" box="[1125,1215,1467,1492]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
always have a low but visible lobe behind the external orbital tooth (
|
||
<figureCitation id="13672A02D32D0C1FCB809D95FCE1FA2F" box="[728,800,1505,1531]" captionStart="FIGURE 5" captionStartId="11.[151,256,1974,1998]" captionTargetBox="[151,1436,185,1952]" captionTargetId="figure@11.[151,1436,185,1954]" captionTargetPageId="11" captionText="FIGURE 5. Nasima dotilliformis (Alcock, 1900). A, male (11.7 x 16.5 mm) (NHM 1974.389), Persian Gulf; B, male (10.7 x 15.1 mm) (ZRC), Iraq; C, female (8.7 x 13.0 mm) (NHM 1984.410), Pakistan." httpUri="https://zenodo.org/record/191914/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13672A02D32D0C1FCA749D95FCFAFA2F" box="[812,827,1505,1531]" captionStart="FIGURE 6" captionStartId="12.[151,257,1713,1737]" captionTargetBox="[172,1417,403,1640]" captionTargetId="figure@12.[160,1429,382,1688]" captionTargetPageId="12" captionText="FIGURE 6. Nasima dotilliformis (Alcock, 1900), male (7.5 x 10.6 mm) (NHM 1984.497), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; C, right third maxilliped; D, outer view of right chela; E, lateral view of right chela; F, male abdomen; G, left G 1; H, distal part of left G 1. Scales: A – E, G = 1.0 mm; F, H = 0.5 mm." httpUri="https://zenodo.org/record/191915/files/figure.png" pageId="13" pageNumber="14">6</figureCitation>
|
||
A), while in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA959D95FBB7FA2E" box="[973,1142,1505,1530]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCA959D95FBB7FA2E" box="[973,1142,1505,1530]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
, the anterolateral margin is entire and never develops a tooth or lobe, although the area just behind the external orbital tooth may be crenulated or uneven. This character may not be reliable once a larger series of specimens can be examined. The likelihood that both genera are synonymous is high, but the lack of sexual dimorphism in the chelipeds of
|
||
<taxonomicName id="4C5C4D04D32D0C1FC9CF9E08FF30F941" box="[151,241,1660,1685]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC9CF9E08FF30F941" box="[151,241,1660,1685]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
(distinct in
|
||
<taxonomicName id="4C5C4D04D32D0C1FC8D89E08FDEBF941" box="[384,554,1660,1685]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FC8D89E08FDEBF941" box="[384,554,1660,1685]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
), the presently observed differences in carapace shape and ornamentation, and the presence of a low anterolateral lobe in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB8D9ED7FCF0F968" box="[725,817,1699,1724]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCB8D9ED7FCF0F968" box="[725,817,1699,1724]" italics="true" pageId="13" pageNumber="14">Nasima</emphasis>
|
||
</taxonomicName>
|
||
(absent in
|
||
<taxonomicName id="4C5C4D04D32D0C1FCAE29ED7FBA8F968" box="[954,1129,1699,1724]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928EA95D32D0C1FCAE29ED7FBA8F968" box="[954,1129,1699,1724]" italics="true" pageId="13" pageNumber="14">Leptochryseus</emphasis>
|
||
</taxonomicName>
|
||
) suggest that it is best to separate them for the moment. In addition, it is clear that
|
||
<taxonomicName id="4C5C4D04D32D0C1FCA109EBDFBF3F936" box="[840,1074,1737,1762]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dotillifomis">
|
||
<emphasis id="B928EA95D32D0C1FCA109EBDFBF3F936" box="[840,1074,1737,1762]" italics="true" pageId="13" pageNumber="14">Nasima dotillifomis</emphasis>
|
||
</taxonomicName>
|
||
is a smaller-size species, with females maturing and bearing eggs less than
|
||
<quantity id="4CA49B62D32D0C1FCBF19E84FD3CF8DE" box="[681,765,1776,1802]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="mm" value="10.0">10 mm</quantity>
|
||
in carapace width. In contrast, females of
|
||
<taxonomicName id="4C5C4D04D32D0C1FCDAC9E84FA5DF8DD" box="[1268,1436,1776,1801]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kuwaitensis">
|
||
<emphasis id="B928EA95D32D0C1FCDAC9E84FA5DF8DD" box="[1268,1436,1776,1801]" italics="true" pageId="13" pageNumber="14">L. kuwaitensis</emphasis>
|
||
</taxonomicName>
|
||
<quantity id="4CA49B62D32D0C1FC9CF9F62FF28F8E4" box="[151,233,1814,1840]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="mm" value="25.0">25 mm</quantity>
|
||
in carapace width do not appear to be fully mature. The few differences observed here do not appear to be very significant and key to resolving the question of whether these two genera are distinct may be in obtaining a much larger series for
|
||
<taxonomicName id="4C5C4D04D32D0C1FCB709F10FD11F8A9" box="[552,720,1892,1917]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kuwaitensis">
|
||
<emphasis id="B928EA95D32D0C1FCB709F10FD11F8A9" box="[552,720,1892,1917]" italics="true" pageId="13" pageNumber="14">L. kuwaitensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |