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<document id="0121E2AD98BC2CCD76684AB648FE274B" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625624287261" checkinUser="felipe" docAuthor="Published, First" docDate="2007" docId="03CE87EB49590732FF36FF32FD45DB37" docLanguage="en" docName="zt01404p066.pdf" docOrigin="Zootaxa 1404" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Tokunagaia azari Published 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="44" masterDocId="FFF7FF9349730719FFA1FF88FFC8DE5F" masterDocTitle="Chironomid midges from early Eocene amber of France (Diptera: Chironomidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="43" updateTime="1698992602245" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6483235E406941C94999A4A880A36DBB">Chironomid midges from early Eocene amber of France (Diptera: Chironomidae)</mods:title>
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<mods:date id="53AEAC18AF665D3FC10C0F2FF554FB3C">2007</mods:date>
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<mods:number id="2D68D30D9FBFC24B19556E29411B0515">2007-02-05</mods:number>
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<treatment id="03CE87EB49590732FF36FF32FD45DB37" ID-DOI="http://doi.org/10.5281/zenodo.5076805" ID-Zenodo-Dep="5076805" LSID="urn:lsid:plazi:treatment:03CE87EB49590732FF36FF32FD45DB37" httpUri="http://treatment.plazi.org/id/03CE87EB49590732FF36FF32FD45DB37" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="C37D657649590733FF36FF32FE7CDE8B" box="[151,436,186,212]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BD836FD49590733FF36FF32FE7CDE8B" blockId="42.[151,622,151,247]" box="[151,436,186,212]" pageId="42" pageNumber="43">
<heading id="D090819149590733FF36FF32FE7CDE8B" bold="true" box="[151,436,186,212]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B913EAEF49590733FF36FF32FE7CDE8B" bold="true" box="[151,436,186,212]" pageId="42" pageNumber="43">
<taxonomicName id="4C674D7E49590733FF36FF32FEA2DE8B" ID-CoL="8FGC8" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,362,186,212]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="azari" status="sp. nov.">
<emphasis id="B913EAEF49590733FF36FF32FEA2DE8B" bold="true" box="[151,362,186,212]" italics="true" pageId="42" pageNumber="43">Tokunagaia azari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220579449590733FED3FF32FE7CDE8B" box="[370,436,186,212]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D657649590733FF36FF55FEE7DEA8" box="[151,303,221,247]" pageId="42" pageNumber="43" type="description">
<paragraph id="8BD836FD49590733FF36FF55FEE7DEA8" blockId="42.[151,622,151,247]" box="[151,303,221,247]" pageId="42" pageNumber="43">
(
<figureCitation id="135C2A7849590733FF3EFF55FEC8DEA8" box="[159,256,221,247]" captionStart="FIGURE 38" captionStartId="42.[151,255,1909,1933]" captionText="FIGURE 38. Tokunagaia azari n. sp., holotype PA 11435, photograph of habitus." pageId="42" pageNumber="43">Figs. 38</figureCitation>
­40)
</paragraph>
</subSubSection>
<subSubSection id="C37D657649590733FF36FEA5FAC2D98C" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BD836FD49590733FF36FEA5FB1FDF18" blockId="42.[151,1239,301,327]" box="[151,1239,301,327]" pageId="42" pageNumber="43">
<emphasis id="B913EAEF49590733FF36FEA5FEEFDF18" bold="true" box="[151,295,301,327]" pageId="42" pageNumber="43">Etymology:</emphasis>
We name this species for our colleague Dr. Dany Azar (University of
<collectingCountry id="F370766D49590733FBFEFEA5FB03DF18" box="[1119,1227,301,327]" name="Lebanon" pageId="42" pageNumber="43">Lebanon</collectingCountry>
).
</paragraph>
<caption id="DF18667549590733FF36F8FDFC37D9D3" box="[151,1023,1909,1933]" pageId="42" pageNumber="43" startId="42.[151,255,1909,1933]">
<paragraph id="8BD836FD49590733FF36F8FDFC37D9D3" blockId="42.[151,1290,1909,2004]" box="[151,1023,1909,1933]" pageId="42" pageNumber="43">
<emphasis id="B913EAEF49590733FF36F8FDFEEED9D2" bold="true" box="[151,294,1909,1933]" pageId="42" pageNumber="43">FIGURE 38.</emphasis>
<taxonomicName id="4C674D7E49590733FE8CF8FDFE20D9D3" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[301,488,1909,1932]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="azari" status="sp. nov.">
<emphasis id="B913EAEF49590733FE8CF8FDFE20D9D3" box="[301,488,1909,1932]" italics="true" pageId="42" pageNumber="43">Tokunagaia azari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220579449590733FE4FF8FDFDE1D9D3" box="[494,553,1909,1932]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
, holotype PA 11435, photograph of habitus.
</paragraph>
</caption>
<paragraph id="8BD836FD49590733FF36F810FB80D9F0" blockId="42.[151,1290,1909,2004]" box="[151,1096,1944,1968]" pageId="42" pageNumber="43">
<emphasis id="B913EAEF49590733FF36F810FEEED9EF" bold="true" box="[151,294,1944,1968]" pageId="42" pageNumber="43">FIGURE 39.</emphasis>
<taxonomicName id="4C674D7E49590733FE8CF810FE20D9F0" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[301,488,1944,1967]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="azari" status="sp. nov.">
<emphasis id="B913EAEF49590733FE8CF810FE20D9F0" box="[301,488,1944,1967]" italics="true" pageId="42" pageNumber="43">Tokunagaia azari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220579449590733FE4FF810FDE1D9F0" box="[494,553,1944,1967]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
,
<typeStatus id="54DC885F49590733FD95F810FD59D9F0" box="[564,657,1944,1967]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
PA 11435, photograph of male genitalia.
</paragraph>
<paragraph id="8BD836FD49590733FF36F834FAC2D98C" blockId="42.[151,1290,1909,2004]" box="[151,1290,1980,2004]" pageId="42" pageNumber="43">
<emphasis id="B913EAEF49590733FF36F834FEEED98B" bold="true" box="[151,294,1980,2004]" pageId="42" pageNumber="43">FIGURE 40.</emphasis>
<taxonomicName id="4C674D7E49590733FE8CF835FE20D98B" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[301,488,1981,2004]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="azari" status="sp. nov.">
<emphasis id="B913EAEF49590733FE8CF835FE20D98B" box="[301,488,1981,2004]" italics="true" pageId="42" pageNumber="43">Tokunagaia azari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220579449590733FE4FF834FDE1D98C" box="[494,553,1980,2003]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
,
<typeStatus id="54DC885F49590733FD95F834FD59D98C" box="[564,657,1980,2003]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
PA 11435, drawing of male genitalia (scale bar =
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).
</paragraph>
</subSubSection>
<subSubSection id="C37D657649580732FF67FF1FFF36DF5D" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BD836FD49580732FF67FF1FFF36DF5D" blockId="43.[151,1438,151,1384]" pageId="43" pageNumber="44">
<emphasis id="B913EAEF49580732FF67FF1FFE80DEEE" bold="true" box="[198,328,151,177]" pageId="43" pageNumber="44">Diagnosis:</emphasis>
Eye with short wedge­shaped dorsomedial extension; antennal ratio 0.46; wing membrane without macrotrichia; anal point absent; gonostylus simple, with low crista dorsalis, tergite IX with less than 20 setae.
</paragraph>
</subSubSection>
<subSubSection id="C37D657649580732FF67FE87FBFDDD88" pageId="43" pageNumber="44" type="description">
<paragraph id="8BD836FD49580732FF67FE87FEDEDFAB" blockId="43.[151,1438,151,1384]" pageId="43" pageNumber="44">
<emphasis id="B913EAEF49580732FF67FE87FEA8DF76" bold="true" box="[198,352,271,297]" pageId="43" pageNumber="44">Description:</emphasis>
Head
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long, deformed; ocelli absent; antenna
<quantity id="4C9F9B1849580732FBA6FE98FBBCDF75" box="[1031,1140,272,298]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="43" pageNumber="44" unit="mm" value="0.41">0.41 mm</quantity>
long, much longer than head, distinctly hairy, with 13 flagellomeres covered with long setae (shortest
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long, longest
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long), pedicel broad and short, rounded, 13
<superScript id="7C129BB549580732FD3BFEE8FD6EDF31" attach="left" box="[666,678,352,366]" fontSize="6" pageId="43" pageNumber="44">th</superScript>
flagellomere
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long; antennal ratio 0.46; eye bare, with small dorsomedial extension, with 2 rows of ommatidia at minimum width; mouthparts lacking functional mandibles; 5 palpomeres with setae; 1 postocular; frontal and inner and outer vertical setae not visible, probably absent.
</paragraph>
<paragraph id="8BD836FD49580732FF67FD8AFBFDDD88" blockId="43.[151,1438,151,1384]" pageId="43" pageNumber="44">
Thorax
<quantity id="4C9F9B1849580732FE82FD8AFE45DC43" box="[291,397,514,540]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="43" pageNumber="44" unit="mm" value="0.34">0.34 mm</quantity>
long,
<quantity id="4C9F9B1849580732FE7BFD8AFD8CDC43" box="[474,580,514,540]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="43" pageNumber="44" unit="mm" value="0.19">0.19 mm</quantity>
wide,
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high; postnotum bare, with longitudinal median groove; surface of scutellum bare; without scutal tubercle; acrostichals absent, dorsocentrals uniserial; scutum without median longitudinal groove; 2 prealar setae, no supraalar setae; epimeron II and preepisternum bare. Wing macropterous,
<quantity id="4C9F9B1849580732FEE9FDF2FE7BDCCB" box="[328,435,634,660]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="43" pageNumber="44" unit="mm" value="0.88">0.88 mm</quantity>
long,
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wide, hyaline, membrane bare, but with coarse punctation; anal vein An
<subScript id="17E334B849580732FF1AFD3BFF0BDC9E" attach="left" box="[187,195,691,705]" fontSize="6" pageId="43" pageNumber="44">2</subScript>
absent; radius with 3 branches R
<subScript id="17E334B849580732FDC1FD3BFDA0DC9E" attach="left" box="[608,616,691,705]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
, R and R
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, R
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not divided into R
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and R
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; only M
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and M
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present; cross­vein MCu absent; squama with 2 setae. Halter
<quantity id="4C9F9B1849580732FCD6FD45FC1ADCB8" box="[887,978,717,743]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="mm" value="0.1">0.1 mm</quantity>
long. Fore femur
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long, tibia
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long, tarsus
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long; mid femur
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long, tibia
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long, tarsus
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long; hind femur
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long, tibia
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long, tarsus
<quantity id="4C9F9B1849580732FD73FC95FCF2DD68" box="[722,826,797,823]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="43" pageNumber="44" unit="mm" value="0.38">0.38 mm</quantity>
long; ta4 of all legs cylindrical, not cordiform; fore tibia with 1 long spur, mid and hind tibiae with 2 long spurs, hind tibial comb well developed; small pulvilli present. Abdomen
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long; tergite IX with about 5­10 setae; gonostylus
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long,
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wide, simple, elongate, with low crista dorsalis, hinged to gonocoxite and folded inward; gonocoxite
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long,
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wide, with long setae apically and on outer surface; anal point absent.
</paragraph>
</subSubSection>
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<subScript id="17E334B849580732FD2FFD3BFD6FDC9E" attach="left" box="[654,679,691,705]" fontSize="6" pageId="43" pageNumber="44">2+3</subScript>
</paragraph>
<subSubSection id="C37D657649580732FF67FC6DFA85DB1F" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BD836FD49580732FF67FC6DFA85DB1F" blockId="43.[151,1438,151,1384]" pageId="43" pageNumber="44">
<emphasis id="B913EAEF49580732FF67FC6DFE9CDDA0" bold="true" box="[198,340,997,1023]" pageId="43" pageNumber="44">Discussion:</emphasis>
In the key to Holarctic orthocladiine genera of Cranston
<emphasis id="B913EAEF49580732FBB0FC6EFB80DDA0" box="[1041,1096,998,1023]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
. (1989), this
<taxonomicName id="4C674D7E49580732FB4FFC6DFA54DDA0" authorityName="Edwards" authorityYear="1929" box="[1262,1436,997,1023]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subFamily" subFamily="Orthocladiinae">Orthocladiinae</taxonomicName>
falls in the genus
<taxonomicName id="4C674D7E49580732FEC4FB86FE26DA78" authorityName="Goetghebuer" authorityYear="1932" box="[357,494,1038,1063]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAEF49580732FEC4FB86FE26DA78" box="[357,494,1038,1063]" italics="true" pageId="43" pageNumber="44">Tokunagaia</emphasis>
</taxonomicName>
because of the following characters: wing well developed; without thick clavus; scutum without median longitudinal groove; eye bare; wing membrane without macrotrichia; epimeron II and preepisternum bare; no lateral spines on spurs of mid and hind tibiae; anal point without strong lamellate setae laterally and apicodorsally; squama with at least 1 seta; small pulvilli, never as long as half claw length; hind tibial comb well developed; tergite IX without strongly elevated ridge; anal point absent; gonostylus simple; and eye with short wedge­shaped dorsomedial extension. The species is near the group of extant species including
<taxonomicName id="4C674D7E49580732FEACFB76FE54DB48" baseAuthorityName="Brundin" baseAuthorityYear="1956" box="[269,412,1278,1303]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="B913EAEF49580732FEACFB76FE54DB48" box="[269,412,1278,1303]" italics="true" pageId="43" pageNumber="44">T. scutellata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D7E49580732FE0AFB76FDE7DB48" baseAuthorityName="Saether" baseAuthorityYear="1969" box="[427,559,1278,1303]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="rowensis">
<emphasis id="B913EAEF49580732FE0AFB76FDE7DB48" box="[427,559,1278,1303]" italics="true" pageId="43" pageNumber="44">T. rowensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C674D7E49580732FDD1FB76FCE2DB48" baseAuthorityName="Goetghebuer" baseAuthorityYear="1940" box="[624,810,1278,1303]" class="Insecta" family="Chironomidae" genus="Tokunagaia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B913EAEF49580732FDD1FB76FCE2DB48" box="[624,810,1278,1303]" italics="true" pageId="43" pageNumber="44">T. rectangularis</emphasis>
</taxonomicName>
because it has no anal point, a gonostylus with a low crista dorsalis, antennal ratio 0.46, and tergite IX with fewer than 20 setae (
<bibRefCitation id="EFF64B0C49580732FBA9FAAEFA88DB1F" author="Halvorsen, G. A. &amp; Saether, O. A." box="[1032,1344,1318,1344]" pageId="43" pageNumber="44" pagination="173 - 188" refId="ref37849" refString="Halvorsen, G. A. &amp; Saether, O. A. (1987) Redefinition and revision of the genus Tokunagaia Saether, 1973 (Diptera: Chironomidae). Entomologica Scandinavica, (Supplement), 29, 173 - 188." type="journal article" year="1987">Halvorsen &amp; Saether 1987</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D657649580732FF67FAC5FD45DB37" box="[198,653,1357,1384]" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BD836FD49580732FF67FAC5FD45DB37" blockId="43.[151,1438,151,1384]" box="[198,653,1357,1384]" pageId="43" pageNumber="44">
<emphasis id="B913EAEF49580732FF67FAC5FEF5DB38" bold="true" box="[198,317,1357,1383]" pageId="43" pageNumber="44">Material:</emphasis>
<typeStatus id="54DC885F49580732FEE5FAC6FE79DB37" box="[324,433,1358,1384]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
PA 11435, (male).
</paragraph>
</subSubSection>
</treatment>
</document>