treatments-xml/data/03/D3/87/03D387CFFFC49C33209297AFFD7AFB9A.xml

<document id="779412E84204DAFC1E1BA07BAA3CBC3A" ID-CLB-Dataset="302489" ID-DOI="10.11646/phytotaxa.343.2.11" ID-GBIF-Dataset="c6269787-0d8c-4d00-8add-afc0588ab6e1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13718587" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725654591077" checkinUser="felipe" docAuthor="Averyanov, Leonid V., Nuraliev, Maxim S., Kuznetsov, Andrey N. &amp; Kuznetsova, Svetlana P." docDate="2018" docId="03D387CFFFC49C33209297AFFD7AFB9A" docLanguage="en" docName="phytotaxa.343.2.11.pdf" docOrigin="Phytotaxa 343 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.343.2.11" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Biermannia longicheila Aver. &amp; Nuraliev 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="197" masterDocId="FFEAFFB7FFC49C30201A921BFFEFFFFB" masterDocTitle="Biermannia longicheila (Orchidaceae, Aeridinae), a new species from southern Vietnam" masterLastPageNumber="198" masterPageNumber="194" pageNumber="194" updateTime="1725665457995" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="23DA0185555CA4888F3A8F8BB2D614DB">Biermannia longicheila (Orchidaceae, Aeridinae), a new species from southern Vietnam</mods:title>
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<mods:namePart id="C9BDCDA151BA5838A2DE04AD352C3A6D">Averyanov, Leonid V.</mods:namePart>
<mods:affiliation id="77821BE9481D53751E28375543EC686F">Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov str., 197376 St. Petersburg, Russia;</mods:affiliation>
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<mods:namePart id="381E2022116F3B1DCE5CA77B012D7CC2">Nuraliev, Maxim S.</mods:namePart>
<mods:affiliation id="F13DA0E56412C4D068A5ECB44B006614">Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam &amp; Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia</mods:affiliation>
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<mods:namePart id="F328E37B75CB866EDDC07183201CBE73">Kuznetsov, Andrey N.</mods:namePart>
<mods:affiliation id="CAA5B947C2E24D64D866A16E98B9EC90">Joint Russian-Vietnamese Tropical Scientific and Technological Center, Cau Giay, Hanoi, Vietnam &amp; Severtsov Institute of Ecology and Evolution Problems of the Russian Academy of Sciences, 33, Leninsky Avenue, 119071 Moscow, Russia</mods:affiliation>
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<heading id="D08D81B5FFC49C30209297AFFCBFFA34" box="[136,848,1460,1487]" fontSize="11" level="2" pageId="0" pageNumber="194" reason="3">
<taxonomicName id="4C7A4D5AFFC49C30209297AFFD8BFA34" ID-CoL="8WRR2" authority="Aver. &amp; Nuraliev" authorityName="Aver. &amp; Nuraliev" authorityYear="2018" box="[136,612,1460,1487]" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="194" phylum="Tracheophyta" rank="species" species="longicheila" status="sp. nov.">
<emphasis id="B90EEACBFFC49C30209297AFFE74FA34" bold="true" box="[136,411,1460,1487]" italics="true" pageId="0" pageNumber="194">Biermannia longicheila</emphasis>
Aver. &amp; Nuraliev
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, 
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<taxonomicNameLabel id="A23D57B0FFC49C30227597ADFD2BFA34" box="[623,708,1462,1487]" pageId="0" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="13412A5CFFC49C3022C897ADFCFAFA34" box="[722,789,1462,1487]" captionStart="FIGURE 1" captionStartId="1.[136,229,1800,1821]" captionTargetBox="[151,1436,187,1772]" captionTargetId="figure-13@1.[151,1436,187,1772]" captionTargetPageId="1" captionText="FIGURE 1. Biermannia longicheila Aver. et Nuraliev. Digital epitype, Averyanov &amp; Nuraliev 0302 made from the type, Nuraliev et al. 1726. Photos by M. Nuraliev, correction and design by L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.13718589" httpUri="https://zenodo.org/record/13718589/files/figure.png" pageId="0" pageNumber="194">Fig. 1</figureCitation>
&amp; 
<figureCitation id="13412A5CFFC49C30232397ADFCA7FA34" box="[825,840,1462,1487]" captionStart="FIGURE 2" captionStartId="2.[136,229,1957,1978]" captionTargetBox="[216,1368,195,1925]" captionTargetId="figure-14@2.[215,1373,190,1931]" captionTargetPageId="2" captionText="FIGURE 2. Biermannia longicheila. A.Flowering plant.B. Inflorescence. C. Rachis.D. Intact flowers, frontal and side views.E. Flattened flower, lip and pedicelate ovary. F. Intact lip, view from above. G. Intact lip, side view and sagittal section. H. Intact column and column foot, frontal view. I. Intact column and column foot, side view. J. Operculum, frontal view, side view and view from below. K. Pollinarium, frontal view and view from behind. All drawn from the type (Nuraliev et al. 1726) by L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.13718591" httpUri="https://zenodo.org/record/13718591/files/figure.png" pageId="0" pageNumber="194">2</figureCitation>
)
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Type:— 
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. 
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: K’Bang District, K’rong Municipality, Kon Ka Kinh National Park, 
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NW of K’Bang Town, forest, not far from river, 
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, 
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, 
<date id="FFC41019FFC49C302307941BFC77F9EE" box="[797,920,1536,1557]" pageId="0" pageNumber="194" value="2017-05-09">9 May 2017</date>
, 
<emphasis id="B90EEACBFFC49C3023BE941BFAF4F9EE" box="[932,1307,1536,1557]" italics="true" pageId="0" pageNumber="194">Nuraliev, Kuznetsov, Kuznetsova 1726</emphasis>
(
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: 
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01041427).
</paragraph>
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Digital 
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:—Plate d-EXSICCATES 
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0302/Nuraliev 1726 (
<collectionCode id="ED6BAE1CFFC49C3023E39453FBC6F9A6" box="[1017,1065,1608,1629]" country="Netherlands" name="Museum Wasmann" pageId="0" pageNumber="194">MW</collectionCode>
: MW-DigiPic0000007) (
<figureCitation id="13412A5CFFC49C30250D9453FAA0F9A6" box="[1303,1359,1608,1629]" captionStart="FIGURE 1" captionStartId="1.[136,229,1800,1821]" captionTargetBox="[151,1436,187,1772]" captionTargetId="figure-13@1.[151,1436,187,1772]" captionTargetPageId="1" captionText="FIGURE 1. Biermannia longicheila Aver. et Nuraliev. Digital epitype, Averyanov &amp; Nuraliev 0302 made from the type, Nuraliev et al. 1726. Photos by M. Nuraliev, correction and design by L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.13718589" httpUri="https://zenodo.org/record/13718589/files/figure.png" pageId="0" pageNumber="194">Fig. 1</figureCitation>
).
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Image of 
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at 
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as well as the drawing available at https://depo.msu.ru/open/public/item/MW-DigiPic0000007 and https://depo. msu.ru/open/public/item/MW-DigiPic0000008
</paragraph>
<paragraph id="8BC536D9FFC49C30209294CDFF29F83C" blockId="0.[136,1452,1750,1991]" pageId="0" pageNumber="194">
The new species may be close to the Himalayan 
<taxonomicName id="4C7A4D5AFFC49C3022BD94CDFF51F8E8" authority="(King &amp; Pantling 1895: 340) King &amp; Pantling (1898 b: 200)" authorityName="King &amp; Pantling" authorityPageNumber="200" authorityYear="1898" baseAuthorityName="King &amp; Pantling" baseAuthorityPageNumber="340" baseAuthorityYear="1895" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="194" phylum="Tracheophyta" rank="species" species="bimaculata">
<emphasis id="B90EEACBFFC49C3022BD94CDFCABF914" box="[679,836,1750,1775]" italics="true" pageId="0" pageNumber="194">B. bimaculata</emphasis>
(
<bibRefCitation id="EFEB4B28FFC49C30234994CDFB68F914" author="King, G. &amp; Pantling, R." box="[851,1159,1750,1775]" pageId="0" pageNumber="194" pagination="329 - 344" refId="ref2166" refString="King, G. &amp; Pantling, R. (1895) On some new orchids from Sikkim. Journal of the Asiatic Society of Bengal 64: 329 - 344." type="journal article" year="1895">King &amp; Pantling 1895: 340</bibRefCitation>
) 
<bibRefCitation id="EFEB4B28FFC49C30248F94CDFF51F8E8" author="King, G. &amp; Pantling, R." pageId="0" pageNumber="194" pagination="1 - 342" refId="ref2229" refString="King, G. &amp; Pantling, R. (1898 b) The orchids of the Sikkim-Himalaya. Annals of the Royal Botanic Garden 8: 1 - 342." type="journal article" year="1898">King &amp; Pantling (1898b: 200)</bibRefCitation>
</taxonomicName>
, but differs from the latter by its twice larger, widely opening flowers about 
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across (vs. hardly opening flowers, 
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in diameter), long narrow sepals, 
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(vs. narrowly ovate sepals, 
<quantity id="4C829B3CFFC49C30248C9505FB17F8CC" box="[1174,1272,1822,1847]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="0" pageNumber="194" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
), long narrowly conoidal, acute lip 
<quantity id="4C829B3CFFC49C30217F9559FE30F8A0" box="[357,479,1858,1883]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="0" pageNumber="194" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
long (vs. shortly conoidal 
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long), concave disc with broad half-ovoid callus bearing small central cavity (leading to the spur) at the base (vs. disc with two elongate basal calli without cavities and spur), and by the large, prominent subglobular callus at the front of column foot apex (vs. no swellings on column foot).
</paragraph>
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<paragraph id="8BC536D9FFC59C3120929513FCCBF8BA" blockId="1.[136,1452,1800,1857]" pageId="1" pageNumber="195">
<emphasis id="B90EEACBFFC59C3120929513FF10F8E6" bold="true" box="[136,255,1800,1821]" pageId="1" pageNumber="195">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4D5AFFC59C31211F9513FD0BF8E6" authority="Aver. et Nuraliev. Digital" authorityName="Aver. et Nuraliev. Digital" box="[261,740,1800,1821]" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="195" phylum="Tracheophyta" rank="species" species="longicheila">
<emphasis id="B90EEACBFFC59C31211F9513FE06F8E6" box="[261,489,1800,1821]" italics="true" pageId="1" pageNumber="195">Biermannia longicheila</emphasis>
Aver. et Nuraliev. Digital
</taxonomicName>
epitype, 
<emphasis id="B90EEACBFFC59C3123249513FBA1F8E6" box="[830,1102,1800,1821]" italics="true" pageId="1" pageNumber="195">Averyanov &amp; Nuraliev 0302</emphasis>
made from the type, 
<emphasis id="B90EEACBFFC59C31253A9513FF57F8BA" italics="true" pageId="1" pageNumber="195">Nuraliev et al. 1726</emphasis>
. Photos by M. Nuraliev, correction and design by L. Averyanov.
</paragraph>
</caption>
<paragraph id="8BC536D9FFC59C3320A7956AFC24FD02" blockId="1.[136,1459,1905,2074]" lastBlockId="3.[136,1452,160,1121]" lastPageId="3" lastPageNumber="197" pageId="1" pageNumber="195">
Monopodial, epiphytic herbs, generally glabrous. Stem (0.6)1.0–2.5(3.0) cm long, densely covered by distichous leaf sheaths, slightly flattened, with many pale green wiry roots clustered near stem base. Leaves (4)5–7(8), sessile, jointed, narrowly elliptic, sometimes slightly falcate, (4.5)5.0–8.0(8.5) × (1.4)1.6–2.2(2.4) cm, apex with two acute oblique unequal lobes. Inflorescence rather fleshy, glabrous; peduncle terete, (4)5–8(10) mm long, with 1–3(4) distant sterile triangular acute bracts, bracts (1.0)1.2–1.5(2.0) mm long and wide; rachis flattened (rectangular in cross section) with (4)6–8(10) distichous triangular floral bracts; floral bracts separated by (2.5)3.0–4.0(4.5) mm, triangular, acute, (2.0)2.2–2.4(2.8) mm long and wide; 1–3 flowers blooming simultaneously; inflorescence developing for a long time and thus flowers occupy its apical half. Flowers widely opening, fugacious, resupinate or not. Pedicel and ovary light green, hairy with short, scurfy, scattered brownish hairs, terete, triangular, 6-grooved, (3.8)4.0–4.4(4.8) mm long, 0.8–1.0 mm in diameter. Sepals subequal, broadly lanceolate to narrowly ovate, apiculate, straight, later recurved, (10.0)10.5–12.5(13.0) × (3.0)3.2–3.4(3.6) mm, pure white or with light yellowish-green tint near base; lateral sepals attached to column foot. Petals triangular, broadly lanceolate, acute, straight, as broad as sepals but slightly shorter, pure white. Lip at base with hemispheric, spur-like protrusion 0.8–1.0 mm in diameter, trilobed, firmly adnate at right angle to column foot, lying almost parallel to column, (8.0)8.5–9.0(10.0) × (3.2)3.5–4.0(4.2) mm, white, with four pale brown blotches near base; lateral lobes narrowly triangular, slightly falcate, erect, forward directed, acute, 1.0– 
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long; median lobe narrowly conoidal, (5.6)5.8–6.2(6.5) mm long, 2.0– 
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in diameter. Disc concave, at base with broad half-ovoid callus bearing small central cavity (leading to the spur) and with 2 pillow-like flattened, forwardly directed cushions at base of midlobe. Column white 2.2–2.4 × 1.8–2.0 mm; stigma almost circular, concave; column foot as long as column, at apex with white or pale purple prominent subglobular callus, 1.0– 
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in diameter, slightly bifid at apex. Operculum hemispheric, 
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in diameter, with an insignificant beak. Pollinia 2, globular, 
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in diameter, with a hardly visible cavity; stipe broadly ovate, concave, 0.5–0.6 × 
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wide, attenuate at base, viscidium half this size, ovate. Fruit and seeds unknown.
</paragraph>
</subSubSection>
<caption id="DF056651FFC69C32209295BEFBA5F7DD" ID-DOI="http://doi.org/10.5281/zenodo.13718591" ID-Zenodo-Dep="13718591" httpUri="https://zenodo.org/record/13718591/files/figure.png" pageId="2" pageNumber="196" startId="2.[136,229,1957,1978]" targetBox="[216,1368,195,1925]" targetPageId="2" targetType="figure">
<paragraph id="8BC536D9FFC69C32209295BEFBA5F7DD" blockId="2.[136,1452,1957,2086]" pageId="2" pageNumber="196">
<emphasis id="B90EEACBFFC69C32209295BEFF13F841" bold="true" box="[136,252,1957,1978]" pageId="2" pageNumber="196">FIGURE 2.</emphasis>
<taxonomicName id="4C7A4D5AFFC69C32211A95BEFE0DF841" box="[256,482,1957,1978]" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="196" phylum="Tracheophyta" rank="species" species="longicheila">
<emphasis id="B90EEACBFFC69C32211A95BEFE0DF841" box="[256,482,1957,1978]" italics="true" pageId="2" pageNumber="196">Biermannia longicheila</emphasis>
</taxonomicName>
. A. Flowering plant. B. Inflorescence. C. Rachis. D. Intact flowers, frontal and side views. E. Flattened flower, lip and pedicelate ovary. F. Intact lip, view from above. G. Intact lip, side view and sagittal section. H. Intact column and column foot, frontal view. I. Intact column and column foot, side view. J. Operculum, frontal view, side view and view from below. K. Pollinarium, frontal view and view from behind. All drawn from the type (
<emphasis id="B90EEACBFFC69C3222CD9A0AFC78F7DD" box="[727,919,2065,2086]" italics="true" pageId="2" pageNumber="196">Nuraliev et al. 1726</emphasis>
) by L. Averyanov.
</paragraph>
</caption>
<subSubSection id="C3606552FFC79C3320A7911FFE94FC72" pageId="3" pageNumber="197" type="etymology">
<paragraph id="8BC536D9FFC79C3320A7911FFBC1FCE6" blockId="3.[136,1452,160,1121]" box="[189,1070,772,797]" pageId="3" pageNumber="197">
<emphasis id="B90EEACBFFC79C3320A7911FFE8BFCE6" bold="true" box="[189,356,772,797]" pageId="3" pageNumber="197">Etymology:—</emphasis>
The specific epithet refers to the long narrow lip of this species.
</paragraph>
<paragraph id="8BC536D9FFC79C3320A79133FE94FC72" blockId="3.[136,1452,160,1121]" pageId="3" pageNumber="197">
<emphasis id="B90EEACBFFC79C3320A79133FD05FCBA" bold="true" box="[189,746,808,833]" pageId="3" pageNumber="197">Habitat, phenology and conservation status:—</emphasis>
Monopodial canopy epiphyte. Primary evergreen broad-leaved lowland forest along river valleys, 
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.
</specimenCount>
Flowering April–May. Known only from a single plant. Estimated IUCN Red List status—DD.
</paragraph>
</subSubSection>
<subSubSection id="C3606552FFC79C3320A7918FFD7AFB9A" pageId="3" pageNumber="197" type="distribution">
<paragraph id="8BC536D9FFC79C3320A7918FFCFCFC56" blockId="3.[136,1452,160,1121]" box="[189,787,916,941]" pageId="3" pageNumber="197">
<emphasis id="B90EEACBFFC79C3320A7918FFE9AFC56" bold="true" box="[189,373,916,941]" pageId="3" pageNumber="197">Distribution:—</emphasis>
Endemic to the 
<typeStatus id="54C1887BFFC79C33223C918FFDB9FC56" box="[550,598,916,941]" pageId="3" pageNumber="197">type</typeStatus>
locality thus far.
</paragraph>
<paragraph id="8BC536D9FFC79C3320A791A3FD7AFB9A" blockId="3.[136,1452,160,1121]" pageId="3" pageNumber="197">
<emphasis id="B90EEACBFFC79C3320A791A3FEE6FC2A" bold="true" box="[189,265,952,977]" pageId="3" pageNumber="197">Notes:</emphasis>
—The two species of the genus previously recorded in 
<collectingCountry id="F36D7649FFC79C33238891A3FC1BFC2A" box="[914,1012,952,977]" name="Vietnam" pageId="3" pageNumber="197">Vietnam</collectingCountry>
, i.e. common 
<taxonomicName id="4C7A4D5AFFC79C33248091A3FEE3FC0E" authority="Averyanov (1988: 429)" authorityName="Averyanov" authorityPageNumber="429" authorityYear="1988" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="197" phylum="Tracheophyta" rank="species" species="calcarata">
<emphasis id="B90EEACBFFC79C33248091A3FAC7FC2A" box="[1178,1320,952,977]" italics="true" pageId="3" pageNumber="197">B. calcarata</emphasis>
<bibRefCitation id="EFEB4B28FFC79C33252B91A3FEE3FC0E" author="Averyanov, L. V." pageId="3" pageNumber="197" pagination="423 - 432" refId="ref1787" refString="Averyanov, L. V. (1988) New taxa and nomenclatural changes in the Orchidaceae family of Vietnamese flora. Botanichesky Zhurnal 73: 423 - 432. [in Russian]" type="journal article" year="1988">Averyanov (1988: 429)</bibRefCitation>
</taxonomicName>
and insufficiently known 
<taxonomicName id="4C7A4D5AFFC79C33222E91C7FCD5FC0E" authority="Seidenfaden" authorityName="Seidenfaden" box="[564,826,988,1013]" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="197" phylum="Tracheophyta" rank="species" species="sigaldii">
<emphasis id="B90EEACBFFC79C33222E91C7FD47FC0E" box="[564,680,988,1013]" italics="true" pageId="3" pageNumber="197">B. sigaldii</emphasis>
Seidenfaden
</taxonomicName>
(1992: 433; Seidenfaden 1992, 
<bibRefCitation id="EFEB4B28FFC79C3324B991C7FA8FFC0E" author="Averyanov, L. V." box="[1187,1376,988,1013]" pageId="3" pageNumber="197" refId="ref1821" refString="Averyanov, L. V. (1994) Identification guide to Vietnamese orchids (Orchidaceae Juss.). Mir i Semya, St. Petersburg, 432 pp. [In Russian]" type="book" year="1994">Averyanov 1994</bibRefCitation>
, Pham Hoang Ho 2000, 
<bibRefCitation id="EFEB4B28FFC79C33215F961BFD48FBE2" author="Averyanov, L. V. &amp; Averyanova, A. L." box="[325,679,1024,1049]" pageId="3" pageNumber="197" refId="ref1856" refString="Averyanov, L. V. &amp; Averyanova, A. L. (2003) Updated checklist of the orchids of Vietnam. Vietnam National University Publishing House, Hanoi, 101 pp." type="book" year="2003">Averyanov &amp; Averyanova 2003</bibRefCitation>
, Nguyen Tien Ban 
<emphasis id="B90EEACBFFC79C332361961BFC5DFBE2" box="[891,946,1024,1049]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
2005) have no morphological similarity to 
<taxonomicName id="4C7A4D5AFFC79C332589961BFEE9FBC6" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="197" phylum="Tracheophyta" rank="species" species="longicheila">
<emphasis id="B90EEACBFFC79C332589961BFEE9FBC6" italics="true" pageId="3" pageNumber="197">B. longicheila</emphasis>
</taxonomicName>
. Records of 
<taxonomicName id="4C7A4D5AFFC79C33218C963FFDDCFBC6" authorityName="King &amp; Pantling" authorityYear="1898" baseAuthorityName="King &amp; Pantling" baseAuthorityYear="1895" box="[406,563,1060,1085]" class="Liliopsida" family="Orchidaceae" genus="Biermannia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="197" phylum="Tracheophyta" rank="species" species="bimaculata">
<emphasis id="B90EEACBFFC79C33218C963FFDDCFBC6" box="[406,563,1060,1085]" italics="true" pageId="3" pageNumber="197">B. bimaculata</emphasis>
</taxonomicName>
for 
<collectingCountry id="F36D7649FFC79C33227B963FFD2FFBC6" box="[609,704,1060,1085]" name="Vietnam" pageId="3" pageNumber="197">Vietnam</collectingCountry>
(Xu 
<emphasis id="B90EEACBFFC79C3322E0963FFCDDFBC6" box="[762,818,1060,1085]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
2010, Zhou 
<emphasis id="B90EEACBFFC79C3323DB963FFC15FBC6" box="[961,1018,1060,1085]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
2016) remain highly doubtful because they are not supported by any cited specimens.
</paragraph>
</subSubSection>
</treatment>
</document>