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<document id="C2A19C7205D5A742643DE60800F65D92" ID-DOI="10.11646/zootaxa.3980.2.8" ID-GBIF-Dataset="0b486dd1-c024-4898-a82d-ea06453554b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="244901" ID-ZooBank="BD6CE38A-4EB1-4932-B780-F9EAC0973AAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461241423396" checkinUser="plazi" docAuthor="Abstract, Joko Pamungkas" docDate="2015" docId="03D387B5FFEFFFCBBC92FF14FB69FECD" docLanguage="en" docName="zt03980p285.pdf" docOrigin="Zootaxa 3980 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polymastigos javaensis Abstract, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="284" masterDocId="FFEAFFCDFFEDFFCEBC05FF83FFEAFFD0" masterDocTitle="The description of a new species Polymastigos javaensis n. sp. (Annelida: Capitellidae) from the Segara Anakan mangroves, Central Java, Indonesia" masterLastPageNumber="285" masterPageNumber="279" pageNumber="281" updateTime="1698605196287" updateUser="plazi">
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<mods:title id="3532AD0EF9A7F4D5CA5A7ECE49F99E79">The description of a new species Polymastigos javaensis n. sp. (Annelida: Capitellidae) from the Segara Anakan mangroves, Central Java, Indonesia</mods:title>
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<mods:name id="8CE42CC6B382C515AC8F03B42ADF2D76" type="personal">
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<mods:namePart id="AAD8B83124D17C85F9447C3DDA571105">Abstract, Joko Pamungkas</mods:namePart>
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<mods:title id="09129B6AC7290C71AF640BF0C77C6540">Zootaxa</mods:title>
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<mods:part id="269A13E97BA244399E99CE10F6381677">
<mods:date id="250806D678DA6917F7E6120755A4B5C6">2015</mods:date>
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<mods:number id="9EEBB88382A9B759F89BB8A670FB491E">3980</mods:number>
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<mods:identifier id="6047BC5EEC3B1BA0FF4DE2C7DC74E85E" type="DOI">10.11646/zootaxa.3980.2.8</mods:identifier>
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<treatment id="03D387B5FFEFFFCBBC92FF14FB69FECD" ID-DOI="http://doi.org/10.5281/zenodo.6106051" ID-GBIF-Taxon="119633938" ID-Zenodo-Dep="6106051" LSID="urn:lsid:plazi:treatment:03D387B5FFEFFFCBBC92FF14FB69FECD" httpUri="http://treatment.plazi.org/id/03D387B5FFEFFFCBBC92FF14FB69FECD" lastPageId="5" lastPageNumber="284" pageId="2" pageNumber="281">
<subSubSection id="C3606528FFEFFFCCBC92FF14FE1CFF62" box="[151,502,151,178]" pageId="2" pageNumber="281" type="nomenclature">
<paragraph id="8BC536A3FFEFFFCCBC92FF14FE1CFF62" blockId="2.[151,502,151,178]" box="[151,502,151,178]" pageId="2" pageNumber="281">
<heading id="D08D81CFFFEFFFCCBC92FF14FE1CFF62" bold="true" box="[151,502,151,178]" fontSize="11" level="1" pageId="2" pageNumber="281" reason="1">
<emphasis id="B90EEAB1FFEFFFCCBC92FF14FE1CFF62" bold="true" box="[151,502,151,178]" pageId="2" pageNumber="281">
<taxonomicName id="4C7A4D20FFEFFFCCBC92FF14FE47FF61" ID-CoL="4LB3W" box="[151,429,151,177]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="2" pageNumber="281" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFEFFFCCBC92FF14FE47FF61" bold="true" box="[151,429,151,177]" italics="true" pageId="2" pageNumber="281">Polymastigos javaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57CAFFEFFFCCBDB1FF1BFE1CFF62" box="[436,502,152,178]" pageId="2" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3606528FFEFFFCCBC92FF5EFE2BFEEE" pageId="2" pageNumber="281" type="materials_examined">
<paragraph id="8BC536A3FFEFFFCCBC92FF5EFE2BFEEE" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
<emphasis id="B90EEAB1FFEFFFCCBC92FF5EFE6EFF26" bold="true" box="[151,388,221,246]" pageId="2" pageNumber="281">Material examined.</emphasis>
SA-01 (
<typeStatus id="54C18801FFEFFFCCBDEEFF5EFDBBFF26" box="[491,593,221,246]" pageId="2" pageNumber="281" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED6BAE66FFEFFFCCBE64FF5DFD4BFF25" box="[609,673,222,245]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00132); SA-01 (1
<typeStatus id="54C18801FFEFFFCCBFC5FF5DFBC9FF26" box="[960,1059,222,246]" pageId="2" pageNumber="281" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED6BAE66FFEFFFCCB836FF5DFB99FF25" box="[1075,1139,222,245]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00133); SA-01 (13,
<collectionCode id="ED6BAE66FFEFFFCCBC92FE81FF3DFEC9" box="[151,215,258,281]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00134); SA-02 (6,
<collectionCode id="ED6BAE66FFEFFFCCBDEEFE81FDC1FEC9" box="[491,555,258,281]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00135), SA-01 (1,
<collectionCode id="ED6BAE66FFEFFFCCBF3BFE81FC94FEC9" box="[830,894,258,281]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00136), SA-03 (7,
<collectionCode id="ED6BAE66FFEFFFCCB897FE81FB38FEC9" box="[1170,1234,258,281]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00137), SA- 0 4 (11,
<collectionCode id="ED6BAE66FFEFFFCCBCE9FEA5FEC6FEED" box="[236,300,294,317]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="281">MZB</collectionCode>
. Pol. 00138).
</paragraph>
</subSubSection>
<subSubSection id="C3606528FFEFFFCDBCC2FECAFEB8FE58" lastPageId="3" lastPageNumber="282" pageId="2" pageNumber="281" type="description">
<paragraph id="8BC536A3FFEFFFCCBCC2FECAFCE7FDEA" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
<emphasis id="B90EEAB1FFEFFFCCBCC2FECAFEB3FEB2" bold="true" box="[199,345,329,354]" pageId="2" pageNumber="281">Description.</emphasis>
<typeStatus id="54C18801FFEFFFCCBD60FECAFE27FEB2" box="[357,461,329,354]" pageId="2" pageNumber="281" type="holotype">Holotype</typeStatus>
complete with about 197 chaetigers,
<quantity id="4C829B46FFEFFFCCBF90FECAFC00FEB1" box="[917,1002,329,353]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="281" unit="mm" value="30.0">30 mm</quantity>
long by 1.0 mm wide in abdomen.
<typeStatus id="54C18801FFEFFFCCBC92FEEDFF13FE56" box="[151,249,366,390]" pageId="2" pageNumber="281" type="paratype">Paratype</typeStatus>
complete with about 251 chaetigers,
<quantity id="4C829B46FFEFFFCCBEAFFEEEFD16FE55" box="[682,764,365,390]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="281" unit="mm" value="35.0">35 mm</quantity>
long by 1.0 mm wide in abdomen. Other adult specimens complete or anterior fragments, ranging from
<quantity id="4C829B46FFEFFFCCBEA7FE12FD0FFE79" box="[674,741,401,425]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="281" unit="mm" value="8.0">8 mm</quantity>
long by
<quantity id="4C829B46FFEFFFCCBF49FE12FC4EFE79" box="[844,932,401,425]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="281" unit="mm" value="0.8">0.8 mm</quantity>
wide (32 chaetigers), to
<quantity id="4C829B46FFEFFFCCB8C7FE12FAF8FE79" box="[1218,1298,401,426]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="281" unit="mm" value="75.0">75 mm</quantity>
long by
<quantity id="4C829B46FFEFFFCCB97CFE12FF29FE1D" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="281" unit="mm" value="1.2">1.2 mm</quantity>
wide (about 366 chaetigers). Less mature specimens complete, with
<quantity id="4C829B46FFEFFFCCBFD4FE36FBCBFE1D" box="[977,1057,437,462]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="281" unit="mm" value="35.0">35 mm</quantity>
long by
<quantity id="4C829B46FFEFFFCCB883FE35FB37FE1D" box="[1158,1245,438,462]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="281" unit="mm" value="0.8">0.8 mm</quantity>
wide (about 224 chaetigers) and
<quantity id="4C829B46FFEFFFCCBD55FE5AFE49FE21" box="[336,419,473,498]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="281" unit="mm" value="53.0">53 mm</quantity>
long by
<quantity id="4C829B46FFEFFFCCBE14FE5AFD81FE21" box="[529,619,473,497]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="281" unit="mm" value="0.8">0.8 mm</quantity>
wide (about 237 chaetigers). Original colour in field reddish pink, in alcohol light brown. Prostomium conical with palpode. Eyespots and nuchal organs not seen. Proboscis partially everted, globular, with papillae. Peristomium achaetous.
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FDC6FEAFFD3E" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Thorax with 12 segments, including 1 achaetous peristomium followed posteriorly by 11 chaetigers with exclusively bilimbate capillary chaetae in both rami (
<figureCitation id="13412A26FFEFFFCCBF07FDEAFCA3FD52" box="[770,841,617,642]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Fig. 2</figureCitation>
A). All chaetigers, including chaetiger 1, biramous. Notopodia oriented laterally in first chaetiger, gradually moving dorsally in subsequent segments. Parapodia reduced. Chaetae embedded in middle part of thoracic segments, except in chaetiger 11 embedded more posteriorwards.
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FD7AFB2BFC72" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Peristomium to chaetiger 4 or 5 with areolated epithelium; remaining thorax smooth. Intersegmental groove between chaetiger 4 and 5 somewhat deeper than previous chaetigers. Based on these two characteristics, i.e. epithelium texture and intersegmental groove, thorax appears divided into two portions, i.e. areolated and smooth thorax (
<figureCitation id="13412A26FFEFFFCCBCF4FCE6FED1FCAE" box="[241,315,869,894]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Figs 2</figureCitation>
A &amp; 3AB, D). Thoracic segments biannulated. Areolated thorax less biannulated with obscure lateral line; smooth thorax more obviously biannulated with strong lateral line (
<figureCitation id="13412A26FFEFFFCCB815FC0AFBBCFC72" box="[1040,1110,905,930]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Figs 2</figureCitation>
A &amp; 3A).
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FC2EFCBEFC3A" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Lateral organs difficult to see. Genital pores lie on last four thoracic chaetigers, i.e. on the intersegmental grooves between chaetigers
<date id="FFC41063FFEFFFCCBDD6FC51FE13FC3A" box="[467,505,978,1002]" pageId="2" pageNumber="281" value="2011-07">7/8</date>
,
<date id="FFC41063FFEFFFCCBE00FC51FDC6FC3A" box="[517,556,978,1002]" pageId="2" pageNumber="281" value="2011-08">8/9</date>
,
<date id="FFC41063FFEFFFCCBE32FC51FD34FC3A" box="[567,734,977,1002]" pageId="2" pageNumber="281" value="2011-09-10" valueMax="2011-09-10" valueMin="2011-09-10">9/10 and 10/11</date>
(
<figureCitation id="13412A26FFEFFFCCBEE8FC52FCDBFC3A" box="[749,817,977,1002]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FC76FDA7FB86" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Transition between thorax and abdomen clearly marked by deep intersegmental groove between chaetiger 11 and 12 (
<figureCitation id="13412A26FFEFFFCCBCF0FB9AFEDDFBE2" box="[245,311,1049,1074]" captionStart="FIGURE 3" captionStartId="4.[151,250,1278,1300]" captionTargetBox="[182,1410,187,1255]" captionTargetId="figure@4.[594,1014,706,1256]" captionTargetPageId="4" captionText="FIGURE 3. Polymastigos javaensis n. sp. Methyl green staining pattern. A D = holotype, E F = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 5 19. D = ventral view, chaetiger 4 16. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view." httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="2" pageNumber="281">Fig. 3</figureCitation>
AD), distinct dorsal abdominal tori (
<figureCitation id="13412A26FFEFFFCCBEE2FB9AFCC5FBE2" box="[743,815,1049,1074]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Figs 2</figureCitation>
B &amp; 3BC), and changes to neurochaetae (
<figureCitation id="13412A26FFEFFFCCB91FFB9AFAB4FBE2" box="[1306,1374,1049,1074]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Fig. 2</figureCitation>
AB) and methyl green staining (
<figureCitation id="13412A26FFEFFFCCBDCDFBBEFDE0FB86" box="[456,522,1085,1110]" captionStart="FIGURE 3" captionStartId="4.[151,250,1278,1300]" captionTargetBox="[182,1410,187,1255]" captionTargetId="figure@4.[594,1014,706,1256]" captionTargetPageId="4" captionText="FIGURE 3. Polymastigos javaensis n. sp. Methyl green staining pattern. A D = holotype, E F = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 5 19. D = ventral view, chaetiger 4 16. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view." httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="2" pageNumber="281">Fig. 3</figureCitation>
AF).
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FBE2FB3EFAA6" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Dorsal surface of first several abdominal segments appear uni-annulate (sometimes with some weak furrows), changing posteriorly to more bi-annulate appearance in middle and posterior abdomen (sometimes with additional weak furrows), due to extension of ventral intra-annular groove dorsalwards. Ventral abdominal segments appear bi-annulate (
<figureCitation id="13412A26FFEFFFCCBD21FB4EFE86FB36" box="[292,364,1229,1254]" captionStart="FIGURE 2" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[180,1371,477,1904]" captionTargetId="figure@3.[151,1436,447,1934]" captionTargetPageId="3" captionText="FIGURE 2. Polymastigois javaensis n. sp. A B = holotype; C = MZB. Pol. 00136. A = lateral view. B = dorsal view, chaetiger 8 17; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: A B = 1 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="2" pageNumber="281">Figs 2</figureCitation>
B, 3D &amp; 3F). Abdominal segments tapering posteriorly; width and length of posterior segments smaller than those of anterior ones.
<typeStatus id="54C18801FFEFFFCCBE45FB72FD42FADA" box="[576,680,1265,1290]" pageId="2" pageNumber="281" type="holotype">Holotype</typeStatus>
with 5 transitional abdominal segments with bilimbate capillary notochaetae and neurohooks (chaetiger 12 to 16).
<typeStatus id="54C18801FFEFFFCCBEC3FA95FCC2FAFE" box="[710,808,1302,1326]" pageId="2" pageNumber="281" type="paratype">Paratype</typeStatus>
with 2 transitional abdominal segments (chaetiger 12 to 13); chaetiger
<date id="FFC41063FFEFFFCCBD3BFAB9FE01FA82" box="[318,491,1337,1362]" pageId="2" pageNumber="281">14 and 15 may</date>
have lost capillaries due to damage. Non-type materials vary, with 0 (less mature specimens) to 7 transitional abdominal segments. Remaining chaetigers with hooks in both rami.
</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2FA01FEDAF99E" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">In anterior abdomen, number of notohooks within a fascicle (about 20 hooks) fewer than neurohooks (about 43 hooks). Hooks within fascicle decrease in number posteriorly, about 5 notohooks and 23 neurohooks in posterior chaetigers. Abdominal hooks subequal. Notohooks with longer shaft than neurohooks, about two times. Length of anterior and posterior shaft nearly equal with an obvious node in middle of shaft. Hooks with 3 teeth above main fang, with narrow neck and shoulder. Short vague hood (hood with weak outline) observed in neurohooks, not seen in notohooks.</paragraph>
<paragraph id="8BC536A3FFEFFFCCBCC2F9DAFDE1F8D2" blockId="2.[151,1437,221,2010]" pageId="2" pageNumber="281">
Branchiae simple digitate, may be retractable, observed on dorsal region of posterior chaetigers, arise from intersegmental grooves, evident posteriorly from about chaetiger
<quantity id="4C829B46FFEFFFCCBF77F9FDFC41F946" box="[882,939,1661,1686]" metricMagnitude="0" metricUnit="m" metricValue="2.1082" pageId="2" pageNumber="281" unit="in" value="83.0">83 in</quantity>
<typeStatus id="54C18801FFEFFFCCBFB7F9FEFBFEF946" box="[946,1044,1661,1686]" pageId="2" pageNumber="281" type="holotype">holotype</typeStatus>
and about chaetiger
<quantity id="4C829B46FFEFFFCCB8F9F9FEFADFF946" box="[1276,1333,1661,1686]" metricMagnitude="0" metricUnit="m" metricValue="2.1844" pageId="2" pageNumber="281" unit="in" value="86.0">86 in</quantity>
<typeStatus id="54C18801FFEFFFCCB939F9FDFA76F946" box="[1340,1436,1662,1686]" pageId="2" pageNumber="281" type="paratype">paratype</typeStatus>
(
<figureCitation id="13412A26FFEFFFCCBC9AF922FF0BF96A" box="[159,225,1697,1722]" captionStart="FIGURE 3" captionStartId="4.[151,250,1278,1300]" captionTargetBox="[182,1410,187,1255]" captionTargetId="figure@4.[594,1014,706,1256]" captionTargetPageId="4" captionText="FIGURE 3. Polymastigos javaensis n. sp. Methyl green staining pattern. A D = holotype, E F = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 5 19. D = ventral view, chaetiger 4 16. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view." httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="2" pageNumber="281">Fig. 3</figureCitation>
E). Pygidium retracted in
<typeStatus id="54C18801FFEFFFCCBE04F922FD8CF96A" box="[513,614,1697,1722]" pageId="2" pageNumber="281" type="holotype">holotype</typeStatus>
, everted in
<typeStatus id="54C18801FFEFFFCCBEE3F921FCA3F96A" box="[742,841,1698,1722]" pageId="2" pageNumber="281" type="paratype">paratype</typeStatus>
, with mid-ventral caudal cirrus (
<figureCitation id="13412A26FFEFFFCCB8B1F922FB1CF96A" box="[1204,1270,1697,1722]" captionStart="FIGURE 3" captionStartId="4.[151,250,1278,1300]" captionTargetBox="[182,1410,187,1255]" captionTargetId="figure@4.[594,1014,706,1256]" captionTargetPageId="4" captionText="FIGURE 3. Polymastigos javaensis n. sp. Methyl green staining pattern. A D = holotype, E F = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 5 19. D = ventral view, chaetiger 4 16. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view." httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="2" pageNumber="281">Fig. 3</figureCitation>
F). Middle and posterior part of body with eggs in coelom. Egg oval, dark grey, around 300 µm and 800 µm, minimum and maximum diameter, respectively.
</paragraph>
<paragraph id="8BC536A3FFEFFFCDBCC2F88EFA80FECC" blockId="2.[151,1437,221,2010]" lastBlockId="3.[151,1437,151,392]" lastPageId="3" lastPageNumber="282" pageId="2" pageNumber="281">
Methyl green stain complete on thorax, including lateral line, intersegmental and intra-annular grooves. Dorsal region of proboscis, as well as prostomium to about half of chaetiger 2, stained greenish blue with medium intensity. Chaetiger 3, slightly extending to about half of chaetiger 2, stained dark blue. Chaetiger 4 with slightly brighter blue intensity than chaetiger 3. Ventral region of proboscis stained the darkest blue. In general, ventral side of areolated thorax stained darker than dorsal and lateral sides. Smooth thorax stained blue with rather more intensity than areolated thorax. Weakly stained areas on neuropodial lobes of chaetiger 4 to 11, forming oval appearance. Dorsal abdominal tori weakly stained with scattered blue spots. Area around notopodial lobes stained brightly, creating circle patterns; area around mid-dorsal line close to intersegmental areas stained darkly, forming longitudinal line. On abdominal ventrum, prechaetal annuli stained darkly. Mid-ventral region of posterior part with two dark longitudinal lines, interrupted by intersegmental ring. Pygidium rim stained darkly (
<figureCitation id="13412A26FFEEFFCDB8E0FE87FACDFECC" box="[1253,1319,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1278,1300]" captionTargetBox="[182,1410,187,1255]" captionTargetId="figure@4.[594,1014,706,1256]" captionTargetPageId="4" captionText="FIGURE 3. Polymastigos javaensis n. sp. Methyl green staining pattern. A D = holotype, E F = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 5 19. D = ventral view, chaetiger 4 16. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view." httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="3" pageNumber="282">Fig. 3</figureCitation>
AF).
</paragraph>
<paragraph id="8BC536A3FFEEFFCDBCC2FEABFEB8FE58" blockId="3.[151,1437,151,392]" pageId="3" pageNumber="282">Less mature specimens with 5 capillary chaetae (remaining chaetigers with hooks), smaller body, either absence or a few number of eggs in abdominal cavity, and show no branchiae. MGSP with similar appearance as in adult specimens.</paragraph>
</subSubSection>
<caption id="DF05662BFFEEFFCDBC92F813FCB8F833" httpUri="https://zenodo.org/record/244902/files/figure.png" pageId="3" pageNumber="282" targetBox="[180,1371,477,1904]" targetPageId="3">
<paragraph id="8BC536A3FFEEFFCDBC92F813FCB8F833" blockId="3.[151,1436,1936,2020]" pageId="3" pageNumber="282">
<emphasis id="B90EEAB1FFEEFFCDBC92F813FEE7F875" bold="true" box="[151,269,1936,1958]" pageId="3" pageNumber="282">FIGURE 2</emphasis>
.
<taxonomicName id="4C7A4D20FFEEFFCDBD1CF813FDE7F876" box="[281,525,1936,1958]" class="Polychaeta" family="Capitellidae" genus="Polymastigois" kingdom="Animalia" pageId="3" pageNumber="282" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFEEFFCDBD1CF813FDE7F876" box="[281,525,1936,1958]" italics="true" pageId="3" pageNumber="282">Polymastigois javaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57CAFFEEFFCDBE11F813FDA7F875" box="[532,589,1936,1957]" pageId="3" pageNumber="282" rank="species">
<emphasis id="B90EEAB1FFEEFFCDBE11F813FDA7F875" bold="true" box="[532,589,1936,1957]" pageId="3" pageNumber="282">n. sp.</emphasis>
</taxonomicNameLabel>
AB = holotype; C = MZB.Pol. 00136.A = lateral view. B = dorsal view, chaetiger 817; some abdominal capillaries lost during the removing process of mucus. C = neurohook (left) and notohook (right) of the anterior abdominal segment. Scale bars: AB = 1 mm; C = 0.01 mm.
</paragraph>
</caption>
<caption id="DF05662BFFE9FFCABC92FB7DFE23FA82" httpUri="https://zenodo.org/record/244903/files/figure.png" pageId="4" pageNumber="283" targetBox="[182,1410,187,1255]" targetPageId="4">
<paragraph id="8BC536A3FFE9FFCABC92FB7DFE23FA82" blockId="4.[151,1436,1278,1362]" pageId="4" pageNumber="283">
<emphasis id="B90EEAB1FFE9FFCABC92FB7DFEE6FAC4" bold="true" box="[151,268,1278,1300]" pageId="4" pageNumber="283">FIGURE 3</emphasis>
.
<taxonomicName id="4C7A4D20FFE9FFCABD1CFB7CFDEFFAC4" box="[281,517,1279,1300]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCABD1CFB7CFDEFFAC4" box="[281,517,1279,1300]" italics="true" pageId="4" pageNumber="283">Polymastigos javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCABE09FB7CFDAFFAC4" bold="true" box="[524,581,1279,1300]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCABE09FB7CFDAFFAC4" box="[524,581,1279,1300]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Methyl green staining pattern. AD = holotype, EF = paratype. A = lateral view. B = dorsal view. C = dorsal view, chaetiger 519. D = ventral view, chaetiger 416. E = anterior end with branchiae, dorsal view. F = anterior end, ventral view.
</paragraph>
</caption>
<subSubSection id="C3606528FFE9FFCABCC2FAF5FB36FA63" pageId="4" pageNumber="283" type="distribution">
<paragraph id="8BC536A3FFE9FFCABCC2FAF5FB36FA63" blockId="4.[151,1437,1398,2035]" pageId="4" pageNumber="283">
<emphasis id="B90EEAB1FFE9FFCABCC2FAF5FE8AFA5F" bold="true" box="[199,352,1398,1423]" pageId="4" pageNumber="283">Distribution.</emphasis>
Eastern area of the Segara Anakan mangrove habitat (nearby industrial area), Cilacap, Central Java Province,
<collectingCountry id="F36D7633FFE9FFCABD44FA19FE58FA63" box="[321,434,1434,1459]" name="Indonesia" pageId="4" pageNumber="283">Indonesia</collectingCountry>
. All specimens were collected in sandy clay sediment at
<quantity id="4C829B46FFE9FFCAB82BFA18FB65FA62" box="[1070,1167,1435,1459]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="4" pageNumber="283" unit="cm" value="15.0" valueMax="30.0" valueMin="0.0">030 cm</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C3606528FFE9FFCABCC2FA3DFBBFFA2B" pageId="4" pageNumber="283" type="etymology">
<paragraph id="8BC536A3FFE9FFCABCC2FA3DFBBFFA2B" blockId="4.[151,1437,1398,2035]" pageId="4" pageNumber="283">
<emphasis id="B90EEAB1FFE9FFCABCC2FA3DFEA4FA07" bold="true" box="[199,334,1470,1495]" pageId="4" pageNumber="283">Etymology.</emphasis>
The species is named based on its locality, i.e. Segara Anakan mangroves, Java Island,
<collectingCountry id="F36D7633FFE9FFCAB92DFA3DFA73FA07" box="[1320,1433,1470,1495]" name="Indonesia" pageId="4" pageNumber="283">Indonesia</collectingCountry>
. It is also the place where I spent so much time studying polychaetes for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C3606528FFE9FFCBBCC2F985FB69FECD" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283" type="discussion">
<paragraph id="8BC536A3FFE9FFCABCC2F985FB3DF823" blockId="4.[151,1437,1398,2035]" pageId="4" pageNumber="283">
<emphasis id="B90EEAB1FFE9FFCABCC2F985FEDEF9CF" bold="true" box="[199,308,1542,1567]" pageId="4" pageNumber="283">Remarks</emphasis>
. The new species
<taxonomicName id="4C7A4D20FFE9FFCABE12F984FD4AF9CE" box="[535,672,1543,1566]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCABE12F984FD4AF9CE" box="[535,672,1543,1566]" italics="true" pageId="4" pageNumber="283">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCABEA9F985FD1BF9CF" bold="true" box="[684,753,1542,1567]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCABEA9F985FD1BF9CF" box="[684,753,1542,1567]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resembles
<emphasis id="B90EEAB1FFE9FFCABF7FF984FC3CF9CE" box="[890,982,1542,1567]" italics="true" pageId="4" pageNumber="283">P. re i s h i</emphasis>
in having more than four transitional abdominal segments and in the position of their genital pores on the intersegmental grooves between chaetigers
<date id="FFC41063FFE9FFCAB983F9A8FF4DF9B7" pageId="4" pageNumber="283" value="2011-07">7/ 8</date>
,
<date id="FFC41063FFE9FFCABCB6F9CCFF33F9B7" box="[179,217,1615,1639]" pageId="4" pageNumber="283" value="2011-08">8/9</date>
,
<date id="FFC41063FFE9FFCABCE0F9CCFE65F9B7" box="[229,399,1614,1639]" pageId="4" pageNumber="283" value="2011-09-10" valueMax="2011-09-10" valueMin="2011-09-10">9/10 and 10/11</date>
, but the new species differs in the following features. First, chaetiger 1 of
<taxonomicName id="4C7A4D20FFE9FFCAB8D4F9CCFABFF9B6" box="[1233,1365,1615,1638]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCAB8D4F9CCFABFF9B6" box="[1233,1365,1615,1638]" italics="true" pageId="4" pageNumber="283">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCAB959F9CDFA76F9B7" bold="true" box="[1372,1436,1614,1639]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCAB959F9CDFA76F9B7" box="[1372,1436,1614,1639]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is biramous, whereas it is uniramous in
<emphasis id="B90EEAB1FFE9FFCABE65F9F7FD52F95A" box="[608,696,1650,1675]" italics="true" pageId="4" pageNumber="283">P. re i s h i</emphasis>
. Further, Green did not mention segment width difference as an important character to define
<emphasis id="B90EEAB1FFE9FFCABDE2F914FDD4F97E" box="[487,574,1686,1711]" italics="true" pageId="4" pageNumber="283">P. re i s h i</emphasis>
, but in the authors observation, the feature is a distinctive characteristic of
<taxonomicName id="4C7A4D20FFE9FFCABC92F938FEF9F902" box="[151,275,1723,1746]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCABC92F938FEF9F902" box="[151,275,1723,1746]" italics="true" pageId="4" pageNumber="283">P.javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCABD1EF939FEB1F903" bold="true" box="[283,347,1722,1747]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCABD1EF939FEB1F903" box="[283,347,1722,1747]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The character makes the whole body of
<taxonomicName id="4C7A4D20FFE9FFCABF22F93FFC46F902" box="[807,940,1723,1747]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCABF22F93FFC46F902" box="[807,940,1723,1747]" italics="true" pageId="4" pageNumber="283">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCABFB6F939FC19F903" bold="true" box="[947,1011,1722,1747]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCABFB6F939FC19F903" box="[947,1011,1722,1747]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
seem to be divided into four portions when seen from a dorsal view, i.e. areolated thorax (with weak lateral line), smooth thorax (with strong lateral line), anterior abdominal region (without branchiae) and posterior abdominal region (with branchiae and pygidium). Another difference is that dorsal surface of
<emphasis id="B90EEAB1FFE9FFCABE86F8ABFCE9F8EF" box="[643,771,1832,1855]" italics="true" pageId="4" pageNumber="283">P. j a v a en s i s</emphasis>
<emphasis id="B90EEAB1FFE9FFCABF0BF8A5FCA4F8EF" bold="true" box="[782,846,1830,1855]" pageId="4" pageNumber="283">n. sp.</emphasis>
first several abdominal segments look uni-annulate, changing to more bi-annulate appearance in mid and posterior segments, whereas Green described the dorsum of
<taxonomicName id="4C7A4D20FFE9FFCAB982F8CFFF3FF857" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="reishi">
<emphasis id="B90EEAB1FFE9FFCAB982F8CFFF3FF857" italics="true" pageId="4" pageNumber="283">P. reishi</emphasis>
</taxonomicName>
s abdominal segments as multi-annulate. The annular pattern of the anterior abdominal ventrum between them is also different (this can be more clearly seen after staining with methyl green).
<taxonomicName id="4C7A4D20FFE9FFCAB857F811FABDF87A" box="[1106,1367,1938,1963]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE9FFCAB857F811FABDF87A" box="[1106,1367,1938,1963]" italics="true" pageId="4" pageNumber="283">Polymastigos javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE9FFCAB958F811FA76F87B" bold="true" box="[1373,1436,1938,1963]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A23D57CAFFE9FFCAB958F811FA76F87B" box="[1373,1436,1938,1963]" pageId="4" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shows prechaetal annuli that are slightly similar to diamond (with a large angle around the mid-ventral line and a low angle around lateral sides—Fig.3D), whereas
<taxonomicName id="4C7A4D20FFE9FFCABECFF85FFCCEF823" box="[714,804,2010,2035]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="reishi">
<emphasis id="B90EEAB1FFE9FFCABECFF85FFCCEF823" box="[714,804,2010,2035]" italics="true" pageId="4" pageNumber="283">P. reishi</emphasis>
</taxonomicName>
has rectangular-like prechaetal annuli.
</paragraph>
<paragraph id="8BC536A3FFE8FFCBBCC2FF1BFB69FECD" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="284">
Next, the hooks of
<taxonomicName id="4C7A4D20FFE8FFCBBDA6FF1AFDC6FF60" box="[419,556,153,176]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE8FFCBBDA6FF1AFDC6FF60" box="[419,556,153,176]" italics="true" pageId="5" pageNumber="284">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE8FFCBBE30FF14FD9FFF60" bold="true" box="[565,629,151,176]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A23D57CAFFE8FFCBBE30FF14FD9FFF60" box="[565,629,151,176]" pageId="5" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are either without hood (notohooks) or with relatively small and short hood (seen in neurohooks), while
<taxonomicName id="4C7A4D20FFE8FFCBBE14FF3EFD85FF05" box="[529,623,188,213]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="reishi">
<emphasis id="B90EEAB1FFE8FFCBBE14FF3EFD85FF05" box="[529,623,188,213]" italics="true" pageId="5" pageNumber="284">P. reishi</emphasis>
</taxonomicName>
s hooks with long hood with relatively long fringe. In
<taxonomicName id="4C7A4D20FFE8FFCBB8D5FF3EFABDFF04" box="[1232,1367,189,212]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE8FFCBB8D5FF3EFABDFF04" box="[1232,1367,189,212]" italics="true" pageId="5" pageNumber="284">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE8FFCBB958FF3FFA76FF05" bold="true" box="[1373,1436,188,213]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A23D57CAFFE8FFCBB958FF3FFA76FF05" box="[1373,1436,188,213]" pageId="5" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the number of hooks within a fascicle decrease in number posteriorly, but they increase in
<taxonomicName id="4C7A4D20FFE8FFCBB8B4FF62FAF9FF28" box="[1201,1299,223,248]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="reishi">
<emphasis id="B90EEAB1FFE8FFCBB8B4FF62FAF9FF28" box="[1201,1299,223,248]" italics="true" pageId="5" pageNumber="284">P. reishi</emphasis>
</taxonomicName>
. Lastly, the MGSP between
<taxonomicName id="4C7A4D20FFE8FFCBBD4BFE86FE3FFECC" box="[334,469,261,284]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B90EEAB1FFE8FFCBBD4BFE86FE3FFECC" box="[334,469,261,284]" italics="true" pageId="5" pageNumber="284">P. javaensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAB1FFE8FFCBBDD9FE87FDF1FECD" bold="true" box="[476,539,260,285]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A23D57CAFFE8FFCBBDD9FE87FDF1FECD" box="[476,539,260,285]" pageId="5" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C7A4D20FFE8FFCBBE57FE86FD5BFECD" box="[594,689,260,285]" class="Polychaeta" family="Capitellidae" genus="Polymastigos" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="reishi">
<emphasis id="B90EEAB1FFE8FFCBBE57FE86FD5BFECD" box="[594,689,260,285]" italics="true" pageId="5" pageNumber="284">P. reishi</emphasis>
</taxonomicName>
is also very different as explained above.
</paragraph>
</subSubSection>
</treatment>
</document>