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<mods:title id="6ACD7CCBCB152B556EF9166791D51B12">Four new species of Macrostomus Wiedemann and a checklist of Empididae s. str (Diptera) from Peru</mods:title>
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<mods:namePart id="150FD8310EF72F353407ED8C5FDD88CB">Rafael, José Albertino</mods:namePart>
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<mods:affiliation id="0C839EAAAA65F7CC318B80D12DD8FBCF">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Manaus, Amazonas, Brazil. jarafael @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<mods:namePart id="6B5CDE8AA0309E51F5E26427E2D165FF">Marques, Dayse Willkenia A.</mods:namePart>
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<mods:affiliation id="8D96D1C6FBF9FF5635EC2D13363335BC">Instituto Nacional de Pesquisas da Amazônia, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas, Brazil. willkenia @ gmail. com; https: // orcid. org / 0000 - 0001 - 7260 - 5760</mods:affiliation>
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<heading id="D0AB81F6FFCEFFF275D1AF8EC4AFA9C7" bold="true" box="[151,706,1410,1437]" fontSize="11" level="1" pageId="13" pageNumber="519" reason="1">
<taxonomicName id="4C5C4D19FFCEFFF275D1AF8EC4AFA9C7" ID-CoL="6R472" authority="Rafael &amp; Cumming" authorityName="Rafael &amp; Cumming" authorityYear="2006" box="[151,706,1410,1437]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="ciliaticosta">
<emphasis id="B928EA88FFCEFFF275D1AF8EC4AFA9C7" bold="true" box="[151,706,1410,1437]" pageId="13" pageNumber="519">
<emphasis id="B928EA88FFCEFFF275D1AF8EC7A8A9C6" bold="true" box="[151,453,1410,1437]" italics="true" pageId="13" pageNumber="519">Macrostomus ciliaticosta</emphasis>
Rafael &amp; Cumming
</emphasis>
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<paragraph id="8BE3369AFFCEFFF275D1AFABC759A99B" blockId="13.[151,706,1410,1472]" box="[151,308,1446,1472]" pageId="13" pageNumber="519">
(
<figureCitation id="13672A1FFFCEFFF275D9AFABC741A99B" box="[159,300,1446,1472]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Figs 10AG</figureCitation>
)
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<paragraph id="8BE3369AFFCEFFF275D1AFE6C209AA58" blockId="13.[151,1124,1514,1570]" box="[151,1124,1514,1539]" pageId="13" pageNumber="519">
<treatmentCitationGroup id="AB4C11B4FFCEFFF275D1AFE6C209AA58" box="[151,1124,1514,1539]" pageId="13" pageNumber="519">
<treatmentCitation id="0AFD108BFFCEFFF275D1AFE6C4A3AA59" ID-DOI="http://doi.org/10.5281/zenodo.6493798" ID-Zenodo-Dep="6493798" author="Rafael, J. A. &amp; Cumming, J. M." box="[151,718,1514,1539]" httpUri="http://treatment.plazi.org/id/163B5B4D3464FFE5FEF7CBE5FD67453E" page="52" pageId="13" pageNumber="519" year="2006">
<taxonomicName id="4C5C4D19FFCEFFF275D1AFE6C4A3AA59" ID-CoL="6R472" authority="Rafael &amp; Cumming, 2006: 52" authorityName="Rafael &amp; Cumming" authorityPageNumber="52" authorityYear="2006" box="[151,718,1514,1539]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="ciliaticosta">
<emphasis id="B928EA88FFCEFFF275D1AFE6C7FAAA58" box="[151,407,1515,1539]" italics="true" pageId="13" pageNumber="519">Macrostomus ciliaticosta</emphasis>
<materialsCitation id="3B343CC7FFCEFFF274DBAFE6C4A3AA59" ID-GBIF-Occurrence="3312253323" box="[413,718,1514,1539]" country="Peru" location="Quincemil" pageId="13" pageNumber="519" specimenCount="1" stateProvince="Cusco">
<bibRefCitation id="EFCD4B6BFFCEFFF274DBAFE6C4A3AA59" author="Rafael, J. A. &amp; Cumming, J. M." box="[413,718,1514,1539]" pageId="13" pageNumber="519" pagination="49 - 64" refId="ref17230" refString="Rafael, J. A. &amp; Cumming, J. M. (2006) New species of Macrostomus Wiedemann (Diptera, Empididae) from Colombia, Ecuador and Peru. Zootaxa, 1136 (1), 49 - 64. https: // doi. org / 10.11646 / zootaxa. 1136.1.3" type="journal article" year="2006">Rafael &amp; Cumming, 2006: 52</bibRefCitation>
</materialsCitation>
</taxonomicName>
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.
<typeStatus id="54E78838FFCEFFF27791AFE6C564AA58" box="[727,777,1515,1539]" pageId="13" pageNumber="519">Type</typeStatus>
locality:
<collectingCountry id="F34B760AFFCEFFF2762FAFE6C5F7AA59" box="[873,922,1515,1538]" name="Peru" pageId="13" pageNumber="519">Peru</collectingCountry>
,
<collectingRegion id="4998F878FFCEFFF276E3AFE6C585AA58" box="[933,1000,1515,1539]" country="Peru" name="Cusco" pageId="13" pageNumber="519">Cuzco</collectingRegion>
,
<location id="8E836041FFCEFFF276B4AFE6C20DAA58" LSID="urn:lsid:plazi:treatment:03F5878CFFCEFFF175D1AF8EC423AD47:8E836041FFCEFFF276B4AFE6C20DAA58" box="[1010,1120,1515,1539]" country="Peru" name="Quincemil" pageId="13" pageNumber="519" stateProvince="Cusco">Quincemil</location>
.
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</paragraph>
<paragraph id="8BE3369AFFCEFFF275D1AC07C435AA7A" blockId="13.[151,1124,1514,1570]" box="[151,600,1545,1570]" pageId="13" pageNumber="519">
<treatmentCitationGroup id="AB4C11B4FFCEFFF275D1AC07C435AA7A" box="[151,600,1545,1570]" pageId="13" pageNumber="519">
<treatmentCitation id="0AFD108BFFCEFFF275D1AC07C47BAA7A" author="Yang, D. &amp; Zhang, K. &amp; Yao, G. &amp; Zhang, J." box="[151,534,1545,1570]" page="145" pageId="13" pageNumber="519" year="2007">
<taxonomicName id="4C5C4D19FFCEFFF275D1AC07C47BAA7A" ID-CoL="6R472" authority=": Yang et al., 2007: 145" authorityName="Yang" authorityPageNumber="145" authorityYear="2007" box="[151,534,1545,1570]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="ciliaticosta">
<emphasis id="B928EA88FFCEFFF275D1AC07C747AA7A" box="[151,298,1545,1569]" italics="true" pageId="13" pageNumber="519">M. ciliaticosta</emphasis>
:
<bibRefCitation id="EFCD4B6BFFCEFFF27470AC07C47BAA7A" author="Yang, D. &amp; Zhang, K. &amp; Yao, G. &amp; Zhang, J." box="[310,534,1545,1570]" pageId="13" pageNumber="519" refId="ref17703" refString="Yang, D., Zhang, K., Yao, G. &amp; Zhang, J. (2007) World Catalog of Empididae (Insecta: Diptera). China Agricultural University Press, Beijing, 599 pp." type="book" year="2007">
Yang
<emphasis id="B928EA88FFCEFFF27437AC07C7F0AA7A" box="[369,413,1545,1569]" italics="true" pageId="13" pageNumber="519">et al</emphasis>
., 2007: 145
</bibRefCitation>
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(cat.).
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</paragraph>
</subSubSection>
<subSubSection id="C3466511FFCEFFF175D1AC42C4A2ACEA" lastPageId="14" lastPageNumber="520" pageId="13" pageNumber="519" type="diagnosis">
<paragraph id="8BE3369AFFCEFFF275D1AC42C33EAB3D" blockId="13.[151,1437,1615,2038]" pageId="13" pageNumber="519">
<emphasis id="B928EA88FFCEFFF275D1AC42C730AA31" bold="true" box="[151,349,1615,1642]" pageId="13" pageNumber="519">Diagnosis. Male.</emphasis>
Two pairs of ocellar setae; thorax yellow with scutum and scutellum brown to black (
<figureCitation id="13672A1FFFCEFFF27060AC42C3E2AA31" box="[1318,1423,1615,1642]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10A</figureCitation>
); hind tibia (
<figureCitation id="13672A1FFFCEFFF27451AC7EC7ECAAD5" box="[279,385,1651,1678]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10A</figureCitation>
) with 56 dorsal setae; wing brown infuscated along costal cell and apex of cell r
<subScript id="17D834DFFFCEFFF27005AC8BC331AACF" attach="left" box="[1347,1372,1670,1684]" fontSize="6" pageId="13" pageNumber="519">2+3</subScript>
(
<figureCitation id="13672A1FFFCEFFF27029AC7EC6A6AAE9" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10A</figureCitation>
); cell dm somewhat acute; veins M
<subScript id="17D834DFFFCEFFF2771EACA7C40DAAE3" attach="left" box="[600,608,1706,1720]" fontSize="6" pageId="13" pageNumber="519">1</subScript>
, M
<subScript id="17D834DFFFCEFFF277C1ACA7C4E2AAE3" attach="left" box="[647,655,1706,1720]" fontSize="6" pageId="13" pageNumber="519">2</subScript>
and CuA+CuP somewhat distinct; tergite 8 (
<figureCitation id="13672A1FFFCEFFF271C0AC9AC37DAAE9" box="[1158,1296,1687,1714]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Figs 10B, C</figureCitation>
) with somewhat acute truncated distal projection; epandrium (
<figureCitation id="13672A1FFFCEFFF27789ACB6C55EAA8D" box="[719,819,1723,1750]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10B</figureCitation>
) with anterodorsal and posterodorsal lobes of subequal height; anterior cercus (
<figureCitation id="13672A1FFFCEFFF274E4ACD2C46BAAA1" box="[418,518,1759,1786]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10B</figureCitation>
) somewhat narrow; median cercus with slender setae; posterior cercus (
<figureCitation id="13672A1FFFCEFFF27076ACD2C3F9AAA1" box="[1328,1428,1759,1786]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10B</figureCitation>
) with apex acute and with a few long slender setae at margin, with wide ventral protuberance and many marginal yellow setae; hypandrium (
<figureCitation id="13672A1FFFCEFFF27481AD2AC440AB19" box="[455,557,1831,1858]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10D</figureCitation>
) with distinct bifid branch posteriorly, with many setae along posterior margin and distinct membranous area; in posterior view with apical lateral projections inwardly directed (
<figureCitation id="13672A1FFFCEFFF271A7AD46C32AAB3D" box="[1249,1351,1867,1894]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="8BE3369AFFCEFFF27581AD7DC3F8ABAE" blockId="13.[151,1437,1615,2038]" pageId="13" pageNumber="519">
<emphasis id="B928EA88FFCEFFF27581AD7DC748ABD1" bold="true" box="[199,293,1904,1930]" pageId="13" pageNumber="519">Female.</emphasis>
Hind tibia with 4 long dorsal setae and short dorsal pinnate setae along distal half; hind tarsomere 1 with 1 slender dorsal seta; sternite 8 (
<figureCitation id="13672A1FFFCEFFF27702AD9EC4CAABF5" box="[580,679,1939,1966]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Fig. 10F</figureCitation>
) with a subapical paired sclerotized protuberance, medially with a longitudinal somewhat membranous line, laterally with median third less sclerotized; genital fork (
<figureCitation id="13672A1FFFCEFFF271ABADBAC31EAB89" box="[1261,1395,1975,2002]" captionStart="FIGURE 10" captionStartId="14.[151,250,1227,1251]" captionTargetBox="[159,1437,312,1202]" captionTargetId="figure-52@14.[150,1437,308,1202]" captionTargetPageId="14" captionText="FIGURE 10. Macrostomus ciliaticosta Rafael &amp; Cumming, 2006. ♂ (AE). A. Habitus, lateral view; B. Apex of abdomen, lateral view; C. Tergite 8, dorsal view; D. Hypandrium and phallus, lateral view (black arrow indicating the membranous area); E. Phallus, posterior view. ♀ (FG). F. Apex of abdomen, lateral view; G. Apex of abdomen, dorsal pieces in dorsal view, ventral pieces in ventral view. Scale bars, figures A = 1 mm; B, F, G = 0.3 mm; CE = 0.2 mm.Abbreviations: a ce, anterior cercus; epand, epandrium; gen fk, genital fork; gen ch, genital chamber; m ce, median cercus; p ce, posterior cercus; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921377" httpUri="https://zenodo.org/record/4921377/files/figure.png" pageId="13" pageNumber="519">Figs 10F, G</figureCitation>
) Yshaped with rather strong lateral arms and a median short protuberance ventrally, with genital chamber membranous.
</paragraph>
<paragraph id="8BE3369AFFCDFFF17581AA9BC4A2ACEA" blockId="14.[151,1436,150,285]" box="[199,719,150,177]" pageId="14" pageNumber="520">
<emphasis id="B928EA88FFCDFFF17581AA9BC7BCACEB" bold="true" box="[199,465,150,176]" pageId="14" pageNumber="520">Geographical records.</emphasis>
<collectingCountry id="F34B760AFFCDFFF1749EAA9BC472ACEB" box="[472,543,150,176]" name="Brazil" pageId="14" pageNumber="520">Brazil</collectingCountry>
,
<collectingCountry id="F34B760AFFCDFFF1776CAA9BC4E4ACEB" box="[554,649,150,176]" name="Ecuador" pageId="14" pageNumber="520">Ecuador</collectingCountry>
,
<collectingCountry id="F34B760AFFCDFFF177D2AA9AC4A6ACEA" box="[660,715,151,177]" name="Peru" pageId="14" pageNumber="520">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3466511FFCDFFF17581AAB7C53AAC8E" box="[199,855,186,213]" pageId="14" pageNumber="520" type="materials_examined">
<paragraph id="8BE3369AFFCDFFF17581AAB7C53AAC8E" blockId="14.[151,1436,150,285]" box="[199,855,186,213]" pageId="14" pageNumber="520">
<emphasis id="B928EA88FFCDFFF17581AAB7C7DCAC8F" bold="true" box="[199,433,186,212]" pageId="14" pageNumber="520">Material examined.</emphasis>
Listed in
<bibRefCitation id="EFCD4B6BFFCDFFF17767AAB7C53EAC8E" author="Rafael, J. A. &amp; Cumming, J. M." box="[545,851,186,213]" pageId="14" pageNumber="520" pagination="49 - 64" refId="ref17230" refString="Rafael, J. A. &amp; Cumming, J. M. (2006) New species of Macrostomus Wiedemann (Diptera, Empididae) from Colombia, Ecuador and Peru. Zootaxa, 1136 (1), 49 - 64. https: // doi. org / 10.11646 / zootaxa. 1136.1.3" type="journal article" year="2006">Rafael &amp; Cumming (2006)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3466511FFCDFFF17581AAD3C423AD47" pageId="14" pageNumber="520" type="discussion">
<paragraph id="8BE3369AFFCDFFF17581AAD3C423AD47" blockId="14.[151,1436,150,285]" pageId="14" pageNumber="520">
<emphasis id="B928EA88FFCDFFF17581AAD3C756ACA3" bold="true" box="[199,315,222,248]" pageId="14" pageNumber="520">Remarks.</emphasis>
This species was described based on specimens from
<collectingCountry id="F34B760AFFCDFFF176EBAAD3C599ACA3" box="[941,1012,222,248]" name="Brazil" pageId="14" pageNumber="520">Brazil</collectingCountry>
,
<collectingCountry id="F34B760AFFCDFFF17147AAD3C233ACA3" box="[1025,1118,222,248]" name="Ecuador" pageId="14" pageNumber="520">Ecuador</collectingCountry>
and
<collectingCountry id="F34B760AFFCDFFF171DFAAD2C2A1ACA2" box="[1177,1228,223,249]" name="Peru" pageId="14" pageNumber="520">Peru</collectingCountry>
and no additional specimens of this species were studied.
</paragraph>
</subSubSection>
</treatment>
</document>