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<document id="30A7EC740A76057D697D0E2785CF184E" ID-CLB-Dataset="300697" ID-DOI="10.11646/phytotaxa.638.3.5" ID-GBIF-Dataset="b0cf002f-542a-4b60-a5f6-99f322aa2c40" ID-ISSN="1179-3163" ID-Zenodo-Dep="13213030" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722602205828" checkinUser="felipe" docAuthor="Wojtas, K. Philip, Bandara, Champika &amp; Kumar, Pankaj" docDate="2024" docId="03D387F1AF760731D1EBFB25347DFEB5" docLanguage="en" docName="phytotaxa.638.3.5.pdf" docOrigin="Phytotaxa 638 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.638.3.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Holcoglossum clausum Wojtas, C. Bandara &amp; Kumar, sp. nov." docType="treatment" docVersion="2" lastPageNumber="263" masterDocId="FFEAFF89AF770736D163FFA83117FFD9" masterDocTitle="A new species of Holcoglossum (Orchidaceae, Aeridinae) from Southern Shan State, Myanmar with taxonomic notes on Holcoglossum himalaicum" masterLastPageNumber="267" masterPageNumber="257" pageNumber="258" updateTime="1722802138542" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="33E090D93CC34D247F3C4E8E9F1EAB60">A new species of Holcoglossum (Orchidaceae, Aeridinae) from Southern Shan State, Myanmar with taxonomic notes on Holcoglossum himalaicum</mods:title>
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<mods:namePart id="CED4070F6FB1695B60C3884C0EDA0466">Bandara, Champika</mods:namePart>
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<mods:affiliation id="6BA359C91F265FCB2D3645F1DA8C1A07">Faculty of Graduate Studies, University of Sri Jayewardenepura, Gangodawila, Nugegoda 10250, Sri Lanka.</mods:affiliation>
<mods:nameIdentifier id="AA4F3C7F82C53C3CF61D43E651C0F7DF" type="email">cshbandara@gmail.com</mods:nameIdentifier>
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<mods:namePart id="136F08D4841F87ED727E940F62047323">Kumar, Pankaj</mods:namePart>
<mods:nameIdentifier id="F8FDE50B48452E93D3B69549F89424EC" type="ORCID">0000-0002-3469-5731</mods:nameIdentifier>
<mods:affiliation id="21F415903461F1B2F4449F6C14C804B2">Department of Plant and Soil Science, Texas Tech University, Lubbock, Texas 79409, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="0C4E682C5954195C556EE579F0739EB3" type="email">sahanipankaj@gmail.com</mods:nameIdentifier>
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<taxonomicName id="4C7A4D64AF760737D1EBFB2533EAFB71" authority="Wojtas, C.Bandara &amp; Kumar" authorityName="Wojtas, C. Bandara &amp; Kumar" box="[136,765,1165,1192]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="clausum" status="sp. nov.">
<emphasis id="B90EEAF5AF760737D1EBFB253081FB71" bold="true" box="[136,406,1165,1192]" italics="true" pageId="1" pageNumber="258">Holcoglossum clausum</emphasis>
Wojtas, C.Bandara &amp; Kumar
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,
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<taxonomicNameLabel id="A23D578EAF760737D269FB263277FB71" box="[778,864,1166,1192]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
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<collectionCode id="ED6BAE22AF760737D1EBFB1D31C2FB12" box="[136,213,1205,1227]" pageId="1" pageNumber="258">TYPES</collectionCode>
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:
<collectingCountry id="F36D7677AF760737D186FB1D3071FB13" box="[229,358,1205,1226]" name="Myanmar" pageId="1" pageNumber="258">MYANMAR</collectingCountry>
.
<collectingRegion id="49BEF805AF760737D011FB1D30C9FB13" box="[370,478,1205,1227]" country="Myanmar" name="Shan" pageId="1" pageNumber="258">Shan State</collectingRegion>
,
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:
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<location id="8EA5603CAF760737D3C5FB1D321FFB12" LSID="urn:lsid:plazi:treatment:03D387F1AF760731D1EBFB25347DFEB5:8EA5603CAF760737D3C5FB1D321FFB12" box="[678,776,1205,1227]" country="Myanmar" municipality="Taunggyi District" name="Saw Lwin" pageId="1" pageNumber="258" stateProvince="Shan">Saw Lwin</location>
s.n.
</emphasis>
(
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<specimenCode id="DBDC9E9CAF760737D227FB1D32DAFB13" box="[836,973,1205,1226]" collectionCode="SING" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="1" pageNumber="258" type="Herbarium">SING0237310</specimenCode>
</emphasis>
) (
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!-
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)
</materialsCitation>
;
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<emphasis id="B90EEAF5AF760737D5C3FB1D35F6FB13" box="[1184,1249,1205,1226]" italics="true" pageId="1" pageNumber="258">ibidem</emphasis>
,
<collectingMunicipality id="6BA1AC9DAF760737D592FB1D34BFFB12" box="[1265,1448,1205,1227]" pageId="1" pageNumber="258">Pindaya Township</collectingMunicipality>
,
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,
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<collectingDate id="EF80E9CFAF760737D031FB713305FB37" box="[338,530,1241,1263]" pageId="1" pageNumber="258" value="2008-11-30">30 November 2008</collectingDate>
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,
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et al. 080131
</emphasis>
(
<collectionCode id="ED6BAE22AF760737D39CFB71322EFB37" box="[767,825,1241,1262]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15870" lsid="urn:lsid:biocol.org:col:15870" name="Makino Botanical Garden" pageId="1" pageNumber="258" type="Herbarium">MBK</collectionCode>
,
<collectionCode id="ED6BAE22AF760737D227FB713248FB37" box="[836,863,1241,1262]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="258">TI</collectionCode>
! (
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<specimenCode id="DBDC9E9CAF760737D217FB7132FEFB37" box="[884,1001,1241,1262]" collectionCode="TI" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="258">TI00070510</specimenCode>
</emphasis>
),
<collectorName id="268F5331AF760737D29CFB713588FB36" box="[1023,1183,1241,1263]" pageId="1" pageNumber="258">Paratypes-image</collectorName>
seen)
</materialsCitation>
;
<materialsCitation id="3B123CBAAF760737D58AFB713062FAEE" collectionCode="HEID" country="Myanmar" location="Germany" municipality="Thailand" pageId="1" pageNumber="258" specimenCode="HEID204659" specimenCount="1" typeStatus="paratype">
plant obtained from
<collectingMunicipality id="6BA1AC9DAF760737D1DEFB553005FACA" box="[189,274,1277,1299]" pageId="1" pageNumber="258">Thailand</collectingMunicipality>
and introduced as
<emphasis id="B90EEAF5AF760737D0ADFB563303FACA" box="[462,532,1278,1299]" italics="true" pageId="1" pageNumber="258">in vitro</emphasis>
culture to
<location id="8EA5603CAF760737D3E2FB5533CCFACB" LSID="urn:lsid:plazi:treatment:03D387F1AF760731D1EBFB25347DFEB5:8EA5603CAF760737D3E2FB5533CCFACB" box="[641,731,1277,1298]" country="Myanmar" municipality="Thailand" name="Germany" pageId="1" pageNumber="258">Germany</location>
, flowered in cultivation,
<emphasis id="B90EEAF5AF760737D2B2FB553577FACB" box="[977,1120,1277,1298]" italics="true" pageId="1" pageNumber="258">KPW230426-1</emphasis>
(
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<specimenCode id="DBDC9E9CAF760737D50CFB5535E6FACB" box="[1135,1265,1277,1298]" collectionCode="HEID" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15548" lsid="urn:lsid:biocol.org:col:15548" name="University of Heidelberg" pageId="1" pageNumber="258" type="Herbarium">HEID204659</specimenCode>
</emphasis>
) (
<collectionCode id="ED6BAE22AF760737D464FB553453FACB" box="[1287,1348,1277,1298]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15548" lsid="urn:lsid:biocol.org:col:15548" name="University of Heidelberg" pageId="1" pageNumber="258" type="Herbarium">HEID</collectionCode>
!-
<typeStatus id="54C18845AF760737D42EFB5534BFFACB" box="[1357,1448,1277,1298]" pageId="1" pageNumber="258" type="paratype">Paratype</typeStatus>
, preserved in spirit)
</materialsCitation>
;
<materialsCitation id="3B123CBAAF760737D01EFA8932EAFA83" collectionCode="C" country="Thailand" elevation="1900" location="Doi Inthanond" pageId="1" pageNumber="258" specimenCount="1" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingCountry id="F36D7677AF760737D01EFA8930EBFAEF" box="[381,508,1313,1334]" name="Thailand" pageId="1" pageNumber="258">THAILAND</collectingCountry>
.
<collectingRegion id="49BEF805AF760737D367FA8933C4FAEE" box="[516,723,1313,1335]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="258">Chiang Mai Province</collectingRegion>
,
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,
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,
<emphasis id="B90EEAF5AF760737D2A5FA89350EFAEF" box="[966,1049,1313,1334]" italics="true" pageId="1" pageNumber="258">GT 8289</emphasis>
(spirit number 9604) (
<collectionCode id="ED6BAE22AF760737D593FA893417FAEF" box="[1264,1280,1313,1334]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="258" type="Herbarium">C</collectionCode>
!-
<typeStatus id="54C18845AF760737D469FA893470FAEF" box="[1290,1383,1313,1334]" pageId="1" pageNumber="258" type="paratype">Paratype</typeStatus>
, image seen, location according to
<bibRefCitation id="EFEB4B16AF760737D0A6FAED33A3FA83" author="Seidenfaden, G." box="[453,692,1349,1371]" pageId="1" pageNumber="258" pagination="1 - 356" refId="ref7293" refString="Seidenfaden, G. (1988) Orchid Genera in Thailand 14: Fifty-nine Vandoid genera. Opera Botanica 95: 1 - 356." type="journal article" year="1988">Seidenfaden (1988: 317)</bibRefCitation>
, see note below). (
<figureCitation id="13412A62AF760737D204FAED32D4FA83" box="[871,963,1349,1371]" captionStart="FIGURE 1" captionStartId="2.[136,229,1820,1842]" captionTargetBox="[151,1436,187,1793]" captionTargetId="figure-27@2.[151,1436,187,1793]" captionTargetPageId="2" captionText="FIGURE 1. Holcoglossum clausum Wojtas, C.Bandara &amp; Kumar. A. A plant. B. Front view of flower. C. Side view of flower. D. Floral bract. E. Dorsal sepal. F. Petal. G. Lateral sepal. H. Labellum. I. Side view of flower (dorsal and lateral sepal and petal removed). J. Side lobe of labellum. K. Front view of column. L. Pollinarium. M. Dorsal and ventral view of anther cap. [Compilation by C. Bandara, photographs by K.P. Wojtas, based on fresh specimen and paratype: KPW230426-1 (HEID!)]." figureDoi="http://doi.org/10.5281/zenodo.13213032" httpUri="https://zenodo.org/record/13213032/files/figure.png" pageId="1" pageNumber="258">Figures 1</figureCitation>
,
<figureCitation id="13412A62AF760737D2ACFAED32CAFA83" box="[975,989,1349,1370]" captionStart="FIGURE 2" captionStartId="3.[136,229,1858,1880]" captionTargetBox="[151,1435,190,1834]" captionTargetId="figure-17@3.[151,1435,190,1834]" captionTargetPageId="3" captionText="FIGURE 2. Holcoglossum clausum Wojtas, C.Bandara &amp; Kumar. A. Close up of inflorescence. B. Side view of flower, C. Floral bract. D. Dorsal sepal. E. Petal. F. Lateral sepal. G. Labellum. H. Labellum with spur. I. Side view of labellum. J. Immature fruit. K. Anther cap. [Compilation and photographs by Champika Bandara, based on holotype: Saw Lwin s.n. - SING0237310 (SING!) with permission from SING Herbarium, Singapore Botanic Gardens]" figureDoi="http://doi.org/10.5281/zenodo.13213036" httpUri="https://zenodo.org/record/13213036/files/figure.png" pageId="1" pageNumber="258">2</figureCitation>
,
<figureCitation id="13412A62AF760737D284FAED32E3FA83" box="[999,1012,1349,1370]" captionStart="FIGURE 3" captionStartId="4.[136,229,1825,1847]" captionTargetBox="[154,1435,193,1798]" captionTargetId="figure-27@4.[151,1435,190,1801]" captionTargetPageId="4" captionText="FIGURE 3. Holcoglossum clausum Wojtas, C.Bandara &amp; Kumar. A. A plant. B. Close up of inflorescence. C. Front view of flower. D. Side view of flower. E. Floral bract. F. Dorsal sepal. G. Petal. H. Lateral sepal. I. Labellum. J. Side view of labellum. K. Close up of side lobe. L. Labellum with spur. M. Front view of column. N. Anther cap. O. Pollinarium. [Drawn by Champika Bandara from Saw Lwin s.n. - SING0237310 (SING!-Holotype) and KPW230426-1]." figureDoi="http://doi.org/10.5281/zenodo.13213038" httpUri="https://zenodo.org/record/13213038/files/figure.png" pageId="1" pageNumber="258">3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C360656CAF760737D1EBFAC130A9F93F" pageId="1" pageNumber="258" type="diagnosis">
<paragraph id="8BC536E7AF760737D1EBFAC130A9F93F" blockId="1.[136,1452,1165,1767]" pageId="1" pageNumber="258">
<emphasis id="B90EEAF5AF760737D1EBFAC131FCFAA6" bold="true" box="[136,235,1385,1407]" pageId="1" pageNumber="258">Diagnosis</emphasis>
:
<taxonomicName id="4C7A4D64AF760737D19BFAC130C0FAA7" box="[248,471,1385,1406]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF760737D19BFAC130C0FAA7" box="[248,471,1385,1406]" italics="true" pageId="1" pageNumber="258">Holcoglossum clausum</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="4C7A4D64AF760737D3B7FAC23274FAA7" authorityName="Averyanov" authorityYear="1988" baseAuthorityName="Deb, Sengupta &amp; Malick" box="[724,867,1385,1407]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF760737D3B7FAC23274FAA7" box="[724,867,1385,1407]" italics="true" pageId="1" pageNumber="258">H. himalaicum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A4D64AF760737D2F1FAC2355CFAA7" authorityName="R. Rice" authorityYear="2018" baseAuthorityName="Seidenfaden" baseAuthorityYear="1970" box="[914,1099,1385,1407]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF760737D2F1FAC2355CFAA7" box="[914,1099,1385,1407]" italics="true" pageId="1" pageNumber="258">H. semiteretifolium</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C7A4D64AF760737D5D8FAC2343FFAA7" box="[1211,1320,1385,1407]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF760737D5D8FAC2343FFAA7" box="[1211,1320,1385,1407]" italics="true" pageId="1" pageNumber="258">H. clausum</emphasis>
</taxonomicName>
has subterete leaves, comparatively the shortest stem (up to
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long), comparatively shorter inflorescence (up to
<quantity id="4C829B02AF760737D5F8FA2535DDFA7B" box="[1179,1226,1421,1442]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="cm" value="7.0">7 cm</quantity>
long) bearing up to 17 flowers, longest pedicel and ovary (
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long), largest sepals (≥
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long) and petals (≥
<quantity id="4C829B02AF760737D55FFA193564FA1F" box="[1084,1139,1457,1478]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="mm" value="7.0">7 mm</quantity>
long) of all three species, larger midlobe of the labellum (up to
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long,
<quantity id="4C829B02AF760737D313FA7D33AEFA33" box="[624,697,1493,1515]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="258" unit="mm" value="3.5">3.5 mm</quantity>
wide), shorter side lobe (≤
<quantity id="4C829B02AF760737D2B2FA7D351FFA33" box="[977,1032,1493,1514]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="258" unit="mm" value="2.0">2 mm</quantity>
long) of white color, spur of intermediate length (up to
<quantity id="4C829B02AF760737D023FA513096F9D7" box="[320,385,1529,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="258" unit="cm" value="1.3">1.3 cm</quantity>
long), smaller column (
<quantity id="4C829B02AF760737D308FA5133A1F9D7" box="[619,694,1529,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="258" unit="mm" value="1.5">1.5 mm</quantity>
×
<quantity id="4C829B02AF760737D3B2FA51320AF9D7" box="[721,797,1529,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="258" unit="mm" value="1.5">1.5 mm</quantity>
) and the plants flower during NovemberDecember.
<taxonomicName id="4C7A4D64AF760737D441FA51304FF9EB" authorityName="R. Rice" authorityYear="2018" baseAuthorityName="Seidenfaden" baseAuthorityYear="1970" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF760737D441FA51304FF9EB" italics="true" pageId="1" pageNumber="258">Holcoglossum semiteretifolium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A4D64AF760737D0E7F9B63306F9EB" authorityName="Averyanov" authorityYear="1988" baseAuthorityName="Deb, Sengupta &amp; Malick" box="[388,529,1565,1587]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF760737D0E7F9B63306F9EB" box="[388,529,1565,1587]" italics="true" pageId="1" pageNumber="258">H. himalaicum</emphasis>
</taxonomicName>
have subterete and semiterete leaves respectively, longer stem (≥
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long), longer inflorescence (up to
<quantity id="4C829B02AF760737D199F9E93023F98F" box="[250,308,1601,1623]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="258" unit="cm" value="15.0">15 cm</quantity>
long) bearing up to 15 flowers in former and up to 39 flowers in latter, shorter pedicel and ovary (≤
<quantity id="4C829B02AF760737D582F9E93435F98F" box="[1249,1314,1601,1623]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="258" unit="cm" value="1.5">1.5 cm</quantity>
long), smaller sepals (≤
<quantity id="4C829B02AF760737D07EF9CD3043F9A3" box="[285,340,1637,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="mm" value="7.0">7 mm</quantity>
long), smaller petals (≤
<quantity id="4C829B02AF760737D324F9CD3369F9A3" box="[583,638,1637,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="mm" value="7.0">7 mm</quantity>
long), shorter midlobe (up to
<quantity id="4C829B02AF760737D2C9F9CD32F6F9A3" box="[938,993,1637,1658]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="258" unit="mm" value="4.0">4 mm</quantity>
long, up to
<quantity id="4C829B02AF760737D53FF9CD35B2F9A3" box="[1116,1189,1637,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="258" unit="mm" value="2.5">2.5 mm</quantity>
wide), longer side lobe (≥
<quantity id="4C829B02AF760737D1DEF92131E3F947" box="[189,244,1673,1694]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="258" unit="mm" value="2.0">2 mm</quantity>
long) which is yellowish in former and pink in latter, spur shortest in former (
<quantity id="4C829B02AF760737D2B6F9213552F947" box="[981,1093,1673,1695]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="258" unit="cm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 cm</quantity>
), longest in latter (
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), larger column in both (2.0 mm × 2.0 mm) and both species have different flowering time (former in March and latter in OctoberJanuary) (see
<tableCitation id="C6F8035CAF760737D189F9793026F93F" box="[234,305,1745,1767]" captionStart="TABLE 1" captionStartId="6.[136,229,554,580]" captionText="TABLE 1. Morphological comparison between Holcoglossum clausum, H. himalaicum and H. semiteretifolium." pageId="1" pageNumber="258">Table 1</tableCitation>
and
<figureCitation id="13412A62AF760737D03CF97930A5F93F" box="[351,434,1745,1767]" captionStart="FIGURE 4" captionStartId="5.[136,229,1049,1071]" captionTargetBox="[151,1436,190,1025]" captionTargetId="figure-17@5.[151,1436,190,1025]" captionTargetPageId="5" captionText="FIGURE 4. Comparison of pink-floweredAsia-mainland Holcoglossum species.A. H.semiteritifolium. B. H. himalaicum. C. H.clausum. [Compilation by C. Bandara, picture of H. semiteretifolium by K. Keller, other pictures by K. P. Wojtas.]" figureDoi="http://doi.org/10.5281/zenodo.13213040" httpUri="https://zenodo.org/record/13213040/files/figure.png" pageId="1" pageNumber="258">Figure 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C360656CAF760730D1EBF8BD3211FAF0" lastPageId="6" lastPageNumber="263" pageId="1" pageNumber="258" type="description">
<paragraph id="8BC536E7AF760733D1EBF8BD321EF9F0" blockId="1.[136,1452,1813,2092]" lastBlockId="5.[136,1452,1154,2081]" lastPageId="5" lastPageNumber="262" pageId="1" pageNumber="258">
<emphasis id="B90EEAF5AF760737D1EBF8BD3005F8F6" bold="true" box="[136,274,1813,1839]" pageId="1" pageNumber="258">Description</emphasis>
: Monopodial, epiphytic herb, pendant, roots many, at the base of stem, silvery-grey, rigid and flexuous, creeping and densely appressed to substrate. Stem short up to
<quantity id="4C829B02AF760737D238F8923282F88D" box="[859,917,1850,1876]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="258" unit="cm" value="4.0">4 cm</quantity>
long and
<quantity id="4C829B02AF760737D56EF892354EF88D" box="[1037,1113,1850,1876]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="cm" value="0.7">0.7 cm</quantity>
broad; leaves 45, subterete, channelled on ventral side, up to
<quantity id="4C829B02AF760737D09AF8F53329F8A1" box="[505,574,1885,1912]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="258" unit="cm" value="45.0">45 cm</quantity>
long,
<quantity id="4C829B02AF760737D3E7F8F53216F8A1" box="[644,769,1885,1912]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="258" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 cm</quantity>
wide, acuminate, sheathing at the base, leaf sheath ca
<quantity id="4C829B02AF760737D403F8F634BCF8A1" box="[1376,1451,1886,1912]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="258" unit="cm" value="1.8">1.8 cm</quantity>
long, clasping the stem. Inflorescence bending to be horizontal, bracteate raceme, up to
<quantity id="4C829B02AF760737D536F82A359BF845" box="[1109,1164,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="258" unit="cm" value="7.0">7 cm</quantity>
long, rachis up to
<quantity id="4C829B02AF760737D43BF82A34B0F845" box="[1368,1447,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="258" unit="cm" value="4.8">4.8 cm</quantity>
, bearing 517 flowers. Pedicel with ovary clavate, ca.
<quantity id="4C829B02AF760737D3BCF80E3201F819" box="[735,790,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="258" unit="cm" value="3.0">3 cm</quantity>
long, 1.0
<quantity id="4C829B02AF760737D2EDF80E32F4F819" box="[910,995,1958,1984]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="258" unit="mm" value="1.7">1.7 mm</quantity>
in diameter, glabrous, ridged on surface. Floral bracts narrowly triangular, ca. 2.0 mm long, acute. Flowers not widely open, resupinate, ca.
<quantity id="4C829B02AF760737D585F8623454F83D" box="[1254,1347,1994,2020]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="258" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long, ca.
<quantity id="4C829B02AF760737D1EBF84531F2F7D1" box="[136,229,2029,2056]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="258" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
wide, pink or white coloured with darker markings on the dorsal surface along the mid-veins of petals and sepals. Dorsal sepal elliptic,
<quantity id="4C829B02AF760737D0AFF7B93344F7F5" box="[460,595,2065,2092]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.5" metricValueMin="7.2" pageId="1" pageNumber="258" unit="mm" value="7.35" valueMax="7.5" valueMin="7.2">7.27.5 mm</quantity>
long, up to
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wide, obtuse, 5-veined, pink; lateral sepals ellipticovate, slightly oblique, 7.89.0 mm long,
<quantity id="4C829B02AF720733D30EFB2A33E3FB44" box="[621,756,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="5" pageNumber="262" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.14.2 mm</quantity>
wide, acute, 3-veined, pink; petals elliptic, slightly oblique,
<quantity id="4C829B02AF720733D1EBFB0E3018FB18" box="[136,271,1190,1217]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.2" metricValueMin="7.1" pageId="5" pageNumber="262" unit="mm" value="7.15" valueMax="7.2" valueMin="7.1">7.17.2 mm</quantity>
long, 3.0
<quantity id="4C829B02AF720733D0EFFB0E30F6FB18" box="[396,481,1190,1217]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="262" unit="mm" value="3.1">3.1 mm</quantity>
wide, obtuse, 5-veined, pink; labellum 3-lobed, 5.56.0 mm long, up to
<quantity id="4C829B02AF720733D440FB0E3473FB18" box="[1315,1380,1190,1217]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="262" unit="mm" value="5.0">5 mm</quantity>
wide; midlobe tongue-shaped, 5.56.0 mm long,
<quantity id="4C829B02AF720733D313FB6233E0FB3C" box="[624,759,1226,1253]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="5" pageNumber="262" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.53.6 mm</quantity>
wide, succulent, retuse at the apex giving a bilobed look, 5- veined, pink, pointed forwards; side lobes erect, broadly ovate-orbicular,
<quantity id="4C829B02AF720733D2ACFB463541FAD0" box="[975,1110,1262,1289]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="5" pageNumber="262" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.61.7 mm</quantity>
long,
<quantity id="4C829B02AF720733D5C3FB463430FAD0" box="[1184,1319,1262,1289]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="5" pageNumber="262" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
wide, apex broadly obtuse, white; column very short and stout, 1.5 ×
<quantity id="4C829B02AF720733D273FABA327FFAF4" box="[784,872,1298,1325]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="262" unit="mm" value="1.5">1.5 mm</quantity>
, partly hidden by the side lobes of labellum; stigma orbicular to slightly cordate, 1.0 ×
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; rostellum with two short horns directed upwards; clinandrium round, almost flat, foot absent; spur located at the base of the column, slightly inflated on side,
<quantity id="4C829B02AF720733D502FAF235C9FAAC" box="[1121,1246,1370,1397]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="5" pageNumber="262" unit="cm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 cm</quantity>
long, 1.0
<quantity id="4C829B02AF720733D435FAF234BBFAAC" box="[1366,1452,1370,1397]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="262" unit="mm" value="1.2">1.2 mm</quantity>
wide, rounded at the apex, pink, slightly bent forwards; anther cap trapezoid, cucullate, 1.0 mm long,
<quantity id="4C829B02AF720733D473FAD63471FA40" box="[1296,1382,1406,1433]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="262" unit="mm" value="1.2">1.2 mm</quantity>
wide, dark pink, with two lobes inside; pollinarium 1 with two lobes,
<quantity id="4C829B02AF720733D23FFA0A32F4FA64" box="[860,995,1442,1469]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="262" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.21.4 mm</quantity>
long,
<quantity id="4C829B02AF720733D548FA0A3596FA64" box="[1067,1153,1442,1469]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="262" unit="mm" value="0.8">0.8 mm</quantity>
wide, each lobe fabiform,
<quantity id="4C829B02AF720733D1EBFA6E31C9FA38" box="[136,222,1478,1505]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="262" unit="mm" value="0.6">0.6 mm</quantity>
long,
<quantity id="4C829B02AF720733D041FA6E306FFA38" box="[290,376,1478,1505]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="262" unit="mm" value="0.4">0.4 mm</quantity>
wide, dark red, with a depression on the outer side; stipe elliptic with recurved margin, broader at the base, narrower above,
<quantity id="4C829B02AF720733D0CAFA4230E8F9DC" box="[425,511,1514,1541]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="262" unit="mm" value="0.8">0.8 mm</quantity>
long,
<quantity id="4C829B02AF720733D320FA42338EF9DC" box="[579,665,1514,1541]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="262" unit="mm" value="0.3">0.3 mm</quantity>
wide at base,
<quantity id="4C829B02AF720733D251FA42329FF9DC" box="[818,904,1514,1541]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="262" unit="mm" value="0.1">0.1 mm</quantity>
wide towards apex, white, translucent; viscidium trapezoid,
<quantity id="4C829B02AF720733D19DF9A63090F9F0" box="[254,391,1550,1577]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="5" pageNumber="262" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
, white, translucent. Fruit not seen.
</paragraph>
<caption id="DF05666FAF750734D1EBF8B4351BF844" ID-DOI="http://doi.org/10.5281/zenodo.13213032" ID-Zenodo-Dep="13213032" httpUri="https://zenodo.org/record/13213032/files/figure.png" pageId="2" pageNumber="259" startId="2.[136,229,1820,1842]" targetBox="[151,1436,187,1793]" targetPageId="2" targetType="figure">
<paragraph id="8BC536E7AF750734D1EBF8B4351BF844" blockId="2.[136,1452,1820,1950]" pageId="2" pageNumber="259">
<emphasis id="B90EEAF5AF750734D1EBF8B431E9F8E8" bold="true" box="[136,254,1820,1842]" pageId="2" pageNumber="259">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4D64AF750734D067F8B4322EF8EB" authority="Wojtas, C. Bandara &amp; Kumar. A. A" authorityName="Wojtas, C. Bandara &amp; Kumar. A. A" box="[260,825,1820,1842]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF750734D067F8B430F3F8E8" box="[260,484,1820,1841]" italics="true" pageId="2" pageNumber="259">Holcoglossum clausum</emphasis>
Wojtas, C.Bandara &amp; Kumar.
<emphasis id="B90EEAF5AF750734D26FF8B4320AF8E8" bold="true" box="[780,797,1820,1841]" pageId="2" pageNumber="259">A</emphasis>
. A
</taxonomicName>
plant.
<emphasis id="B90EEAF5AF750734D21FF8B4329BF8E8" bold="true" box="[892,908,1820,1841]" pageId="2" pageNumber="259">B</emphasis>
. Front view of flower.
<emphasis id="B90EEAF5AF750734D50FF8B4356AF8EB" bold="true" box="[1132,1149,1820,1842]" pageId="2" pageNumber="259">C</emphasis>
. Side view of flower.
<emphasis id="B90EEAF5AF750734D437F8B43472F8E8" bold="true" box="[1364,1381,1820,1841]" pageId="2" pageNumber="259">D</emphasis>
. Floral bract.
<emphasis id="B90EEAF5AF750734D1A5F8E831C1F88C" bold="true" box="[198,214,1856,1877]" pageId="2" pageNumber="259">E</emphasis>
. Dorsal sepal.
<emphasis id="B90EEAF5AF750734D008F8E8306DF88C" bold="true" box="[363,378,1856,1877]" pageId="2" pageNumber="259">F</emphasis>
. Petal.
<emphasis id="B90EEAF5AF750734D0A6F8E830CFF88F" bold="true" box="[453,472,1856,1878]" pageId="2" pageNumber="259">G</emphasis>
. Lateral sepal.
<emphasis id="B90EEAF5AF750734D313F8E83394F88C" bold="true" box="[624,643,1856,1877]" pageId="2" pageNumber="259">H</emphasis>
. Labellum.
<emphasis id="B90EEAF5AF750734D399F8E83214F88C" bold="true" box="[762,771,1856,1877]" pageId="2" pageNumber="259">I</emphasis>
. Side view of flower (dorsal and lateral sepal and petal removed).
<emphasis id="B90EEAF5AF750734D4FAF8E834B2F88C" bold="true" box="[1433,1445,1856,1877]" pageId="2" pageNumber="259">J</emphasis>
. Side lobe of labellum.
<emphasis id="B90EEAF5AF750734D000F8CC3061F8A0" bold="true" box="[355,374,1892,1913]" pageId="2" pageNumber="259">K</emphasis>
. Front view of column.
<emphasis id="B90EEAF5AF750734D303F8CC3367F8A0" bold="true" box="[608,624,1892,1913]" pageId="2" pageNumber="259">L</emphasis>
. Pollinarium.
<emphasis id="B90EEAF5AF750734D399F8CC3206F8A0" bold="true" box="[762,785,1892,1913]" pageId="2" pageNumber="259">M</emphasis>
. Dorsal and ventral view of anther cap. [Compilation by C. Bandara, photographs by K.P. Wojtas, based on fresh specimen and paratype:
<emphasis id="B90EEAF5AF750734D27BF82032B1F844" box="[792,934,1928,1949]" italics="true" pageId="2" pageNumber="259">KPW230426-1</emphasis>
(HEID!)].
</paragraph>
</caption>
<caption id="DF05666FAF740735D1EBF8EA335DF81C" ID-DOI="http://doi.org/10.5281/zenodo.13213036" ID-Zenodo-Dep="13213036" httpUri="https://zenodo.org/record/13213036/files/figure.png" pageId="3" pageNumber="260" startId="3.[136,229,1858,1880]" targetBox="[151,1435,190,1834]" targetPageId="3" targetType="figure">
<paragraph id="8BC536E7AF740735D1EBF8EA335DF81C" blockId="3.[136,1452,1858,1989]" pageId="3" pageNumber="260">
<emphasis id="B90EEAF5AF740735D1EBF8EA31EAF88E" bold="true" box="[136,253,1858,1880]" pageId="3" pageNumber="260">FIGURE 2.</emphasis>
<taxonomicName id="4C7A4D64AF740735D061F8EB3241F880" authority="Wojtas, C. Bandara &amp; Kumar. A. Close" box="[258,854,1858,1881]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="260" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF740735D061F8EB30F7F881" box="[258,480,1859,1880]" italics="true" pageId="3" pageNumber="260">Holcoglossum clausum</emphasis>
Wojtas, C.Bandara &amp; Kumar.
<emphasis id="B90EEAF5AF740735D261F8EA3204F88E" bold="true" box="[770,787,1858,1879]" pageId="3" pageNumber="260">A</emphasis>
. Close
</taxonomicName>
up of inflorescence.
<emphasis id="B90EEAF5AF740735D57FF8EB353BF881" bold="true" box="[1052,1068,1859,1880]" pageId="3" pageNumber="260">B</emphasis>
. Side view of flower,
<emphasis id="B90EEAF5AF740735D59DF8EA3418F881" bold="true" box="[1278,1295,1858,1880]" pageId="3" pageNumber="260">C</emphasis>
. Floral bract.
<emphasis id="B90EEAF5AF740735D4F7F8EB34B2F881" bold="true" box="[1428,1445,1859,1880]" pageId="3" pageNumber="260">D</emphasis>
. Dorsal sepal.
<emphasis id="B90EEAF5AF740735D06FF8CF300BF8A5" bold="true" box="[268,284,1895,1916]" pageId="3" pageNumber="260">E</emphasis>
. Petal.
<emphasis id="B90EEAF5AF740735D006F8CF3063F8A5" bold="true" box="[357,372,1895,1916]" pageId="3" pageNumber="260">F</emphasis>
. Lateral sepal.
<emphasis id="B90EEAF5AF740735D36BF8CE330CF8A5" bold="true" box="[520,539,1894,1916]" pageId="3" pageNumber="260">G</emphasis>
. Labellum.
<emphasis id="B90EEAF5AF740735D3ECF8CF33B5F8A5" bold="true" box="[655,674,1895,1916]" pageId="3" pageNumber="260">H</emphasis>
. Labellum with spur.
<emphasis id="B90EEAF5AF740735D215F8CF3268F8A5" bold="true" box="[886,895,1895,1916]" pageId="3" pageNumber="260">I</emphasis>
. Side view of labellum.
<emphasis id="B90EEAF5AF740735D50FF8CF356FF8A5" bold="true" box="[1132,1144,1895,1916]" pageId="3" pageNumber="260">J</emphasis>
. Immature fruit.
<emphasis id="B90EEAF5AF740735D47EF8CF3427F8A5" bold="true" box="[1309,1328,1895,1916]" pageId="3" pageNumber="260">K</emphasis>
. Anther cap. [Compilation and photographs by Champika Bandara, based on holotype:
<emphasis id="B90EEAF5AF740735D23BF823356AF879" box="[856,1149,1931,1952]" italics="true" pageId="3" pageNumber="260">Saw Lwin s.n. - SING0237310</emphasis>
(SING!) with permission from SING Herbarium, Singapore Botanic Gardens]
</paragraph>
</caption>
<caption id="DF05666FAF730732D1EBF88933B5F87B" ID-DOI="http://doi.org/10.5281/zenodo.13213038" ID-Zenodo-Dep="13213038" httpUri="https://zenodo.org/record/13213038/files/figure.png" pageId="4" pageNumber="261" startId="4.[136,229,1825,1847]" targetBox="[154,1435,193,1798]" targetPageId="4" targetType="figure">
<paragraph id="8BC536E7AF730732D1EBF88933B5F87B" blockId="4.[136,1452,1825,1955]" pageId="4" pageNumber="261">
<emphasis id="B90EEAF5AF730732D1EBF88931E9F8EF" bold="true" box="[136,254,1825,1847]" pageId="4" pageNumber="261">FIGURE 3.</emphasis>
<taxonomicName id="4C7A4D64AF730732D066F889322BF8EF" authority="Wojtas, C. Bandara &amp; Kumar. A. A" authorityName="Wojtas, C. Bandara &amp; Kumar. A. A" box="[261,828,1825,1847]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="261" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF730732D066F88930F2F8EF" box="[261,485,1825,1846]" italics="true" pageId="4" pageNumber="261">Holcoglossum clausum</emphasis>
Wojtas, C.Bandara &amp; Kumar.
<emphasis id="B90EEAF5AF730732D26CF8893237F8EF" bold="true" box="[783,800,1825,1846]" pageId="4" pageNumber="261">A</emphasis>
. A
</taxonomicName>
plant.
<emphasis id="B90EEAF5AF730732D2E3F8893287F8EF" bold="true" box="[896,912,1825,1846]" pageId="4" pageNumber="261">B</emphasis>
. Close up of inflorescence.
<emphasis id="B90EEAF5AF730732D5C3F88935A6F8EE" bold="true" box="[1184,1201,1825,1847]" pageId="4" pageNumber="261">C</emphasis>
. Front view of flower.
<emphasis id="B90EEAF5AF730732D4F7F88934B2F8EF" bold="true" box="[1428,1445,1825,1846]" pageId="4" pageNumber="261">D</emphasis>
. Side view of flower.
<emphasis id="B90EEAF5AF730732D02CF8ED3048F883" bold="true" box="[335,351,1861,1882]" pageId="4" pageNumber="261">E</emphasis>
. Floral bract.
<emphasis id="B90EEAF5AF730732D086F8ED30E3F883" bold="true" box="[485,500,1861,1882]" pageId="4" pageNumber="261">F</emphasis>
. Dorsal sepal.
<emphasis id="B90EEAF5AF730732D3E1F8ED3382F882" bold="true" box="[642,661,1861,1883]" pageId="4" pageNumber="261">G</emphasis>
. Petal.
<emphasis id="B90EEAF5AF730732D3BFF8ED33F8F883" bold="true" box="[732,751,1861,1882]" pageId="4" pageNumber="261">H</emphasis>
. Lateral sepal.
<emphasis id="B90EEAF5AF730732D2E2F8ED329DF883" bold="true" box="[897,906,1861,1882]" pageId="4" pageNumber="261">I</emphasis>
. Labellum.
<emphasis id="B90EEAF5AF730732D29EF8ED351EF883" bold="true" box="[1021,1033,1861,1882]" pageId="4" pageNumber="261">J</emphasis>
. Side view of labellum.
<emphasis id="B90EEAF5AF730732D590F8ED3411F883" bold="true" box="[1267,1286,1861,1882]" pageId="4" pageNumber="261">K</emphasis>
. Close up of side lobe.
<emphasis id="B90EEAF5AF730732D1DEF8C131DAF8A7" bold="true" box="[189,205,1897,1918]" pageId="4" pageNumber="261">L</emphasis>
. Labellum with spur.
<emphasis id="B90EEAF5AF730732D0FEF8C130A3F8A7" bold="true" box="[413,436,1897,1918]" pageId="4" pageNumber="261">M</emphasis>
. Front view of column.
<emphasis id="B90EEAF5AF730732D3FAF8C133BDF8A7" bold="true" box="[665,682,1897,1918]" pageId="4" pageNumber="261">N</emphasis>
. Anther cap.
<emphasis id="B90EEAF5AF730732D24AF8C1322BF8A6" bold="true" box="[809,828,1897,1919]" pageId="4" pageNumber="261">O</emphasis>
. Pollinarium. [Drawn by Champika Bandara from
<emphasis id="B90EEAF5AF730732D444F8C13008F87B" italics="true" pageId="4" pageNumber="261">Saw Lwin s.n. - SING0237310</emphasis>
(SING!-Holotype) and
<emphasis id="B90EEAF5AF730732D366F8253383F87B" box="[517,660,1933,1954]" italics="true" pageId="4" pageNumber="261">KPW230426-1</emphasis>
].
</paragraph>
</caption>
<caption id="DF05666FAF720733D1EBFBB13565FB8D" ID-DOI="http://doi.org/10.5281/zenodo.13213040" ID-Zenodo-Dep="13213040" httpUri="https://zenodo.org/record/13213040/files/figure.png" pageId="5" pageNumber="262" startId="5.[136,229,1049,1071]" targetBox="[151,1436,190,1025]" targetPageId="5" targetType="figure">
<paragraph id="8BC536E7AF720733D1EBFBB13565FB8D" blockId="5.[136,1451,1049,1108]" pageId="5" pageNumber="262">
<emphasis id="B90EEAF5AF720733D1EBFBB131EBFBF6" bold="true" box="[136,252,1049,1071]" pageId="5" pageNumber="262">FIGURE 4.</emphasis>
Comparison of pink-flowered Asia-mainland
<taxonomicName id="4C7A4D64AF720733D3D2FBB232B7FBF7" box="[689,928,1049,1072]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="B90EEAF5AF720733D3D2FBB2322DFBF6" box="[689,826,1050,1071]" italics="true" pageId="5" pageNumber="262">Holcoglossum</emphasis>
species.
<emphasis id="B90EEAF5AF720733D2ECFBB132B7FBF7" bold="true" box="[911,928,1049,1070]" pageId="5" pageNumber="262">A</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7A4D64AF720733D2C8FBB23548FBF6" box="[939,1119,1050,1071]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="semiteritifolium">
<emphasis id="B90EEAF5AF720733D2C8FBB23548FBF6" box="[939,1119,1050,1071]" italics="true" pageId="5" pageNumber="262">H. semiteritifolium</emphasis>
</taxonomicName>
.
<emphasis id="B90EEAF5AF720733D509FBB2356DFBF6" bold="true" box="[1130,1146,1050,1071]" pageId="5" pageNumber="262">B</emphasis>
.
<taxonomicName id="4C7A4D64AF720733D5E6FBB23405FBF6" authorityName="Averyanov" authorityYear="1988" baseAuthorityName="Deb, Sengupta &amp; Malick" box="[1157,1298,1050,1071]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF720733D5E6FBB23405FBF6" box="[1157,1298,1050,1071]" italics="true" pageId="5" pageNumber="262">H. himalaicum</emphasis>
</taxonomicName>
.
<emphasis id="B90EEAF5AF720733D47EFBB13439FBF6" bold="true" box="[1309,1326,1049,1071]" pageId="5" pageNumber="262">C</emphasis>
.
<taxonomicName id="4C7A4D64AF720733D45AFBB234B2FBF6" box="[1337,1445,1050,1071]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF720733D45AFBB234B2FBF6" box="[1337,1445,1050,1071]" italics="true" pageId="5" pageNumber="262">H. clausum</emphasis>
</taxonomicName>
. [Compilation by C. Bandara, picture of
<taxonomicName id="4C7A4D64AF720733D364FB9633A8FB8A" authorityName="R. Rice" authorityYear="2018" baseAuthorityName="Seidenfaden" baseAuthorityYear="1970" box="[519,703,1086,1107]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF720733D364FB9633A8FB8A" box="[519,703,1086,1107]" italics="true" pageId="5" pageNumber="262">H. semiteretifolium</emphasis>
</taxonomicName>
by K. Keller, other pictures by K. P. Wojtas.]
</paragraph>
</caption>
<paragraph id="8BC536E7AF720730D1DEF99A345DFE25" blockId="5.[136,1452,1154,2081]" lastBlockId="6.[136,1452,158,509]" lastPageId="6" lastPageNumber="263" pageId="5" pageNumber="262">
<emphasis id="B90EEAF5AF720733D1DEF99A30E5F995" bold="true" box="[189,498,1586,1612]" pageId="5" pageNumber="262">Other Specimens studied</emphasis>
:
<emphasis id="B90EEAF5AF720733D365F99A338AF995" bold="true" box="[518,669,1586,1612]" pageId="5" pageNumber="262">
<collectingCountry id="F36D7677AF720733D365F99A338AF995" box="[518,669,1586,1612]" name="Thailand" pageId="5" pageNumber="262">THAILAND</collectingCountry>
</emphasis>
.
<collectingRegion id="49BEF805AF720733D3D3F99B32C3F995" box="[688,980,1586,1613]" country="Thailand" name="Mae Hong Son" pageId="5" pageNumber="262">Mae Hong Son province</collectingRegion>
, Doi Khun Huay Pong,
<quantity id="4C829B02AF720733D599F99A3446F994" box="[1274,1361,1586,1613]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="262" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFC41027AF720733D401F99A31D4F9A9" pageId="5" pageNumber="262" value="1968-03">March 1968</date>
,
<emphasis id="B90EEAF5AF720733D1B2F9FE3021F9A9" box="[209,310,1622,1648]" italics="true" pageId="5" pageNumber="262">GT 7276</emphasis>
(C!-
<typeStatus id="54C18845AF720733D005F9FE30C0F9A9" box="[358,471,1622,1648]" pageId="5" pageNumber="262" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C7A4D64AF720733D361F9FF33CBF9A9" authorityName="R. Rice" authorityYear="2018" baseAuthorityName="Seidenfaden" baseAuthorityYear="1970" box="[514,732,1622,1648]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF720733D361F9FF33CBF9A9" box="[514,732,1622,1648]" italics="true" pageId="5" pageNumber="262">H. semiteretifolium</emphasis>
</taxonomicName>
, images seen);
<collectingRegion id="49BEF805AF720733D2F0F9FE359CF9A9" box="[915,1163,1622,1648]" country="Thailand" name="Chiang Mai" pageId="5" pageNumber="262">Chiang Mai Province</collectingRegion>
, Doi Inthanond,
<quantity id="4C829B02AF720733D433F9FE34B0F9A8" box="[1360,1447,1622,1649]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="262" unit="m" value="1900.0">1900 m</quantity>
,
<emphasis id="B90EEAF5AF720733D1EBF9D330BCF94D" box="[136,427,1658,1686]" italics="true" pageId="5" pageNumber="262">LØjtnant &amp; Chalawit 143</emphasis>
(spirit number 9606) (C!,
<taxonomicName id="4C7A4D64AF720733D381F9D332A6F94D" box="[738,945,1658,1684]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="semitertifolium">
<emphasis id="B90EEAF5AF720733D381F9D332A6F94D" box="[738,945,1658,1684]" italics="true" pageId="5" pageNumber="262">H. semitertifolium</emphasis>
</taxonomicName>
, images seen);
<emphasis id="B90EEAF5AF720733D50BF9D235E8F94D" bold="true" box="[1128,1279,1658,1684]" pageId="5" pageNumber="262">
<collectingCountry id="F36D7677AF720733D50BF9D235E8F94D" box="[1128,1279,1658,1684]" name="Myanmar" pageId="5" pageNumber="262">MYANMAR</collectingCountry>
</emphasis>
. Sima,
<collectingRegion id="49BEF805AF720733D439F9D234BCF94D" box="[1370,1451,1658,1684]" country="Myanmar" name="Kachin" pageId="5" pageNumber="262">Kachin</collectingRegion>
Hills,
<quantity id="4C829B02AF720733D1ADF9363032F960" box="[206,293,1694,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.9809999999999999" pageId="5" pageNumber="262" unit="m" value="1981.0">1981 m</quantity>
,
<date id="FFC41027AF720733D050F93730D9F961" box="[307,462,1694,1721]" pageId="5" pageNumber="262" value="1900-01">January 1900</date>
,
<emphasis id="B90EEAF5AF720733D0B8F9363382F961" box="[475,661,1694,1720]" italics="true" pageId="5" pageNumber="262">Shaik Mokim 13</emphasis>
(
<emphasis id="B90EEAF5AF720733D3C4F9363270F961" box="[679,871,1694,1720]" italics="true" pageId="5" pageNumber="262">CAL0000000490</emphasis>
, https://ivh.bsi.gov.in/phanerogams-Details/en?link =CAL0000000490&amp;column=szBarcode) (CAL!-
<typeStatus id="54C18845AF720733D3CAF96A3206F905" box="[681,785,1730,1756]" pageId="5" pageNumber="262" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7A4D64AF720733D254F96A3544F905" box="[823,1107,1730,1756]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF720733D254F96A3544F905" box="[823,1107,1730,1756]" italics="true" pageId="5" pageNumber="262">Saccolabium himalaicum</emphasis>
</taxonomicName>
, image seen);
<emphasis id="B90EEAF5AF720733D594F96A343DF905" box="[1271,1322,1730,1756]" italics="true" pageId="5" pageNumber="262">ibid.</emphasis>
,
<date id="FFC41027AF720733D45BF96A31D4F8D9" pageId="5" pageNumber="262" value="1899-12">December 1899</date>
,
<emphasis id="B90EEAF5AF720733D1ADF94E3092F8D9" box="[206,389,1766,1792]" italics="true" pageId="5" pageNumber="262">Shaik Mokim 71</emphasis>
, (
<emphasis id="B90EEAF5AF720733D0FEF94E334BF8D9" box="[413,604,1766,1792]" italics="true" pageId="5" pageNumber="262">CAL0000000486</emphasis>
, https://ivh.bsi.gov.in/phanerogams-Details/en?link=CAL0000000486&amp;col umn=szBarcode;
<emphasis id="B90EEAF5AF720733D02CF8A23318F8FD" box="[335,527,1802,1828]" italics="true" pageId="5" pageNumber="262">CAL0000000489</emphasis>
, https://ivh.bsi.gov.in/phanerogams-Details/en?link=CAL0000000489&amp;column=s zBarcode) (CAL!-
<typeStatus id="54C18845AF720733D032F88630D5F891" box="[337,450,1838,1864]" pageId="5" pageNumber="262" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C7A4D64AF720733D089F8863385F891" box="[490,658,1838,1864]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="hamalaicum">
<emphasis id="B90EEAF5AF720733D089F8863385F891" box="[490,658,1838,1864]" italics="true" pageId="5" pageNumber="262">S. hamalaicum</emphasis>
</taxonomicName>
, images seen);
<collectingRegion id="49BEF805AF720733D227F886328FF891" box="[836,920,1838,1864]" country="Myanmar" name="Kachin" pageId="5" pageNumber="262">Kachin</collectingRegion>
, Putao,
<quantity id="4C829B02AF720733D290F886355DF890" box="[1011,1098,1838,1865]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="262" unit="m" value="1220.0">1220 m</quantity>
,
<date id="FFC41027AF720733D534F886340EF890" box="[1111,1305,1838,1865]" pageId="5" pageNumber="262" value="1953-10-29">29 October 1953</date>
,
<emphasis id="B90EEAF5AF720733D446F8873005F8B5" italics="true" pageId="5" pageNumber="262">F. Kingdon-Ward 21528</emphasis>
, (
<emphasis id="B90EEAF5AF720733D04EF8FA30C4F8B5" box="[301,467,1874,1900]" italics="true" pageId="5" pageNumber="262">BM000539014</emphasis>
, https://data.nhm.ac.uk/object/32e4fd7a-9106-4de8-8f23-bffe66455794/168860160000 0) (BM!
<taxonomicName id="4C7A4D64AF720733D18CF8DF3080F849" box="[239,407,1910,1936]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF720733D18CF8DF3080F849" box="[239,407,1910,1936]" italics="true" pageId="5" pageNumber="262">H. himalaicum</emphasis>
</taxonomicName>
, image seen)
<emphasis id="B90EEAF5AF720733D354F8DE33BAF849" bold="true" box="[567,685,1910,1936]" pageId="5" pageNumber="262">
<collectingCountry id="F36D7677AF720733D354F8DE33BAF849" box="[567,685,1910,1936]" name="Bhutan" pageId="5" pageNumber="262">BHUTAN</collectingCountry>
</emphasis>
.
<collectingRegion id="49BEF805AF720733D3DFF8DE3220F849" box="[700,823,1910,1936]" country="Bhutan" name="Shemgang" pageId="5" pageNumber="262">Zhemgang</collectingRegion>
.
<date id="FFC41027AF720733D227F8DE3535F849" box="[836,1058,1910,1936]" pageId="5" pageNumber="262" value="1964-11-18">18 November 1964</date>
,
<emphasis id="B90EEAF5AF720733D54CF8DF35DBF849" box="[1071,1228,1910,1936]" italics="true" pageId="5" pageNumber="262">D.B. Deb 424</emphasis>
, (
<emphasis id="B90EEAF5AF720733D586F8DE34B3F849" box="[1253,1444,1910,1936]" italics="true" pageId="5" pageNumber="262">CAL0000000487</emphasis>
, https://ivh.bsi.gov.in/phanerogams-Details/en?link=CAL0000000487&amp;column=szBarcode) (CAL! -
<typeStatus id="54C18845AF720733D598F8333449F86C" box="[1275,1374,1947,1973]" pageId="5" pageNumber="262" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7A4D64AF720733D4F5F832301BF801" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF720733D4F5F832301BF801" italics="true" pageId="5" pageNumber="262">S. himalaicum</emphasis>
</taxonomicName>
, image seen).
<emphasis id="B90EEAF5AF720733D0CFF816330AF801" bold="true" box="[428,541,1982,2008]" pageId="5" pageNumber="262">
<collectingCountry id="F36D7677AF720733D0CFF816330AF801" box="[428,541,1982,2008]" name="Taiwan" pageId="5" pageNumber="262">TAIWAN</collectingCountry>
</emphasis>
. S. mountains Koarong-Shiusuisei,
<quantity id="4C829B02AF720733D2CEF8163513F800" box="[941,1028,1982,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="262" unit="m" value="1220.0">1220 m</quantity>
,
<date id="FFC41027AF720733D56CF81635B2F801" box="[1039,1189,1982,2009]" pageId="5" pageNumber="262" value="1912-05-24">24 May 1912</date>
,
<emphasis id="B90EEAF5AF720733D5CCF8173441F801" box="[1199,1366,1982,2008]" italics="true" pageId="5" pageNumber="262">W.R. Price 551</emphasis>
(K!,
<taxonomicName id="4C7A4D64AF720733D4F3F81731FDF825" authorityName="L. J. Chen, X. J. Xiao &amp; G. Q. Zhang" baseAuthorityName="Hayata" baseAuthorityYear="1906" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="pumilum">
<emphasis id="B90EEAF5AF720733D4F3F81731FDF825" italics="true" pageId="5" pageNumber="262">H. pumilum</emphasis>
</taxonomicName>
, K000891627, image seen).
<emphasis id="B90EEAF5AF720733D34AF84A3391F825" bold="true" box="[553,646,2018,2044]" pageId="5" pageNumber="262">
<collectingCountry id="F36D7677AF720733D34AF84A3391F825" box="[553,646,2018,2044]" name="China" pageId="5" pageNumber="262">CHINA</collectingCountry>
</emphasis>
.
<collectingRegion id="49BEF805AF720733D3F0F84B33FBF824" box="[659,748,2019,2045]" country="China" name="Yunnan" pageId="5" pageNumber="262">Yunnan</collectingRegion>
, Shweli-Salwin divide,
<quantity id="4C829B02AF720733D297F84A355CF824" box="[1012,1099,2018,2045]" metricMagnitude="3" metricUnit="m" metricValue="2.44" pageId="5" pageNumber="262" unit="m" value="2440.0">2440 m</quantity>
,
<date id="FFC41027AF720733D535F84A341AF825" box="[1110,1293,2018,2044]" pageId="5" pageNumber="262" value="1924-11">November 1924</date>
,
<emphasis id="B90EEAF5AF720733D474F84B34BCF825" box="[1303,1451,2018,2044]" italics="true" pageId="5" pageNumber="262">Forest 26130</emphasis>
(
<emphasis id="B90EEAF5AF720733D1F2F7AE3035F7F9" box="[145,290,2054,2080]" italics="true" pageId="5" pageNumber="262">K000891626</emphasis>
, http://specimens.kew.org/herbarium/K000891626) (K!-
<typeStatus id="54C18845AF720733D2EDF7AE32E1F7F9" box="[910,1014,2054,2080]" pageId="5" pageNumber="262" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7A4D64AF720733D575F7AF3583F7F9" authorityName="Z. H. Tsi" authorityYear="1982" box="[1046,1172,2055,2080]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="B90EEAF5AF720733D575F7AF3583F7F9" box="[1046,1172,2055,2080]" italics="true" pageId="5" pageNumber="262">H.junceum</emphasis>
</taxonomicName>
,image seen), duplicate at E (
<emphasis id="B90EEAF5AF710730D1C4FF36303FFF61" box="[167,296,158,184]" italics="true" pageId="6" pageNumber="263">E00123460</emphasis>
, https://data.rbge.org.uk/herb/E00123460) (E!-
<typeStatus id="54C18845AF710730D24EFF363281FF61" box="[813,918,158,184]" pageId="6" pageNumber="263" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7A4D64AF710730D2D5FF373523FF61" authorityName="Z. H. Tsi" authorityYear="1982" box="[950,1076,159,184]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="B90EEAF5AF710730D2D5FF373523FF61" box="[950,1076,159,184]" italics="true" pageId="6" pageNumber="263">H. junceum</emphasis>
</taxonomicName>
, image seen);
<collectingRegion id="49BEF805AF710730D5ACFF37343FFF60" box="[1231,1320,159,185]" country="China" name="Yunnan" pageId="6" pageNumber="263">Yunnan</collectingRegion>
, Tengchong County, Jietou municipal, Datang Cun, East of Datang on the Western slope of Gaoligong Shan,
<quantity id="4C829B02AF710730D5A9FF6A3436FF04" box="[1226,1313,194,221]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="263" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FFC41027AF710730D44DFF6A31D4FED9" pageId="6" pageNumber="263" value="1998-10-30">30 October 1998</date>
,
<emphasis id="B90EEAF5AF710730D1AFFF4F339DFED9" box="[204,650,230,257]" italics="true" pageId="6" pageNumber="263">H. Li, B. Bartholomew &amp; Z. Dhao 11286</emphasis>
(
<emphasis id="B90EEAF5AF710730D3FBFF4E3209FED9" box="[664,798,230,256]" italics="true" pageId="6" pageNumber="263">
<taxonomicName id="4C7A4D64AF710730D3FBFF4E33DEFED9" authorityName="Baker &amp; Wharton" authorityYear="1952" box="[664,713,230,256]" class="Arachnida" family="Rhinonyssidae" genus="Cas" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="263" phylum="Arthropoda" rank="genus">CAS</taxonomicName>
629868
</emphasis>
, https://www.cch2.org/portal/collections/individual/index. php?occid=4662960&amp;clid=0) (
<taxonomicName id="4C7A4D64AF710730D087FEA23334FEFD" authorityName="Baker &amp; Wharton" authorityYear="1952" box="[484,547,266,292]" class="Arachnida" family="Rhinonyssidae" genus="Cas" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="263" phylum="Arthropoda" rank="genus">CAS</taxonomicName>
!,
<taxonomicName id="4C7A4D64AF710730D350FEA333A3FEFD" authorityName="Z. H. Tsi" authorityYear="1982" box="[563,692,267,292]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="B90EEAF5AF710730D350FEA333A3FEFD" box="[563,692,267,292]" italics="true" pageId="6" pageNumber="263">H. junceum</emphasis>
</taxonomicName>
, image seen);
<collectingRegion id="49BEF805AF710730D234FEA332A7FEFC" box="[855,944,267,293]" country="China" name="Yunnan" pageId="6" pageNumber="263">Yunnan</collectingRegion>
, Kunming,
<date id="FFC41027AF710730D550FEA23418FEFD" box="[1075,1295,266,292]" pageId="6" pageNumber="263" value="1980-11-24">24 November 1980</date>
,
<emphasis id="B90EEAF5AF710730D478FEA334BCFEFD" box="[1307,1451,266,292]" italics="true" pageId="6" pageNumber="263">Z.H.Tsi 0076</emphasis>
(
<emphasis id="B90EEAF5AF710730D1F2FE863033FE91" box="[145,292,302,328]" italics="true" pageId="6" pageNumber="263">PE00200781</emphasis>
, http://nsii.org.cn/2017/specimen.php?id=6475664) (PE!
<taxonomicName id="4C7A4D64AF710730D2CAFE87353DFE91" authorityName="Z. H. Tsi" authorityYear="1982" box="[937,1066,303,328]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="B90EEAF5AF710730D2CAFE87353DFE91" box="[937,1066,303,328]" italics="true" pageId="6" pageNumber="263">H. junceum</emphasis>
</taxonomicName>
, image seen),
<collectingRegion id="49BEF805AF710730D5A9FE873434FE90" box="[1226,1315,303,329]" country="China" name="Yunnan" pageId="6" pageNumber="263">Yunnan</collectingRegion>
, Lianghe,
<date id="FFC41027AF710730D4F2FE86305BFEB4" pageId="6" pageNumber="263" value="2015-11-10">10 November 2015</date>
,
<emphasis id="B90EEAF5AF710730D001FEFA30F9FEB5" box="[354,494,338,364]" italics="true" pageId="6" pageNumber="263">5331220805</emphasis>
(
<emphasis id="B90EEAF5AF710730D36AFEFA33B8FEB5" box="[521,687,338,364]" italics="true" pageId="6" pageNumber="263">IMDY0042615</emphasis>
, http://nsii.org.cn/2017/specimen.php?id=18358911) (IMDY!
<taxonomicName id="4C7A4D64AF710730D4F3FEFB301BFE49" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF710730D4F3FEFB301BFE49" italics="true" pageId="6" pageNumber="263">H. himalaicum</emphasis>
</taxonomicName>
, image seen);
<taxonomicName id="4C7A4D64AF710730D0CCFEDE3316FE49" box="[431,513,374,400]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="genus">
<emphasis id="B90EEAF5AF710730D0CCFEDE3316FE49" bold="true" box="[431,513,374,400]" pageId="6" pageNumber="263">INDIA</emphasis>
</taxonomicName>
. Between Ukhrul and Tusom Khullen,
<date id="FFC41027AF710730D2DAFEDE3583FE49" box="[953,1172,374,400]" pageId="6" pageNumber="263" value="1948-11-21">21 November 1948</date>
,
<emphasis id="B90EEAF5AF710730D5FCFEDF34BCFE49" box="[1183,1451,374,400]" italics="true" pageId="6" pageNumber="263">F. Kingdon-Ward 18339</emphasis>
(
<emphasis id="B90EEAF5AF710730D1F2FE323034FE6D" box="[145,291,410,436]" italics="true" pageId="6" pageNumber="263">NY02650866</emphasis>
, https://sweetgum.nybg.org/science/vh/specimen-details/?irn=3377419) (NY!,
<taxonomicName id="4C7A4D64AF710730D5C9FE333443FE6D" box="[1194,1364,410,436]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF710730D5C9FE333443FE6D" box="[1194,1364,410,436]" italics="true" pageId="6" pageNumber="263">H. himalaicum</emphasis>
</taxonomicName>
, image seen);
<collectingRegion id="49BEF805AF710730D1B2FE163037FE01" box="[209,288,446,472]" country="India" name="Assam" pageId="6" pageNumber="263">Assam</collectingRegion>
, Karong,
<collectingRegion id="49BEF805AF710730D0EDFE1630E7FE01" box="[398,496,446,472]" country="India" name="Manipur" pageId="6" pageNumber="263">Manipur</collectingRegion>
,
<quantity id="4C829B02AF710730D098FE163345FE00" box="[507,594,446,473]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="263" unit="m" value="1070.0">1070 m</quantity>
,
<date id="FFC41027AF710730D33CFE16322CFE00" box="[607,827,446,473]" pageId="6" pageNumber="263" value="1950-11-25">25 November 1950</date>
,
<emphasis id="B90EEAF5AF710730D225FE1632C4FE01" box="[838,979,446,472]" italics="true" pageId="6" pageNumber="263">Koelz 27016</emphasis>
(
<emphasis id="B90EEAF5AF710730D287FE163599FE01" box="[996,1166,446,472]" italics="true" pageId="6" pageNumber="263">MICH1492888</emphasis>
, https://midwestherbaria. org/portal/collections/individual/index.php?occid=30239881&amp;clid=0) (MICH!,
<taxonomicName id="4C7A4D64AF710730D56BFE4B35B9FE25" box="[1032,1198,482,508]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF710730D56BFE4B35B9FE25" box="[1032,1198,482,508]" italics="true" pageId="6" pageNumber="263">H. himalaicum</emphasis>
</taxonomicName>
, image seen).
</paragraph>
<caption id="DF05666FAF710730D1EBFD823466FD9C" box="[136,1393,554,581]" pageId="6" pageNumber="263" startId="6.[136,229,554,580]" targetType="table">
<paragraph id="8BC536E7AF710730D1EBFD823466FD9C" blockId="6.[136,1393,554,581]" box="[136,1393,554,581]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1EBFD823016FD9D" bold="true" box="[136,257,554,580]" pageId="6" pageNumber="263">TABLE 1.</emphasis>
Morphological comparison between
<taxonomicName id="4C7A4D64AF710730D3C0FD8232B0FD9D" box="[675,935,554,580]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF710730D3C0FD8232B0FD9D" box="[675,935,554,580]" italics="true" pageId="6" pageNumber="263">Holcoglossum clausum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7A4D64AF710730D2D5FD83354BFD9D" box="[950,1116,554,581]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF710730D2D5FD8332DDFD9D" box="[950,970,555,580]" italics="true" pageId="6" pageNumber="263">H</emphasis>
.
<emphasis id="B90EEAF5AF710730D2BBFD82354BFD9D" box="[984,1116,554,580]" italics="true" pageId="6" pageNumber="263">himalaicum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A4D64AF710730D5F0FD83347DFD9D" authorityName="R. Rice" authorityYear="2018" baseAuthorityName="Seidenfaden" baseAuthorityYear="1970" box="[1171,1386,554,580]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF710730D5F0FD83347DFD9D" box="[1171,1386,554,580]" italics="true" pageId="6" pageNumber="263">H. semiteretifolium</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC536E7AF710730D1F0FDFD3565FB62" pageId="6" pageNumber="263">
<table id="F97AC447AF71F8C9D1F0FDFD3483FB62" box="[147,1428,597,1211]" gridcols="4" gridrows="16" pageId="6" pageNumber="263">
<tr id="354A34A5AF71F8C9D1F0FDFD3483FDB2" box="[147,1428,597,619]" gridrow="0" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FDFD3057FDB2" box="[147,320,597,619]" gridcol="0" gridrow="0" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1F0FDFD31EBFDB2" bold="true" box="[147,252,597,619]" pageId="6" pageNumber="263">Character</emphasis>
</th>
<th id="769B5DD9AF71F8C9D000FDFD33B0FDB2" box="[355,679,597,619]" gridcol="1" gridrow="0" pageId="6" pageNumber="263">
<taxonomicName id="4C7A4D64AF710730D000FDFE30C1FDB2" box="[355,470,597,619]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF710730D000FDFE30C1FDB2" bold="true" box="[355,470,597,619]" italics="true" pageId="6" pageNumber="263">H. clausum</emphasis>
</taxonomicName>
</th>
<th id="769B5DD9AF71F8C9D3ABFDFD351EFDB2" box="[712,1033,597,619]" gridcol="2" gridrow="0" pageId="6" pageNumber="263">
<taxonomicName id="4C7A4D64AF710730D3ABFDFE3249FDB2" box="[712,862,597,619]" class="Liliopsida" family="Orchidaceae" genus="Saccolabium" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="himalaicum">
<emphasis id="B90EEAF5AF710730D3ABFDFE3249FDB2" bold="true" box="[712,862,597,619]" italics="true" pageId="6" pageNumber="263">H. himalaicum</emphasis>
</taxonomicName>
</th>
<th id="769B5DD9AF71F8C9D550FDFD3483FDB2" box="[1075,1428,597,619]" gridcol="3" gridrow="0" pageId="6" pageNumber="263">
<taxonomicName id="4C7A4D64AF710730D550FDFE35E4FDB2" authority="Growth" authorityName="Growth" box="[1075,1267,597,619]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF710730D550FDFE35E4FDB2" bold="true" box="[1075,1267,597,619]" italics="true" pageId="6" pageNumber="263">H. semiteretifolium</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="354A34A5AF71F8C9D1F0FDD53483FD4A" box="[147,1428,637,659]" gridrow="1" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FDD53057FD4A" box="[147,320,637,659]" gridcol="0" gridrow="1" pageId="6" pageNumber="263">
Growth
<taxonomicName id="4C7A4D64AF710730D180FDD53005FD4A" box="[227,274,637,659]" form="pendant" pageId="6" pageNumber="263" rank="form">form</taxonomicName>
</th>
<td id="769B5DD9AF71F8C9D000FDD533B0FD4A" box="[355,679,637,659]" gridcol="1" gridrow="1" pageId="6" pageNumber="263">pendant</td>
<td id="769B5DD9AF71F8C9D3ABFDD5351EFD4A" box="[712,1033,637,659]" gridcol="2" gridrow="1" pageId="6" pageNumber="263">pendant</td>
<td id="769B5DD9AF71F8C9D550FDD53483FD4A" box="[1075,1428,637,659]" gridcol="3" gridrow="1" pageId="6" pageNumber="263">erect</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FD0C3483FD63" box="[147,1428,676,698]" gridrow="2" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FD0C3057FD63" box="[147,320,676,698]" gridcol="0" gridrow="2" pageId="6" pageNumber="263">Stem</th>
<td id="769B5DD9AF71F8C9D000FD0C33B0FD63" box="[355,679,676,698]" gridcol="1" gridrow="2" pageId="6" pageNumber="263">24 cm</td>
<td id="769B5DD9AF71F8C9D3ABFD0C351EFD63" box="[712,1033,676,698]" gridcol="2" gridrow="2" pageId="6" pageNumber="263">924 cm</td>
<td id="769B5DD9AF71F8C9D550FD0C3483FD63" box="[1075,1428,676,698]" gridcol="3" gridrow="2" pageId="6" pageNumber="263">5 cm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FD633483FD38" box="[147,1428,715,737]" gridrow="3" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FD633057FD38" box="[147,320,715,737]" gridcol="0" gridrow="3" pageId="6" pageNumber="263">Leaves</th>
<td id="769B5DD9AF71F8C9D000FD6333B0FD38" box="[355,679,715,737]" gridcol="1" gridrow="3" pageId="6" pageNumber="263">subterete, 9.545 cm</td>
<td id="769B5DD9AF71F8C9D3ABFD63351EFD38" box="[712,1033,715,737]" gridcol="2" gridrow="3" pageId="6" pageNumber="263">subterete, 3550 cm</td>
<td id="769B5DD9AF71F8C9D550FD633483FD38" box="[1075,1428,715,737]" gridcol="3" gridrow="3" pageId="6" pageNumber="263">semiterete, up to 15 cm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FD5B3483FCD0" box="[147,1428,755,777]" gridrow="4" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FD5B3057FCD0" box="[147,320,755,777]" gridcol="0" gridrow="4" pageId="6" pageNumber="263">Inflorescence</th>
<td id="769B5DD9AF71F8C9D000FD5B33B0FCD0" box="[355,679,755,777]" gridcol="1" gridrow="4" pageId="6" pageNumber="263">1.67.0 cm</td>
<td id="769B5DD9AF71F8C9D3ABFD5B351EFCD0" box="[712,1033,755,777]" gridcol="2" gridrow="4" pageId="6" pageNumber="263">415 cm</td>
<td id="769B5DD9AF71F8C9D550FD5B3483FCD0" box="[1075,1428,755,777]" gridcol="3" gridrow="4" pageId="6" pageNumber="263">89 cm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FCB23483FCE9" box="[147,1428,794,816]" gridrow="5" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FCB23057FCE9" box="[147,320,794,816]" gridcol="0" gridrow="5" pageId="6" pageNumber="263">Pedicel and ovary</th>
<td id="769B5DD9AF71F8C9D000FCB233B0FCE9" box="[355,679,794,816]" gridcol="1" gridrow="5" pageId="6" pageNumber="263">1.7 cm</td>
<td id="769B5DD9AF71F8C9D3ABFCB2351EFCE9" box="[712,1033,794,816]" gridcol="2" gridrow="5" pageId="6" pageNumber="263">1.21.5 cm</td>
<td id="769B5DD9AF71F8C9D550FCB23483FCE9" box="[1075,1428,794,816]" gridcol="3" gridrow="5" pageId="6" pageNumber="263">1.01.2 cm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FCEA3483FC8E" box="[147,1428,834,855]" gridrow="6" pageId="6" pageNumber="263" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="769B5DD9AF71F8C9D1F0FCEA3057FC8E" box="[147,320,834,855]" gridcol="0" gridrow="6" pageId="6" pageNumber="263">Racemose</th>
</tr>
<tr id="354A34A5AF71F8C9D1F0FCF33483FCA8" box="[147,1428,859,881]" gridrow="7" pageId="6" pageNumber="263" rowspan-0="1">
<td id="769B5DD9AF71F8C9D000FCF333B0FCA8" box="[355,679,859,881]" gridcol="1" gridrow="7" pageId="6" pageNumber="263">417 flowers</td>
<td id="769B5DD9AF71F8C9D3ABFCF3351EFCA8" box="[712,1033,859,881]" gridcol="2" gridrow="7" pageId="6" pageNumber="263">1539 flowers</td>
<td id="769B5DD9AF71F8C9D550FCF33483FCA8" box="[1075,1428,859,881]" gridcol="3" gridrow="7" pageId="6" pageNumber="263">515 flowers</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FCDC3483FC53" box="[147,1428,884,906]" gridrow="8" pageId="6" pageNumber="263" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="769B5DD9AF71F8C9D1F0FCDC3057FC53" box="[147,320,884,906]" gridcol="0" gridrow="8" pageId="6" pageNumber="263">inflorescences</th>
</tr>
<tr id="354A34A5AF71F8C9D1F0FC333483FC68" box="[147,1428,923,945]" gridrow="9" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FC333057FC68" box="[147,320,923,945]" gridcol="0" gridrow="9" pageId="6" pageNumber="263">Sepals</th>
<td id="769B5DD9AF71F8C9D000FC3333B0FC68" box="[355,679,923,945]" gridcol="1" gridrow="9" pageId="6" pageNumber="263">7.09.0 mm × 2.44.2 mm</td>
<td id="769B5DD9AF71F8C9D3ABFC33351EFC68" box="[712,1033,923,945]" gridcol="2" gridrow="9" pageId="6" pageNumber="263">4.05.0 mm × 2.02.8 mm</td>
<td id="769B5DD9AF71F8C9D550FC333483FC68" box="[1075,1428,923,945]" gridcol="3" gridrow="9" pageId="6" pageNumber="263">6.07.0 mm × 4 mm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FC6A3483FC00" box="[147,1428,962,985]" gridrow="10" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FC6A3057FC00" box="[147,320,962,985]" gridcol="0" gridrow="10" pageId="6" pageNumber="263">Petals</th>
<td id="769B5DD9AF71F8C9D000FC6A33B0FC00" box="[355,679,962,985]" gridcol="1" gridrow="10" pageId="6" pageNumber="263">7.17.2 mm × 3.03.1 mm</td>
<td id="769B5DD9AF71F8C9D3ABFC6A351EFC00" box="[712,1033,962,985]" gridcol="2" gridrow="10" pageId="6" pageNumber="263">4.05.0 mm × 2.22.4 mm</td>
<td id="769B5DD9AF71F8C9D550FC6A3483FC00" box="[1075,1428,962,985]" gridcol="3" gridrow="10" pageId="6" pageNumber="263">6.07.0 mm × 4 mm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FC423483FBD9" box="[147,1428,1002,1024]" gridrow="11" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FC423057FBD9" box="[147,320,1002,1024]" gridcol="0" gridrow="11" pageId="6" pageNumber="263">Midlobe</th>
<td id="769B5DD9AF71F8C9D000FC4233B0FBD9" box="[355,679,1002,1024]" gridcol="1" gridrow="11" pageId="6" pageNumber="263">5.56 mm × 3.5 mm</td>
<td id="769B5DD9AF71F8C9D3ABFC42351EFBD9" box="[712,1033,1002,1024]" gridcol="2" gridrow="11" pageId="6" pageNumber="263">3.04.0 mm × 2.3 mm</td>
<td id="769B5DD9AF71F8C9D550FC423483FBD9" box="[1075,1428,1002,1024]" gridcol="3" gridrow="11" pageId="6" pageNumber="263">4.0 × 2.5 mm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FBB93483FB9C" box="[147,1428,1041,1093]" gridrow="12" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FBB93057FB9C" box="[147,320,1041,1093]" gridcol="0" gridrow="12" pageId="6" pageNumber="263">Side lobes</th>
<td id="769B5DD9AF71F8C9D000FBB933B0FB9C" box="[355,679,1041,1093]" gridcol="1" gridrow="12" pageId="6" pageNumber="263">white and smooth, 1.61.7 mm × 1.51.6 mm</td>
<td id="769B5DD9AF71F8C9D3ABFBB9351EFB9C" box="[712,1033,1041,1093]" gridcol="2" gridrow="12" pageId="6" pageNumber="263">pink with callus at the base, 2.0 2.5 mm × 1.21.5 mm</td>
<td id="769B5DD9AF71F8C9D550FBB93483FB9C" box="[1075,1428,1041,1093]" gridcol="3" gridrow="12" pageId="6" pageNumber="263">greenish-yellow and smooth, ca. 2.0 mm × 2.6 mm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FBFE3483FBB5" box="[147,1428,1110,1132]" gridrow="13" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FBFE3057FBB5" box="[147,320,1110,1132]" gridcol="0" gridrow="13" pageId="6" pageNumber="263">Spur length</th>
<td id="769B5DD9AF71F8C9D000FBFE33B0FBB5" box="[355,679,1110,1132]" gridcol="1" gridrow="13" pageId="6" pageNumber="263">1.11.3 cm</td>
<td id="769B5DD9AF71F8C9D3ABFBFE351EFBB5" box="[712,1033,1110,1132]" gridcol="2" gridrow="13" pageId="6" pageNumber="263">1.21.5 cm</td>
<td id="769B5DD9AF71F8C9D550FBFE3483FBB5" box="[1075,1428,1110,1132]" gridcol="3" gridrow="13" pageId="6" pageNumber="263">0.50.6 cm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FBD53483FB4A" box="[147,1428,1149,1171]" gridrow="14" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FBD53057FB4A" box="[147,320,1149,1171]" gridcol="0" gridrow="14" pageId="6" pageNumber="263">Column</th>
<td id="769B5DD9AF71F8C9D000FBD533B0FB4A" box="[355,679,1149,1171]" gridcol="1" gridrow="14" pageId="6" pageNumber="263">1.5 mm × 1.5 mm</td>
<td id="769B5DD9AF71F8C9D3ABFBD5351EFB4A" box="[712,1033,1149,1171]" gridcol="2" gridrow="14" pageId="6" pageNumber="263">2.0 mm × 2.0 mm</td>
<td id="769B5DD9AF71F8C9D550FBD53483FB4A" box="[1075,1428,1149,1171]" gridcol="3" gridrow="14" pageId="6" pageNumber="263">2.0 mm × 2.0 mm</td>
</tr>
<tr id="354A34A5AF71F8C9D1F0FB0D3483FB62" box="[147,1428,1189,1211]" gridrow="15" pageId="6" pageNumber="263">
<th id="769B5DD9AF71F8C9D1F0FB0D3057FB62" box="[147,320,1189,1211]" gridcol="0" gridrow="15" pageId="6" pageNumber="263">Flowering period</th>
<td id="769B5DD9AF71F8C9D000FB0D33B0FB62" box="[355,679,1189,1211]" gridcol="1" gridrow="15" pageId="6" pageNumber="263">NovemberDecember</td>
<td id="769B5DD9AF71F8C9D3ABFB0D351EFB62" box="[712,1033,1189,1211]" gridcol="2" gridrow="15" pageId="6" pageNumber="263">OctoberJanuary</td>
<td id="769B5DD9AF71F8C9D550FB0D3483FB62" box="[1075,1428,1189,1211]" gridcol="3" gridrow="15" pageId="6" pageNumber="263">March</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC536E7AF710730D1DEFB433211FAF0" blockId="6.[136,1452,1259,2077]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEFB4330ADFADC" bold="true" box="[189,442,1259,1285]" pageId="6" pageNumber="263">Flowering phenology</emphasis>
: Plants in flower were observed in late November (winter, dry season). Plants with pure white flowers were observed in pictures on social media.
</paragraph>
</subSubSection>
<subSubSection id="C360656CAF710730D1DEFA9B31F6FAA8" pageId="6" pageNumber="263" type="etymology">
<paragraph id="8BC536E7AF710730D1DEFA9B31F6FAA8" blockId="6.[136,1452,1259,2077]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEFA9B3029FA94" bold="true" box="[189,318,1331,1357]" pageId="6" pageNumber="263">Etymology</emphasis>
: Derived from Latin
<taxonomicName id="4C7A4D64AF710730D359FA9B3380FA94" box="[570,663,1331,1357]" class="Liliopsida" family="Orchidaceae" genus="Holcoglossum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="clausum">
<emphasis id="B90EEAF5AF710730D359FA9B3380FA94" box="[570,663,1331,1357]" italics="true" pageId="6" pageNumber="263">clausum</emphasis>
</taxonomicName>
meaning closed, referring to the barely opening flowers of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C360656CAF710731D1DEFAD3347DFEB5" lastPageId="7" lastPageNumber="264" pageId="6" pageNumber="263" type="distribution">
<paragraph id="8BC536E7AF710730D1DEFAD3346DFA4C" blockId="6.[136,1452,1259,2077]" box="[189,1402,1403,1429]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEFAD33058FA4C" bold="true" box="[189,335,1403,1429]" pageId="6" pageNumber="263">Distribution</emphasis>
:
<collectingCountry id="F36D7677AF710730D03DFAD330DCFA4C" box="[350,459,1403,1429]" name="Myanmar" pageId="6" pageNumber="263">Myanmar</collectingCountry>
(Taunggyi District,
<collectingRegion id="49BEF805AF710730D3CDFAD33230FA4C" box="[686,807,1403,1429]" country="Myanmar" name="Shan" pageId="6" pageNumber="263">Shan State</collectingRegion>
) and
<collectingCountry id="F36D7677AF710730D207FAD332DFFA4C" box="[868,968,1403,1429]" name="Thailand" pageId="6" pageNumber="263">Thailand</collectingCountry>
(Doi Inthanon,
<collectingRegion id="49BEF805AF710730D518FAD33479FA4C" box="[1147,1390,1403,1429]" country="Thailand" name="Chiang Mai" pageId="6" pageNumber="263">Chiang Mai province</collectingRegion>
).
</paragraph>
<paragraph id="8BC536E7AF710730D1DEFA373596FA60" blockId="6.[136,1452,1259,2077]" box="[189,1153,1439,1466]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEFA3730BAFA60" bold="true" box="[189,429,1439,1465]" pageId="6" pageNumber="263">Habitat and ecology</emphasis>
: Tropical mixed deciduous forest on limestone at
<quantity id="4C829B02AF710730D2BCFA37356AFA60" box="[991,1149,1439,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="6" pageNumber="263" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">15001600 m</quantity>
.
</paragraph>
<paragraph id="8BC536E7AF710730D1DEFA6B3049F8B0" blockId="6.[136,1452,1259,2077]" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEFA6B30FCFA04" bold="true" box="[189,491,1475,1501]" pageId="6" pageNumber="263">Conservation assessment</emphasis>
: Plants are known to occur in
<collectingCountry id="F36D7677AF710730D23BFA6B32D2FA04" box="[856,965,1475,1501]" name="Myanmar" pageId="6" pageNumber="263">Myanmar</collectingCountry>
(Taunggyi) and
<collectingCountry id="F36D7677AF710730D5F0FA6B35E0FA04" box="[1171,1271,1475,1501]" name="Thailand" pageId="6" pageNumber="263">Thailand</collectingCountry>
(Doi Inthanon) (
<bibRefCitation id="EFEB4B16AF710730D1ECFA4F3069F9D8" author="Ormerod, P. &amp; Kurzweil, H. &amp; Watthana, S." box="[143,382,1511,1537]" pageId="6" pageNumber="263" pagination="1 - 262" refId="ref7046" refString="Ormerod, P., Kurzweil, H. &amp; Watthana, S. (2021) Annotated List of Orchidaceae for Myanmar. Phytotaxa 481: 1 - 262. https: // doi. org / 10.11646 / phytotaxa. 481.1.1" type="journal article" year="2021">
Ormerod
<emphasis id="B90EEAF5AF710730D063FA40302DF9D8" box="[256,314,1511,1537]" italics="true" pageId="6" pageNumber="263">et al.</emphasis>
2021
</bibRefCitation>
,
<bibRefCitation id="EFEB4B16AF710730D0E8FA4F3340F9D8" author="Seidenfaden, G." box="[395,599,1511,1537]" pageId="6" pageNumber="263" pagination="1 - 356" refId="ref7293" refString="Seidenfaden, G. (1988) Orchid Genera in Thailand 14: Fifty-nine Vandoid genera. Opera Botanica 95: 1 - 356." type="journal article" year="1988">Seidenfaden 1988</bibRefCitation>
) in the Indo-Myanmar Biodiversity Hotspot. In
<collectingCountry id="F36D7677AF710730D5E1FA4F35E6F9D8" box="[1154,1265,1511,1537]" name="Myanmar" pageId="6" pageNumber="263">Myanmar</collectingCountry>
, the plants were observed growing on the branches of trees towards the open areas in a tropical mixed deciduous forest by Tanaka (pers. comm.) (see figure 5). Tanaka also informed us that locals in
<collectingRegion id="49BEF805AF710730D2F6F9873519F990" box="[917,1038,1583,1609]" country="Myanmar" name="Shan" pageId="6" pageNumber="263">Shan State</collectingRegion>
collect the plants and grow them at their homes. Plants have been observed to set fruits naturally and most likely recruit too. Plants have also been seen on display or trade on social media (Facebook). The plants in Europe, although procured from
<collectingCountry id="F36D7677AF710730D5AAF9DF343AF948" box="[1225,1325,1655,1681]" name="Thailand" pageId="6" pageNumber="263">Thailand</collectingCountry>
were most probably collected in
<collectingCountry id="F36D7677AF710730D019F93330F0F96C" box="[378,487,1691,1717]" name="Myanmar" pageId="6" pageNumber="263">Myanmar</collectingCountry>
or propagated usings seeds acquired from
<collectingCountry id="F36D7677AF710730D2A0F9333525F96C" box="[963,1074,1691,1717]" name="Myanmar" pageId="6" pageNumber="263">Myanmar</collectingCountry>
. This implies that the species is in immense threat of collection for its high ornamental value. Extent of Occurrence and Area of Occurrence is calculated in GeoCAT (
<bibRefCitation id="EFEB4B16AF710730D077F94B330CF924" author="Bachman, S. &amp; Moat, J." box="[276,539,1763,1789]" pageId="6" pageNumber="263" pagination="11 - 13" refId="ref5949" refString="Bachman, S. &amp; Moat, J. (2012) GeoCAT - An open-source tool for rapid red list assessments. BGjournal 9 (1): 11 - 13. [https: // www. jstor. org / stable / 24811237]" type="journal article" year="2012">Bachman &amp; Moat 2012</bibRefCitation>
) as
<quantity id="4C829B02AF710730D325F94B338EF924" box="[582,665,1763,1789]" metricMagnitude="5" metricUnit="m" metricValue="3.67" pageId="6" pageNumber="263" unit="km" value="367.0">367 km</quantity>
<superScript id="7C0F9BAFAF710730D3FAF94B33B6F928" attach="left" box="[665,673,1763,1777]" fontSize="6" pageId="6" pageNumber="263">2</superScript>
and
<quantity id="4C829B02AF710730D3B6F94B320CF924" box="[725,795,1763,1789]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="263" unit="km" value="16.0">16 km</quantity>
<superScript id="7C0F9BAFAF710730D278F94B3234F928" attach="left" box="[795,803,1763,1777]" fontSize="6" pageId="6" pageNumber="263">2</superScript>
respectively, but, the total number of mature individuals is not known in either country. Due to continuing decline in the number of mature individuals, and presence of 4 location, this species should be best assessed as
<emphasis id="B90EEAF5AF710730D359F88333DBF89C" bold="true" box="[570,716,1835,1861]" pageId="6" pageNumber="263">Endangered</emphasis>
[EN B1ab(v)+B2ab(v)] following
<bibRefCitation id="EFEB4B16AF710730D539F883304DF8B0" author="IUCN Standards &amp; Petitions Committee" pageId="6" pageNumber="263" refId="ref6539" refString="IUCN Standards and Petitions Committee (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed: 23 December 2023)." type="url" year="2022">IUCN Standards and Petitions Committee (2022)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC536E7AF710731D1DEF8DB347DFEB5" blockId="6.[136,1452,1259,2077]" lastBlockId="7.[136,1452,158,365]" lastPageId="7" lastPageNumber="264" pageId="6" pageNumber="263">
<emphasis id="B90EEAF5AF710730D1DEF8DB31E8F854" bold="true" box="[189,255,1907,1933]" pageId="6" pageNumber="263">Note:</emphasis>
Interestingly, the species was already shown by
<bibRefCitation id="EFEB4B16AF710730D242F8DB3538F854" author="Seidenfaden, G." box="[801,1071,1907,1933]" pageId="6" pageNumber="263" pagination="1 - 356" refId="ref7293" refString="Seidenfaden, G. (1988) Orchid Genera in Thailand 14: Fifty-nine Vandoid genera. Opera Botanica 95: 1 - 356." type="journal article" year="1988">Seidenfaden (1988: 397</bibRefCitation>
, pl. 37A) as a photograph with an erroneous determination as
<taxonomicName id="4C7A4D64AF710730D0DCF83F3279F868" authority="Seidenf" authorityName="Seidenf" box="[447,878,1943,1969]" class="Liliopsida" family="Orchidaceae" genus="Ascocentrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF710730D0DCF83F3219F868" box="[447,782,1943,1969]" italics="true" pageId="6" pageNumber="263">Ascocentrum semiteretifolium</emphasis>
Seidenf
</taxonomicName>
.. We believe that the specimen
<emphasis id="B90EEAF5AF710730D5ACF83F3425F868" box="[1231,1330,1943,1969]" italics="true" pageId="6" pageNumber="263">GT 8289</emphasis>
, deposited at C, is depicted in this photograph. Seidenfaden annotated on the specimen “near
<taxonomicName id="4C7A4D64AF710730D574F81335DBF80C" box="[1047,1228,1979,2005]" class="Liliopsida" family="Orchidaceae" genus="Ascocentrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF710730D574F81335DBF80C" box="[1047,1228,1979,2005]" italics="true" pageId="6" pageNumber="263">semiteretifolium</emphasis>
</taxonomicName>
”. This suggests that he may have recognized it as a distinct species. Nevertheless, it remains unclear why he did not describe it or why it was listed under
<taxonomicName id="4C7A4D64AF710730D026F7AB330DF7C4" box="[325,538,2051,2077]" class="Liliopsida" family="Orchidaceae" genus="Ascocentrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="263" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF710730D026F7AB330DF7C4" box="[325,538,2051,2077]" italics="true" pageId="6" pageNumber="263">A. semiteretifolium</emphasis>
</taxonomicName>
. However, there is no further information on the container of the specimen or the inlying note.
<bibRefCitation id="EFEB4B16AF700731D07FFF363327FF61" author="Seidenfaden, G." box="[284,560,158,184]" pageId="7" pageNumber="264" pagination="1 - 356" refId="ref7293" refString="Seidenfaden, G. (1988) Orchid Genera in Thailand 14: Fifty-nine Vandoid genera. Opera Botanica 95: 1 - 356." type="journal article" year="1988">Seidenfaden (1988: 317)</bibRefCitation>
cited this specimen in Opera Botanica 95, giving “
<collectingRegion id="49BEF805AF700731D50BFF3635E5FF61" box="[1128,1266,158,184]" country="Thailand" name="Chiang Mai" pageId="7" pageNumber="264">Chiang Mai</collectingRegion>
, Doi Inthanond” as its locality. But the second specimen at C, collected by Løjtnant &amp; Chalawit (no.
<emphasis id="B90EEAF5AF700731D53EFF6A3590FF05" box="[1117,1159,194,220]" italics="true" pageId="7" pageNumber="264">143</emphasis>
) from the same location, represents
<taxonomicName id="4C7A4D64AF700731D063FF4F30C0FED9" box="[256,471,230,256]" class="Liliopsida" family="Orchidaceae" genus="Ascocentrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="264" phylum="Tracheophyta" rank="species" species="semiteretifolium">
<emphasis id="B90EEAF5AF700731D063FF4F30C0FED9" box="[256,471,230,256]" italics="true" pageId="7" pageNumber="264">H. semiteretifolium</emphasis>
</taxonomicName>
and not the new species. Initially, we had doubts regarding the origin of
<emphasis id="B90EEAF5AF700731D46FFF4F3479FED9" box="[1292,1390,230,257]" italics="true" pageId="7" pageNumber="264">GT 8289</emphasis>
from the specified location also because the photograph appears to depict a cultivated plant. However, in
<date id="FFC41027AF700731D591FEA234B0FEFD" box="[1266,1447,266,292]" pageId="7" pageNumber="264" value="2023-12">December 2023</date>
, during the review process of the manuscript, we received images of flowering plants of the newly described species from Doi Inthanon, confirming the occurrence of the new species in
<collectingCountry id="F36D7677AF700731D2EBFEFA32FBFEB5" box="[904,1004,338,364]" name="Thailand" pageId="7" pageNumber="264">Thailand</collectingCountry>
(Phumsaringkharn pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>